Dyakiidae

{{Short description|Family of gastropods}} {{Automatic taxobox |image = Dyakia_salangana_01.JPG |image_caption = Five views of a shell of ''Dyakia salangana'' |taxon = Dyakiidae |authority = Gude & B. B. Woodward, 1921<ref name="Gude 1921">Gude G. P. L. K. & Woodward B. B. (1921). "On ''Helicella'', Férussac". ''Proceedings of the Malacological Society of London'' '''14'''(5-6): [https://archive.org/stream/proceedingsofmal1415192023mala#page/n213/mode/2up 174]-190. [https://archive.org/stream/proceedingsofmal1415192023mala#page/n223/mode/2up page 185].</ref> |synonyms_ref = |synonyms = |display_parents= 3 }}

'''Dyakiidae''' is a family of air-breathing land snails terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).<ref name="Bouchet 2005">{{Bouchet 2005}}</ref>

Some of the species in this family are sinistral (left-handed) in their shell coiling.

== Distribution == The family Dyakiidae is endemic to Sundaland in Southeast Asia.<ref>{{Cite journal | last1 = Hausdorf | first1 = B. | title = A Preliminary Phylogenetic and Biogeographic Analysis of the Dyakiidae (Gastropoda: Stylommatophora) and a Biogeographic Analysis of Other Sundaland Taxa | doi = 10.1111/j.1096-0031.1995.tb00095.x | journal = Cladistics | volume = 11 | issue = 4 | pages = 359–376 | year = 1995 | doi-access = free | pmid = 34920646 }}.</ref>

==Anatomy== In this family, the number of haploid chromosomes is between 26 and 30 (according to the values in this table).<ref name="Barker">Barker G. M. (2001) ''Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology''. 1-146. In: Barker G. M. (ed.) (2001) ''[https://books.google.com/books?id=WlvX-9Wt0toC&hl The biology of terrestrial molluscs]''. CABI Publishing, Oxon, UK, cited pages: 139-144. {{ISBN|0-85199-318-4}}.</ref>

The family also includes the only known terrestrial gastropods to emit light.<ref>{{Cite journal | last1 = Isobe | first1 = M. | last2 = Uyakul | first2 = D. | last3 = Goto | first3 = T. | last4 = Counsilman | first4 = J. J. | title = Dyakia bioluminescence—1. Bioluminescence and fluorescence spectra of the land snail, D. Striata | doi = 10.1002/bio.1170020204 | journal = Journal of Bioluminescence and Chemiluminescence | volume = 2 | issue = 2 | pages = 73–79 | year = 1988 | pmid = 3213594 }}.</ref><ref>{{Cite journal |last1=Pholyotha |first1=Arthit |last2=Yano |first2=Daichi |last3=Mizuno |first3=Gaku |last4=Sutcharit |first4=Chirasak |last5=Tongkerd |first5=Piyoros |last6=Oba |first6=Yuichi |last7=Panha |first7=Somsak |date=2023-09-13 |title=A new discovery of the bioluminescent terrestrial snail genus Phuphania (Gastropoda: Dyakiidae) |journal=Scientific Reports |language=en |volume=13 |issue=1 |pages=15137 |doi=10.1038/s41598-023-42364-y |issn=2045-2322 |pmc=10499882 |pmid=37704646|bibcode=2023NatSR..1315137P }}</ref>

The Digestive system characteristics are as follows. The buccal mass is small. The jaw is smooth. The stomach is very simple with weak muscles (as is the case in the majority of land snails).<ref name="Barker"/> (These anatomical characteristics also include the family Staffordiidae which was considered part of the Dyakiidae at the time the study was done).

== Genera == The family Dyakiidae includes the following genera, with a steadily increasing number of described species (71 species until 2007<ref name="Tumpeesuwan"/> + 11 new species in 2009<ref name="Liew 2009">{{Cite journal | last1 = Liew | first1 = T. S. | last2 = Schilthuizen | first2 = M. | last3 = Vermeulen | first3 = J. J. N. | doi = 10.1111/j.1096-3642.2009.00526.x | title = Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo | journal = Zoological Journal of the Linnean Society | volume = 157 | issue = 3 | pages = 515 | year = 2009 | doi-access = free }}.</ref>):

# ''Asperitas'' Gray, 1857 - 14 species # ''Bertia'' Ancey, 1887 - at least 4 species # ''Dyakia'' Godwin-Austen, 1891 - type genus, 22 species # ''Elaphroconcha'' Gude, 1911 - 10 species # ''Everettia'' Godwin-Austen, 1891<ref>Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". ''Proceedings of the Zoological Society of London'' [https://archive.org/details/proceedings03londgoog '''for the year 1891''']: [https://archive.org/stream/proceedings03londgoog#page/n50/mode/1up 22]-47, p. [https://archive.org/stream/proceedings03londgoog#page/n69/mode/1up 33].</ref> - 25 species # ''Kalamantania'' Laidlaw, 1931 - only one species: ''Kalamantania whiteheadi'' (Godwin-Austen, 1891)<ref>Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". ''Proceedings of the Zoological Society of London'' [https://archive.org/details/proceedings03londgoog '''for the year 1891''']: [https://archive.org/stream/proceedings03londgoog#page/n50/mode/1up 22]-47, p. [https://archive.org/stream/proceedings03londgoog#page/n60/mode/1up 24].</ref> # ''Phuphania'' Tumpeesuwan, Naggs & Panha, 2007 - 3 species including: ''Phuphania globosa'' Tumpeesuwan, Naggs & Panha, 2007<ref name="Tumpeesuwan">Tumpeesuwan C., Naggs F. & Panha S. (31 August 2007) "A new genus and new species of dyakiid snail (Pulmonata: Dyakiidae) from the Phu Phan range, northeastern Thailand". ''Raffles Bulletin of Zoology'' [http://rmbr.nus.edu.sg/rbz/journal552.php '''55'''(2)] {{webarchive |url=https://web.archive.org/web/20090722022417/http://rmbr.nus.edu.sg/rbz/journal552.php |date=July 22, 2009 }}: 363-369. [http://rmbr.nus.edu.sg/rbz/biblio/55/55rbz363-369.pdf PDF]</ref> # ''Pseudoplecta'' Laidlaw, 1932 - only one species: ''Pseudoplecta bijuga'' (Stoliczka, 1873) # ''Pseudoquantula'' Jirapatrasilp & Panha gen. nov. - only one species: ''Pseudoquantula lenticularis'' Jirapatrasilp & Panha ''sp. nov.''<ref>{{cite journal|author1=Parin Jirapatrasilp|author2=Piyoros Tongkerd|author3=Ekgachai Jeratthitikul|author4=Thor-Seng Liew|author5=Arthit Pholyotha|author6=Chirasak Sutcharit|author7=Somsak Panha|title=Molecular phylogeny of the limacoid snail family Dyakiidae in Southeast Asia, with the description of a new genus and species|journal=Zoological Journal of the Linnean Society|volume=193|issue=1|year= 2021| pages=250–280| url= https://doi.org/10.1093/zoolinnean/zlaa129|doi=10.1093/zoolinnean/zlaa129|url-access=subscription}}</ref> # ''Quantula'' Baker, 1941 - 7 species including: ''Quantula striata'' (Gray, 1834) # ''Rhinocochlis'' Thiele, 1931 - only one species: ''Rhinocochlis nasuta'' (Metcalfe, 1852) # ''Sasakina'' Rensch, 1930 - 5 species

== Cladogram == The following cladogram shows the phylogenic relationships of this family and superfamily with the other families within the limacoid clade:<ref>{{Cite journal | last1 = Hausdorf | first1 = B. | title = Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance events and long-distance dispersal | doi = 10.1046/j.1365-2699.2000.00403.x | journal = Journal of Biogeography | volume = 27 | issue = 2 | pages = 379–390 | year = 2000 | jstor=2656267| bibcode = 2000JBiog..27..379H }}.</ref>

{{clade |label1=limacoid clade |1={{clade |label1=Staffordioidea |1=Staffordiidae |2={{clade |1={{clade |label1='''Dyakioidea''' |1='''Dyakiidae''' |label2=Gastrodontoidea |2={{clade |1=Pristilomatidae |2={{clade |1=Chronidae |2={{clade |1=Euconulidae |2=Trochomorphidae }} |3={{clade |1=Gastrodontidae |2=Oxychilidae }} }} }} }} |2={{clade |label1=Parmacelloidea |1={{clade |1=Trigonochlamydidae |2=Parmacellidae |3=Milacidae }} |2={{clade |1={{clade |label1=Zonitoidea |1=Zonitidae |label2=Helicarionoidea |2={{clade |1=Helicarionidae |2={{clade |1=Ariophantidae |2=Urocyclidae }} }} }} |label2=Limacoidea |2={{clade |1=Vitrinidae |2={{clade |1=Boettgerillidae |2={{clade |1=Limacidae |2=Agriolimacidae }} }} }} }} }} }} }} }}

==References== {{Reflist}}

== Further reading == * Laidlaw F. F. (1931). "On a new sub-family Dyakiinae of the Zonitidae". ''Proceedings of the malacological Society of London'' '''19''': 190-201. [https://archive.today/20120716065513/http://mollus.oxfordjournals.org/cgi/pdf_extract/19/4/190 abstract]. * Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". ''Ruthenica'', Supplement 2. 1309-1466. * Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

== External links == * [https://biodiversitylibrary.org/page/30346901 Gude, G. K. & Woodward, B. B. (1921). On Helicella, Férussac. Proceedings of the Malacological Society of London. 14 (5/6): 174-190. Londo]

Category:Dyakiidae Category:Taxa named by Gerard Kalshoven Gude