{{Short description|Family of tortrix moths}} {{About|the family of moths}} {{Automatic taxobox | fossil_range = {{fossilrange|Eocene|Present}} | image = Bactra.lancealana.jpg | image_caption = ''Bactra lancealana'' | display_parents = 5 | parent_authority = Latreille, 1803 | taxon = Tortricidae | authority = Latreille, 1803 | diversity = Over 1,050 genera<br/>Over 10,350 species | subdivision_ranks = Subfamilies & tribes | subdivision = *Chlidanotinae *Tortricinae *Olethreutinae See also Taxonomy of Tortricidae for full list of genera. | synonyms = Olethreutidae }}
The '''Tortricidae''' are a family of moths, commonly known as '''tortrix moths''' or '''leafroller moths''',<ref>{{cite web |last=McLeod |first=Robin |date=December 31, 2019 |url=https://bugguide.net/node/view/9524 |title=Family Tortricidae - Tortricid Moths |website=BugGuide |access-date=February 7, 2020 |archive-date=May 17, 2024 |archive-url=https://web.archive.org/web/20240517135040/https://bugguide.net/node/view/9524 |url-status=live }}</ref> in the order Lepidoptera. This large family has over 11,000 species described, and is the sole member of the superfamily '''Tortricoidea''', although the genus ''Heliocosma'' is sometimes placed within this superfamily.<ref>{{cite journal| author=van Nieukerken| title=Order Lepidoptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness| journal=Zootaxa| volume=3148| year=2011| pages=212–221| doi=10.11646/zootaxa.3148.1.41| url=http://mapress.com/zootaxa/2011/f/zt03148p221.pdf| display-authors=etal| access-date=2012-01-14| archive-date=2019-10-13| archive-url=https://web.archive.org/web/20191013141544/https://mapress.com/zootaxa/2011/f/zt03148p221.pdf| url-status=live}}</ref><ref>{{cite news |title=Tracking our Taxonomic Progress |url=http://www.tortricidae.com/TORTS_newsletter20(1).pdf |work=Torts. Newsletter of the Troop of Reputed Tortricid Systematists |volume=20 |issue=1 |date=8 January 2019 |pages=12–13 |access-date=8 April 2021 |archive-date=17 May 2024 |archive-url=https://web.archive.org/web/20240517135053/http://www.tortricidae.com/TORTS_newsletter20(1).pdf |url-status=live }}</ref> Many of these are economically important pests. '''Olethreutidae''' is a junior synonym. The typical resting posture is with the wings folded back, producing a rather rounded profile.
Notable tortricids include the codling moth and the spruce budworm, which are among the most well-studied of all insects because of their economic impact.<ref>{{Cite book |url=https://books.google.com/books?id=MkJCsEctl1UC |title=Tortricidae (Lepidoptera) |last=Brown |first=John W. |date=2005 |publisher=Apollo Books |isbn=9788788757415 |language=en |access-date=2016-05-08 |archive-date=2024-05-17 |archive-url=https://web.archive.org/web/20240517135040/https://books.google.com/books?id=MkJCsEctl1UC |url-status=live }}</ref>
==Description== Tortricid moths are generally small, with a wingspan of 3 cm or less.<ref>Hanson, Paul E. (2018). ''Insects and Other Arthropods of Tropical America''. Cornell University Press. {{ISBN|978-0-8014-5694-7}}</ref> Many species are drab and have mottled and marbled brown colors, but some diurnal species are brightly colored and mimic other moths of the families Geometridae and Pyralidae.
==Life cycle and behavior== Tortricid eggs are often flattened and scale-like.
Larvae in the subfamilies Chlidanotinae and Olethreutinae usually feed by boring into stems, roots, buds or seeds. Larvae in the subfamily Tortricinae, however, feed externally and construct leaf rolls. Larvae in the subfamily Tortricinae tend to be more polyphagous than those in Chlidanotinae and Olethreutinae. Tortricinae also possess an anal fork for flicking excrement away from their shelters.
==Some common tortricids== The tortricids include many economically important pests, including: *Summer fruit tortrix moth (''Adoxophyes orana'') *Fruit tree tortrix moth (''Archips podana'') *Rose leaf roller (''Archips rosana'') *''Argyrotaenia ljungiana'', a pest on vines, maize, and fruit trees *Peach moth (''Cydia molesta'') *Codling moth (''Cydia pomonella'') *Plum fruit moth (''Cydia funebrana'') *Pea moth (''Cydia nigricana'') *Chestnut and acorn moth (''Cydia splendana'') *Light brown apple moth (''Epiphyas postvittana'') *Hemp borer (''Grapholita delineana'') *Oriental fruit moth (''Grapholita molesta'') *Cherry fruitworm (''Grapholita packardi'') *European grapevine moth (''Lobesia botrana'') *Barred fruit tree tortrix moth (''Pandemis cerasana'') *Grape berry moth (''Paralobesia viteana'') *Long-palped tortrix or vine leaf roller (''Sparganothis pilleriana'') *Bud moth (''Spilonota ocellana'') *False codling moth (''Thaumatotibia (Cryptophlebia) leucotreta'') *Spruce budworm (Genus ''Choristoneura'')
''See also'' Mexican jumping bean moth (''Cydia saltitans'')
===A typical tortricid – the codling moth=== The Tortricidae are considered to be the single most important family of insects that feed on apples, both economically and in diversity of feeding found on fruit, buds, leaves, and shoots. In New York, no fewer than seventeen species of Tortricidae have gained pest status in regards to apple production.{{Citation needed|date=January 2014}}
The codling moth ''Cydia pomonella'' causes worm-holes in apples. It has been accidentally spread from its original range in Europe and is now found in North and South America, South Africa, Australia, and New Zealand, wherever apples are grown. Control has required the use of the harshest available insecticides – historically lead arsenate and DDT were used for control. These chemicals brought considerable environmental dangers, and in any case the insect gradually developed resistance to them. Currently, organophosphate sprays are favored and are timed carefully to catch the hatching larvae before they can bore into the fruit.
== Gallery == <gallery> Choristoneura fumiferana larva.jpg|''Choristoneura fumiferana'' larva Choristoneura lambertiana pupa.jpg|''Choristoneura lambertiana'' pupa Eumarozia malachitana adult.jpg|''Eumarozia malachitana'' Pandemis limitata caterpillar.jpg|''Pandemis limitata'' larva File:Pandemis limitata pupa.jpg|''Pandemis limitata'' pupa Pandemis limitata adult.jpg|''Pandemis limitata'' adult </gallery>
==See also== * List of Tortricidae genera
==References== {{Reflist}}
==Further reading== * ''Firefly Encyclopedia of Insects and Spiders'', edited by Christopher O'Toole, {{ISBN|1-55297-612-2}}, 2002
==External links== * [http://www.tortricidae.com Tortricid.net] *{{cite web |url=http://insecta.pro/family/tortricidae/ |title=Family Tortricidae |website=Insecta.pro |access-date=February 7, 2020}} * [https://web.archive.org/web/20060207062552/http://nlbif.eti.uva.nl/bis/tortricidae.php Eurasian Tortricidae] * ''Tortricid Fauna of Apple in New York.(Lepidoptera: Tortricidae): Including an Account of Apples' Occurrence in the State, Especially as a Naturalized Plant''. by P. J. Chapman and S. E. Link, Geneva: New York State Agricultural Experiment Station, 1971 * [http://entomology.ifas.ufl.edu/creatures/trees/moths/nantucket_pine_tip_moth.htm ''Rhyacionia frustrana'', Nantucket pine tip moth] on the UF / IFAS Featured Creatures Web site
<br/> {{Lepidoptera}} {{Taxonbar |from=Q28953}} {{Authority control}}
Category:Tortricidae Category:Moth families Category:Taxa named by Pierre André Latreille Category:Extant Eocene first appearances