{{Short description|Order of microorganisms}} {{Automatic taxobox | image = | image_alt = | image_caption = | taxon = Rigifilida | authority = Cavalier-Smith in Yabuki ''et al.'', 2012 | parent_authority = Cavalier-Smith 2008, emend. 2012<ref>{{Cite journal|last=Cavalier-Smith |first=Thomas |title=Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa |journal=European Journal of Protistology|volume=49 |issue=2 |pages=115–178 |doi=10.1016/j.ejop.2012.06.001|pmid=23085100 |year=2013 }}</ref> | synonyms = | synonyms_ref = | subdivision_ranks = Families | subdivision = * Micronucleariidae * Rigifilidae }}
'''Rigifilida''' is a clade of non-ciliate phagotrophic eukaryotes. It consists of two genera: ''Micronuclearia'' and ''Rigifila''.
==Characteristics== Cells of rigifilids are covered with either a single or a double-layered submembrane pellicular lamina that makes them rigid in consistence. Slender branching filopodia emanate from a ventral aperture of the cell and are employed to collect bacteria upon which they feed and to attach the organism to the substratum. Around this aperture, the pellicle is reflexed around forming a peristomial collar. Other notable features are flat and irregular shaped mitocondrial cristae, a single dorsal nucleus and the lack of centrioles and cilia.<ref name="Yabuki 2012">{{cite journal|vauthors=Yabuki A, Ishida KI, Cavalier-Smith T|date=2013|title=Rigifila ramosa n. gen., n. sp., a Filose Apusozoan with a Distinctive Pellicle, is Related to Micronuclearia|journal=Protist|volume=164|pages=75–88|doi=10.1016/j.protis.2012.04.005|issue=1}}</ref>
==Phylogeny== {{clade| style=font-size:90%;line-height:80% |label1=Opimoda |1={{Clade |1=Ancyromonadida <small>Cavalier-Smith 1998 emend. Atkins 2000</small> |2={{Clade |1=Malawimonadea <small>Cavalier-Smith 2003</small> |2={{Clade |label1=Amorphea |1={{Clade |1=Amoebozoa <small>Lühe 1913 emend. Cavalier-Smith 1998</small> |label2=Obazoa |2={{Clade |1=Breviatea <small>Cavalier-Smith 2004</small> |2={{Clade |1=Apusomonadida <small>Karpov & Mylnikov 1989</small> |2=Opisthokonta }} }} }} |label2=CRuMs<ref name="Brown_2018">{{Cite journal |last1=Brown |first1=Matthew W. |last2=Heiss |first2=Aaron A |last3=Kamikawa |first3=Ryoma |last4=Inagaki |first4=Yuji |last5=Yabuki |first5=Akinori |last6=Tice |first6=Alexander K. |last7=Shiratori |first7=Takashi |last8=Ishida |first8=Ken-Ichiro |last9=Hashimoto |first9=Tetsuo |date=2018-01-19 |title=Phylogenomics Places Orphan Protistan Lineages in a Novel Eukaryotic Super-Group |journal=Genome Biology and Evolution |volume=10 |issue=2 |pages=427–433 |doi=10.1093/gbe/evy014 |issn=1759-6653 |pmc=5793813 |pmid=29360967}}</ref> |2={{Clade |1=Mantamonadida <small>Cavalier-Smith 2004</small> |2={{Clade |1=Glissandrida <small>Yazaki ''et al.'' 2025</small><ref name="Yazaki 2025">{{Cite journal |last1=Yazaki |first1=Euki |last2=Harada |first2=Ryo |last3=Isogai |first3=Ryu |last4=Bamba |first4=Kohei |last5=Ishida |first5=Ken-ichiro |last6=Inagaki |first6=Yuji |last7=Shiratori |first7=Takashi |date=2025-06-04 |title=Glissandra oviformis n. sp.: a novel predatory flagellate illuminates the character evolution within the eukaryotic clade CRuMs |journal=Open Biology |volume=15 |issue=6 |article-number=250057 |doi=10.1098/rsob.250057 |doi-access=free|pmc=12133344 |pmid=40460873}}</ref> |2={{Clade |1='''Rigifilida''' <small>Karpov & Mylnikov 1989</small> |2=Diphylleida <small>Cavalier-Smith 1993</small> }} }} }} }} }} }} }}
==Taxonomy==
Rigifilida is currently placed in CRuMs.<ref>{{Cite journal|last1=Adl|first1=Sina M.|last2=Bass|first2=David|last3=Lane|first3=Christopher E.|last4=Lukeš|first4=Julius|last5=Schoch|first5=Conrad L.|last6=Smirnov|first6=Alexey|last7=Agatha|first7=Sabine|last8=Berney|first8=Cedric|last9=Brown|first9=Matthew W.|date=2018-09-26|title=Revisions to the Classification, Nomenclature, and Diversity of Eukaryotes|journal=Journal of Eukaryotic Microbiology|volume=66|issue=1|pages=4–119|language=en|doi=10.1111/jeu.12691|pmid=30257078|pmc=6492006|issn=1066-5234}}</ref>
*'''Order Rigifilida''' <small>Cavalier-Smith 2012</small> [Micronucleariida <small>Cavalier-Smith 2008</small>] ** Family Rigifilidae <small>Yabuki & Cavalier-Smith 2012</small> *** Genus ''Rigifila'' <small>Yabuki & Cavalier-Smith 2012</small> **** Species ''Rigifila ramosa'' <small>Yabuki & Cavalier-Smith 2012</small> ** Family Micronucleariidae <small>Cavalier-Smith 2008</small> *** Genus ''Micronuclearia'' <small>Mikrjukov & Mylnikov 2001</small> **** Species ''Micronuclearia podoventralis'' <small>Mikrjukov & Mylnikov 2001 </small>
== References == {{Reflist}}
{{Eukaryota}} {{Taxonbar|from=Q16989278}}
Category:CRuMs orders
{{Eukaryote-stub}}