{{Short description|Family of aquatic microorganisms}} {{Automatic taxobox | image = 12862_2018_1224_Fig1i.jpg | image_caption = ''Collodictyon triciliatum'' | taxon = Diphylleidae | authority = Cavalier-Smith 1993 | subdivision_ranks = Genera | subdivision = * ''Collodictyon'' * ''Diphylleia'' | synonyms = "Collodictyonidae" <small>Brugerolle et al. 2002<ref>{{Cite journal |last=Cavalier-Smith |first=Thomas |date=2022-05-01 |title=Ciliary transition zone evolution and the root of the eukaryote tree: implications for opisthokont origin and classification of kingdoms Protozoa, Plantae, and Fungi |url=https://doi.org/10.1007/s00709-021-01665-7 |journal=Protoplasma |language=en |volume=259 |issue=3 |pages=487–593 |doi=10.1007/s00709-021-01665-7 |issn=1615-6102 |pmc=9010356 |pmid=34940909 |doi-access=free}}</ref></small> }}

'''Diphylleidae''' (also '''Collodictyonidae''') is a group of aquatic, unicellular eukaryotic organisms with two to four terminal flagella. They feed by phagocytosis, ingesting other unicellular organisms like algae and bacteria. It is the sister group of Sulcomonadidae.<ref>{{Cite journal |last=Cavalier-Smith |first=Thomas |date=2013-05-01 |title=Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa |url=https://www.sciencedirect.com/science/article/pii/S0932473912000508 |journal=European Journal of Protistology |volume=49 |issue=2 |pages=115–178 |doi=10.1016/j.ejop.2012.06.001 |issn=0932-4739|url-access=subscription }}</ref>

Recent molecular analyses place collodictyonids (e.g. ''Collodictyon'') in a clade also containing Rigifilida and Mantamonadidae. This clade has been named CRuMs and is sister to Amorphea.<ref>{{Cite journal|last1=Lax|first1=Gordon|last2=Eglit|first2=Yana|last3=Eme|first3=Laura|last4=Bertrand|first4=Erin M.|last5=Roger|first5=Andrew J.|last6=Simpson|first6=Alastair G. B.|date=2018-11-14|title=Hemimastigophora is a novel supra-kingdom-level lineage of eukaryotes|journal=Nature|volume=564|issue=7736|pages=410–414|language=En|doi=10.1038/s41586-018-0708-8|pmid=30429611|bibcode=2018Natur.564..410L |s2cid=205570993|issn=0028-0836}}</ref><ref name="Brown_2018">{{Cite journal|last1=Brown|first1=Matthew W|last2=Heiss|first2=Aaron A|last3=Kamikawa|first3=Ryoma|last4=Inagaki|first4=Yuji|last5=Yabuki|first5=Akinori|last6=Tice|first6=Alexander K|last7=Shiratori|first7=Takashi|last8=Ishida|first8=Ken-Ichiro|last9=Hashimoto|first9=Tetsuo|date=2018-01-19|title=Phylogenomics Places Orphan Protistan Lineages in a Novel Eukaryotic Super-Group|journal=Genome Biology and Evolution|language=en|volume=10|issue=2|pages=427–433|doi=10.1093/gbe/evy014|issn=1759-6653|pmc=5793813|pmid=29360967}}</ref>

==Phylogeny== {{clade| style=font-size:90%;line-height:80% |label1=Opimoda |1={{Clade |1=Ancyromonadida <small>Cavalier-Smith 1998 emend. Atkins 2000</small> |2={{Clade |1=Malawimonadea <small>Cavalier-Smith 2003</small> |label2=Podiata |2={{Clade |label1=Amorphea |1={{Clade |1=Amoebozoa <small>Lühe 1913 emend. Cavalier-Smith 1998</small> |label2=Obazoa |2={{Clade |1=Breviatea <small>Cavalier-Smith 2004</small> |2={{Clade |1=Apusomonadida <small>Karpov & Mylnikov 1989</small> |2=Opisthokonta }} }} }} |label2=CRuMs<ref name="Brown_2018"/> |2={{Clade |1=Mantamonadida <small>Cavalier-Smith 2004</small> |2={{Clade |1=Glissandrida <small>Yazaki ''et al.'' 2025</small><ref name="Yazaki 2025">{{Cite journal |last1=Yazaki |first1=Euki |last2=Harada |first2=Ryo |last3=Isogai |first3=Ryu |last4=Bamba |first4=Kohei |last5=Ishida |first5=Ken-ichiro |last6=Inagaki |first6=Yuji |last7=Shiratori |first7=Takashi |date=2025-06-04 |title=Glissandra oviformis n. sp.: a novel predatory flagellate illuminates the character evolution within the eukaryotic clade CRuMs |journal=Open Biology |volume=15 |issue=6 |article-number=250057 |doi=10.1098/rsob.250057 |doi-access=free|pmc=12133344 |pmid=40460873}}</ref> |2={{Clade |1=Rigifilida <small>Karpov & Mylnikov 1989</small> |2={{clade |label1=Diphylleida |1={{clade |1='''Diphylleidae''' <small>Cavalier-Smith 1993</small> |2=Sulcomonadidae <small>Cavalier-Smith 2013</small> }} }} }} }} }} }} }} }} }}

==Taxonomy== * '''Family Diphylleidae''' <small>Cavalier-Smith 1993</small> ["Collodictyonidae" <small>Brugerolle et al. 2002</small>] ** Genus ''Diphylleia'' <small>Massart 1920 non Michaux 1803</small> [''Aulacomonas'' <small>Skuja 1939</small>] *** Species ''D. rotans'' <small>Massart 1920</small> [''Aulacomonas submarina'' <small>Skuja 1939</small>] ** Genus ''Collodictyon'' <small>Carter 1865</small> *** Species ''C. hongkongense'' <small>Skvortzov 1968</small> *** Species ''C. indicum'' <small>Iyengar 1981</small> *** Species ''C. oxycareni'' <small>Franchini 1922</small> *** Species ''C. sparsevacuolatum'' <small>Skuja 1956</small> *** Species ''C. triciliatum'' <small>Carter 1865</small> == References == {{Reflist}}

{{Eukaryota}} {{Taxonbar|from1=Q11773610|from2=Q136687984}}

Category:CRuMs families

{{Eukaryote-stub}}