{{Short description|Extinct family of reptiles}} {{Automatic taxobox | taxon = Baurusuchidae | name = Baurusuchids | fossil_range = Santonian – Danian,<ref>{{Cite journal|last1=Piacentini Pinheiro|first1=André E.|last2=Pereira|first2=Paulo Victor Luiz Gomes da Costa|last3=de Souza|first3=Rafael G.|last4=Brum|first4=Arthur S.|last5=Lopes|first5=Ricardo T.|last6=Machado|first6=Alessandra S.|last7=Bergqvist|first7=Lílian P.|last8=Simbras|first8=Felipe M.|date=2018-08-01|editor-last=Piras|editor-first=Paolo|title=Reassessment of the enigmatic crocodyliform ''"Goniopholis" paulistanus'' Roxo, 1936: Historical approach, systematic, and description by new materials|journal=PLOS ONE|language=en|volume=13|issue=8|article-number=e0199984|doi=10.1371/journal.pone.0199984|pmid=30067779|pmc=6070184|bibcode=2018PLoSO..1399984P |issn=1932-6203|doi-access=free}}</ref> <br/> {{Fossil range|86|62}} | image = Baurusuchus salgadoensis (MPMA) 1.jpg | image_caption = Skull of ''Baurusuchus salgadoensis'' | parent_authority = Warren, 1968 | authority = Price, 1945 | subdivision_ranks = Genera | subdivision = * {{extinct}}''Chimaerasuchus''? * {{extinct}}''Comahuesuchus''? * {{extinct}}''Cynodontosuchus'' * {{extinct}}''Eremosuchus''? * {{extinct}}''Gondwanasuchus'' * {{extinct}}''Pehuenchesuchus'' * {{extinct}}''Razanandrongobe''? * {{extinct}}'''Baurusuchinae''' ** {{extinct}}''Aphaurosuchus'' ** {{extinct}}''Aplestosuchus'' ** {{extinct}}''Baurusuchus'' ** {{extinct}}''Pabwehshi'' ** {{extinct}}''Stratiotosuchus'' * {{extinct}}'''Pissarrachampsinae''' ** {{extinct}}''Campinasuchus'' ** {{extinct}}''Pissarrachampsa'' ** {{extinct}}''Wargosuchus'' }}
'''Baurusuchidae''' is a Gondwanan family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America (Argentina and Brazil) and Asia (Pakistan). Baurusuchidae has been, in accordance with the PhyloCode, officially defined as the least inclusive clade containing ''Cynodontosuchus rothi, Pissarrachampsa sera,'' and ''Baurusuchus pachecoi.''<ref name=":0">{{Cite journal|last1=Darlim|first1=Gustavo|last2=Montefeltro|first2=Felipe C.|last3=Langer|first3=Max C.|date=September 2021|title=3D skull modelling and description of a new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous (Bauru Basin) of Brazil|url=https://www.researchgate.net/publication/350966965|journal=Journal of Anatomy|language=en|volume=239|issue=3|pages=622–662|doi=10.1111/joa.13442|issn=0021-8782|pmc=8349455|pmid=33870512}}</ref> Baurusuchids have been placed in the suborder '''Baurusuchia''', and two subfamilies have been proposed: Baurusuchinae and Pissarrachampsinae.
==Genera== Several genera have been assigned to Baurusuchidae. ''Baurusuchus'' was the first, being the namesake of the family. Remains of ''Baurusuchus'' have been found from the Late Cretaceous Bauru Group of Brazil in deposits that are Turonian - Santonian in age.<ref name=CMAR06>{{cite journal |last=Candeiro |first=C.R.A. |author2=Martinelli, A.G. |author3=Avilla, L.S. |author4= Rich, T.H. |year=2006 |title=Tetrapods from the Upper Cretaceous (Turonian-Maastrichtian) Bauru Group of Brazil: a reappraisal |journal=Cretaceous Research |volume=27 |issue=6 |pages=923–946 |doi=10.1016/j.cretres.2006.05.002|bibcode=2006CrRes..27..923C |url=http://doc.rero.ch/record/16221/files/PAL_E3443.pdf }}</ref> In addition to ''Baurusuchus'', five other South American crocodyliforms have been assigned to Baurusuchidae: ''Campinasuchus'', ''Cynodontosuchus'', ''Pissarrachampsa'', ''Stratiotosuchus'', and ''Wargosuchus''. ''Cynodontosuchus'' was the first known baurusuchid, named in 1896 by English paleontologist Arthur Smith Woodward, although it was only recently assigned to Baurusuchidae.<ref name=WAS96>{{cite journal |last=Woodward |first=A.S. |year=1896 |title=. On two Mesozoic crocodilians, ''Notosuchus (genus novum)'' and ''Cynodontosuchus (genus novum)'' from the red sandstones of the Territory of Neuquén (Argentine Republic) |journal=Anales del Museo de la Plata (Paleontología) |volume=4 |pages=1–20}}</ref> ''Wargosuchus'' was described in 2008.<ref name=MP08>{{cite journal |last=Martinelli |first=A.G. |author2=Pais, D.F. |year=2008 |title=A new baurusuchid crocodyliform (Archosauria) from the Late Cretaceous of Patagonia (Argentina) |journal=Comptes Rendus Palevol |volume=7 |issue=6 |pages= 371–381 |doi=10.1016/j.crpv.2008.05.002|bibcode=2008CRPal...7..371M }}</ref> ''Cynodontosuchus'' and ''Wargosuchus'' are known only from fragmentary remains. Both genera are from the Santonian of Argentina.
[[File:Pissarrachampsa sera.png|thumb|right|Skull of ''Pissarrachampsa''.]] A fourth genus, ''Stratiotosuchus'', was assigned to Baurusuchidae in 2001. ''Pabwehshi'' is the youngest genus that has been assigned to Baurusuchidae, and is from the Maastrichtian of Pakistan.<ref name=WMG01>{{cite journal |last=Wilson |first=J.A. |author2=Malkane, M.S. |author3= Gingerich, P.D. |year=2001 |title=New crocodyliform (Reptilia, Mesoeucrocodylia) from the Upper Cretaceous Pab Formation of Vitakri, Balochistan (Pakistan) |journal=Contributions from the Museum of Paleontology, University of Michigan |volume=30 |issue=12 |pages=321–336}}</ref> It was named in 2001 but has since been reassigned as a basal member of Sebecia.<ref name=LAS07/>
A new genus, ''Campinasuchus'', was assigned to the family in May, 2011. It is known from the Turonian-Santonian Adamantina Formation of the Bauru Basin of Brazil.<ref name=Cetal11>{{cite journal |last=Carvalho |first=I.S. |year=2011 |title=''Campinasuchus dinizi'' gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil |journal=Zootaxa |volume=2871 |issue=2871 |pages=19–42. [http://www.mapress.com/zootaxa/2011/f/z02871p042f.pdf Preview] [http://www.mapress.com/zootaxa/2011/f/zt02871p042.pdf PDF]|doi=10.11646/zootaxa.2871.1.2 |display-authors=etal|doi-access=free }}</ref> Soon after, the new genus ''Pissarrachampsa'' was named from the Campanian–Maastrichtian Vale do Rio do Peixe Formation, also in the Bauru Basin.
==Phylogeny== The family Baurusuchidae was named by Brazilian paleontologist Llewellyn Ivor Price in 1945 to include ''Baurusuchus''.<ref name=PLI45>{{cite journal |last=Price |first=L.I. |year=1945 |title=A new reptile from the Cretaceous of Brazil |journal=Rio de Janeiro, Departamento Nacional da Produção Mineral, Notas preliminares e estudos |volume=25 |pages=1–8}}</ref> In 1946, American paleontologist Edwin Harris Colbert erected the group Sebecosuchia, which united Baurusuchidae with the family Sebecidae (represented by the genus ''Sebecus'').<ref name=CEH46>{{cite journal |last=Colbert |first=E.H. |year=1946 |title=''Sebecus'', representative of a peculiar suborder of fossil Crocodilia from Patagonia |journal=Bulletin of the American Museum of Natural History |volume=87 |pages=217–270}}</ref> Both ''Baurusuchus'' and ''Sebecus'' have deep snouts and ziphodont dentitions (teeth that are serrated and laterally compressed).<ref name=CCH05>{{cite journal |last=Carvalho |first=I.S. |author2=Campos, A.C.A. |author3=Nobre, P.H. |year=2005 |title=''Baurusuchus salgadoensis'', a new Crocodylomorpha from the Bauru Basin (Cretaceous), Brazil |journal=Gondwana Research |volume=8 |issue=1 |pages=11–30 |issn=1342-937X |url=http://www.igeo.ufrj.br/ismar/3/3_14.pdf |doi=10.1016/S1342-937X(05)70259-8 |bibcode=2005GondR...8...11C |access-date=2010-12-26 |archive-url=https://web.archive.org/web/20110706162337/http://www.igeo.ufrj.br/ismar/3/3_14.pdf |archive-date=2011-07-06 }}</ref> Other forms were later found that had a close appearance to these two genera, among them ''Cynodontosuchus'', ''Stratiotosuchus'', and ''Wargosuchus''. Several features were used to unite these groups: a deep snout, a ziphodont dentition, a curved tooth row, an enlarged canine-like dentary tooth that fits into a deep notch in the upper jaw, and a groove on the lower jaw.<ref name=MLL11>{{cite journal |last=Montefeltro |first=F.C. |author2=Larsson, H.C.E. |author3= Langer, M.C. |year=2011 |title=A new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous of Brazil and the phylogeny of Baurusuchidae|journal=PLOS ONE |volume=6 |issue=7 |article-number= e21916 |doi=10.1371/journal.pone.0021916 |pmid=21765925 |pmc=3135595|bibcode=2011PLoSO...621916M |doi-access=free }}</ref>
Many phylogenetic analyses within the past decades have supported a close relationship between the two families.<ref name=CCH05/> Baurusuchids and sebecosuchids are both early members of the clade Metasuchia, which includes the subgroups Notosuchia (mainly terrestrial crocodyliforms) and Neosuchia (larger, often semiaquatic crocodyliforms, including living crocodylians). Sebecosuchians, which include both baurusuchids and sebecosuchids, were found to be closely related to notosuchians in several studies.<ref name=GCF91>{{cite journal |last=Gasparini |first=Z.B. |author2=Chiappe, L.M. |author3= Fernandez, M. |year=1991 |title=A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a sinopsis of the South American Cretaceous crocodilians |journal=Journal of Vertebrate Paleontology |volume=11 |issue=3 |pages=316–333 |doi=10.1080/02724634.1991.10011401|bibcode=1991JVPal..11..316G }}</ref> The new genera ''Iberosuchus'' and ''Eremosuchus'' were later assigned to Baurusuchidae, and phylogenetic analyses encompassing these taxa continued to find Baurusuchidae to be closely related to Sebecidae.<ref name=Oetal1996>{{cite journal |last=Ortega |first=F. |author2=Buscaloni, A.D |author3= Gasaparini, Z. |year=1996 |title=Reinterpretation and new denomination of ''Atacisaurus crassiproratus'' (Middle Eocene; Issel, France) as cf. ''Iberosuchus'' (Crocodylomorpha, Metasuchia) |journal=Geobios |volume=29 |issue=3 |pages=353–364 |doi=10.1016/S0016-6995(96)80037-4 |bibcode=1996Geobi..29..353O }}</ref> Both families were allied with notosuchians in the larger group Ziphosuchia, composed of ziphodont crocodyliforms. More recently, sebecosuchians - including baurusuchids - have been placed within Notosuchia as derived members of the clade. Below is a modified cladogram from Ortega ''et al.'' (2000) placing baurusuchids within Notosuchia:<ref name=OBC00>{{cite journal |last=Ortega |first=F. Z. |author2=Buscalioni, A. D. |author3= Calvo, J. O. |year=2000 |title=A new species of ''Araripesuchus'' (Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina) |journal=Journal of Vertebrate Paleontology |volume=20 |issue=1 |pages=57–76 |doi=10.1671/0272-4634(2000)020[0057:ANSOAC]2.0.CO;2|s2cid=85946263 }}</ref>
{{clade| style=font-size:90%;line-height:80% |label1=Notosuchia |1={{clade |1=''Notosuchus'' |2={{clade |1=''Libycosuchus'' |2={{clade |1=''{{bg|#E3F5FF|'''''Baurusuchus'''''}}'' |2={{clade |1=''{{bg|#E3F5FF|'''''Iberosuchus'''''}}'' |2={{clade |1=''Sebecus'' |2={{clade |1=Itaborai crocodile |2=''Bretesuchus'' }} }} }} }} }} }} {{bg|#E3F5FF| }} Baurusuchids}}
In 2004, the superfamily Baurusuchoidea was established to include baurusuchids and sebecids. Phylogenetically, Baurusuchoidea was defined as the most recent common ancestor of ''Baurusuchus'' and ''Sebecus'' and all of its descendants while Baurusuchidae was defined as the most recent common ancestor of ''Baurusuchus'' and ''Stratiotosuchus'' and all of its descendants.<ref name=Cetal04>{{cite journal |last=Carvalho |first=I.S. |author2=Ribeiro, L.C.B. |author3=Avilla, L.S. |year=2004 |title=''Uberabasuchus terrificus sp. nov.'', a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil |journal=Gondwana Research |volume=7 |issue=4 |pages=975–1002 |issn=1342-937X |url=http://acd.ufrj.br/ismar/3/3_15.pdf |doi=10.1016/S1342-937X(05)71079-0 |bibcode=2004GondR...7..975C |archive-url=https://web.archive.org/web/20110706162632/http://acd.ufrj.br/ismar/3/3_15.pdf |archive-date=2011-07-06 }}</ref>
In a 2005 analysis, Sebecidae was found to be a paraphyletic grouping, or a grouping that includes some descendants of a common ancestor but not all. Sebecids formed an assemblage of basal sebecosuchians, while baurusuchids remained a valid grouping of derived sebecosuchians. Below is a modified cladogram from Turner and Calvo (2005):<ref name=TC05>{{cite journal |last=Turner |first=A.H. |author2=Calvo, J.O. |year=2005 |title=A new sebecosuchian crocodyliform from the Late Cretaceous of Patagonia |journal=Journal of Vertebrate Paleontology |volume=25 |issue=1 |pages=87–98 |doi=10.1671/0272-4634(2005)025[0087:ANSCFT]2.0.CO;2|s2cid=86257810 }}</ref>
{{clade| style=font-size:90%;line-height:80% |label1=Metasuchia |1={{clade |1=Neosuchia |2=''Libycosaurus'' |3={{clade |1=Notosuchia |label2=Sebecosuchia |2={{clade |1=''Pehuenchesuchus'' |2={{clade |1=''Eremosuchus'' |2=''Sebecus'' |3={{clade |1=''Iberosuchus'' |2=''Bretesuchus'' |label3='''Baurusuchidae''' |3={{clade |1=''Baurusuchus'' |2=''Pabwehshi'' }} }} }} }} }} }} }}
Later studies noted many features that distinguished baurusuchids from sebecosuchids. Sebecosuchids were often considered to be more closely related to Neosuchia, a group that includes modern crocodylians, while baurusuchids were thought to be a more distantly related clade.<ref name=CJM97>{{cite book |last=Clark |first=J.M. |year=1997 |chapter=Patterns of evolution in Mesozoic Crocodyliformes |title=In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods |editor=Fraser, N.C. |editor2=Sues, H.-D. |publisher=Cambridge University Press |location=Cambridge |pages=84–97}}</ref> In a 1999 phylogenetic analysis, ''Baurusuchus'' formed a clade with notosuchians to the exclusion of other ziphosuchians.<ref name=PD99>{{cite journal |last=Pol |first=D. |year=1999 |title=Basal Mesoeucrocodylian relationships: new clues to old conflicts |journal=Journal of Vertebrate Paleontology |volume=19 |issue=3, Supplement |pages=69A|doi=10.1080/02724634.1999.10011202}}</ref> This placement has been upheld by recent analyses, which place ''Baurusuchus'' within Notosuchia.<ref name=SL09>{{cite journal |last=Sereno |first=P. C. |author2=Larsson, H. C. E. |year=2009 |title=Cretaceous crocodyliforms from the Sahara |journal=ZooKeys |issue=28 |pages=1–143 |url=http://pensoftonline.net/zookeys/index.php/journal/article/view/325 |doi=10.3897/zookeys.28.325 |access-date=2010-08-13 |archive-url=https://web.archive.org/web/20100921055501/http://pensoftonline.net/zookeys/index.php/journal/article/view/325 |archive-date=2010-09-21 |doi-access=free }}</ref>
In 2007, a new clade called Sebecia was erected.<ref name=LAS07>{{cite journal |last=Larsson |first=H. C. E. |author2=Sues, H.-D. |year=2007 |title=Cranial osteology and phylogenetic relationships of ''Hamadasuchus rebouli'' (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco |journal=Zoological Journal of the Linnean Society |volume=149 |issue=4 |pages=533–567 |doi=10.1111/j.1096-3642.2007.00271.x|doi-access=free }}</ref> Sebecia included sebecids and peirosaurids. Peirosauridae, a family of small terrestrial crocodyliforms, had often been placed in or near Neosuchia in previous studies. The assignment of sebecids to Sebecia placed the family closer to Neosuchia than Notosuchia. In this study, baurusuchids were split up, with ''Baurusuchus'' placed as a more basal metasuchian and the remaining baurusuchids (''Bretesuchus'' and ''Pabwehshi'') placed as sebecians. Therefore, the family Baurusuchidae was paraphyletic. Below is a modified cladogram from Larsson and Sues (2007):<ref name=LAS07/>
{{clade| style=font-size:90%;line-height:80% |label1=Metasuchia |1={{clade |label1=Notosuchia |1={{clade |1=''Notosuchus'' |2=''Malawisuchus''}} |2={{clade |1=''Araripesuchus'' |2={{clade |1={{clade |1=''{{bg|#E3F5FF|'''''Baurusuchus'''''}}'' |2={{clade |1=Neosuchia |label2=Sebecia |2={{clade |1=''{{bg|#E3F5FF|'''''Pabwehshi'''''}}'' |2={{clade |label1=Sebecidae |1={{clade |1=''Sebecus'' |2=''Bretesuchus''}} |2=Peirosauridae }} }} }} }} }} }} }} {{bg|#E3F5FF| }} Baurusuchids}}
More recent studies have nested ''Baurusuchus'' deep within Notosuchia, just as the larger group Sebecosuchia once was, while the remaining sebecosuchian genera have been placed more distantly in Metasuchia.<ref name=SL09/> A new baurusuchid called ''Pissarrachampsa'' was named in 2011, and a comprehensive phylogenetic analysis of baurusuchids was conducted along with its description. Montefeltro ''et al.'' (2011) found Baurusuchidae to be a monophyletic group with the genera ''Baurusuchus'', ''Cynodontosuchus'', ''Pissarrachampsa'', ''Stratiotosuchus'', and ''Wargosuchus''. They adopted the name Baurusuchia in a phylogenetic sense to distinguish baurusuchids from related crocodyliforms. Baurusuchia was first erected as an infraorder in 1968, but in the 2011 analysis it was found to be in an identical position to Baurusuchidae in the final tree. The only difference between Baurusuchidae and Baurusuchia is that the former is a node-based taxon and the latter is a stem-based taxon. Baurusuchidae is defined as the least inclusive clade containing ''Cynodontosuchus rothi, Pissarrachampsa sera,'' and ''Baurusuchus pachecoi.''<ref name=":0" /> As in all node-based clades, there is a most recent common ancestor; these genera are all of its known descendants. Baurusuchia is formally defined as the most inclusive clade containing ''Baurusuchus pachecoi'' but not ''Sebecus icaeorhinus'', ''Sphagesaurus huenei'', ''Araripesuchus gomesi'', ''Montealtosuchus arrudacamposi'', or ''Crocodylus niloticus.''<ref name=":0" />
In contrast to the node-based Baurusuchidae, the stem-based Baurusuchia does not include a common ancestor and all its descendants, but rather all forms more closely related to a specific baurusuchid than a non-baurusuchid. As a stem-based taxon, Baurusuchia is more inclusive than Baurusuchidae; a new taxon could potentially be placed outside Baurusuchidae because it is not a descendant of the most recent common ancestor of baurusuchids, but would still be a baurusuchian because it is more closely related to baurusuchids than it is to other crocodyliforms. For now, however, Baurusuchidae and Baurusuchia are almost identical in scope, with Baurusuchia also including ''Pabwehshi,'' based on their reference phylogenies.<ref name=MLL11/> Other analyses however, have recovered additional taxa within Baurusuchia outside of Baurusuchidae. (''Pakasuchus'' and ''Comahuesuchus'')<ref>{{Cite journal|last1=Ruiz|first1=Juan V.|last2=Bronzati|first2=Mario|last3=Ferreira|first3=Gabriel S.|last4=Martins|first4=Kawan C.|last5=Queiroz|first5=Marcos V.|last6=Langer|first6=Max C.|last7=Montefeltro|first7=Felipe C.|date=2021-02-16|title=A new species of ''Caipirasuchus'' (Notosuchia, Sphagesauridae) from the Late Cretaceous of Brazil and the evolutionary history of Sphagesauria|url=https://www.researchgate.net/publication/351109083|journal=Journal of Systematic Palaeontology|volume=19|issue=4|pages=265–287|doi=10.1080/14772019.2021.1888815|s2cid=235172623|issn=1477-2019|hdl=11449/207687|hdl-access=free}}</ref>
Montefeltro ''et al.'' (2011) also divided Baurusuchidae into two subfamilies, Pissarrachampsinae and Baurusuchinae. Pissarrachampsinae includes ''Pissarrachampsa'' and ''Wargosuchus'' while Baurusuchinae includes ''Stratiotosuchus'' and ''Baurusuchus''. ''Cynodontosuchus'' is not a member of either of these subfamilies, but the most basal baurusuchid. Many of the unique features that separate ''Cynodontosuchus'' may also be associated with a juvenile individual. The material that ''Cynodontosuchus'' is based on has been suggested to be a juvenile form of ''Wargosuchus'', and the two taxa may be synonymous.<ref name=MLL11/>
Below is a cladogram from Montefeltro ''et al.'' (2011):<ref name=MLL11/>
{{clade| style=font-size:85%;line-height:80% |label1=Notosuchia |1={{clade |1=''Notosuchus'' |2={{clade |1=''Mariliasuchus'' |2={{clade |1=''Armadillosuchus'' |label2='''Baurusuchidae''' |2={{clade |1=''Cynodontosuchus'' |2={{clade |label1=Pissarrachampsinae |1={{clade |1=''Pissarrachampsa'' |2=''Wargosuchus''}} |label2=Baurusuchinae |2={{clade |1=''Stratiotosuchus'' |label2=''Baurusuchus'' |2={{clade |1=''B. albertoi'' |2={{clade |1=''B. salgadoensis'' |2=''B. pachecoi'' }} }} }} }} }} }} }} }} }}
A sixth genus of baurusuchid, ''Campinasuchus'', was named just a few months before ''Pissarrachampsa'', and was not included in the analysis.<ref name=Campinasuchus>{{cite journal |author=Ismar De Souza Carvalho |author2=Vicente De Paula Antunes Teixeira |author3=Mara Lúcia Da Fonseca Ferraz |author4=Luiz Carlos Borges Ribeiro |author5=Agustín Guillermo Martinelli |author6=Francisco Macedo Neto |author7=Joseph J. W. Sertich |author8=Gabriel Cardoso Cunha |author9=Isabella Cardoso Cunha |author10=Patrícia Fonseca Ferraz |name-list-style=amp |year=2011 |title=''Campinasuchus dinizi'' gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil |url=http://www.mapress.com/zootaxa/2011/f/zt02871p042.pdf |journal=Zootaxa |volume=2871 |pages=19–42 |doi=10.11646/zootaxa.2871.1.2 }}</ref>
Darlim ''et al.'' (2021) described a new baurusuchid, ''Aphaurosuchus,'' and proposed formal definitions for the clades Baurusuchia, Baurusuchidae, Baurusuchinae, and Pissarrachampsinae. In addition to this, the study conducted a phylogenetic analysis to resolve the affinites of the new taxon and provide a reference phylogeny for the newly defined clades. The cladogram of this analysis is shown below.<ref name=":0" />
{{Clade|{{clade |1=''Cynodontosuchus rothi'' |2={{clade |1=''Gondwanasuchus scabrosus'' |2={{clade |label1=Pissarrachampsinae |1={{clade |1=''Campinasuchus dinizi'' |2={{clade |1=''Pissarrachampsa sera'' |2=''Wargosuchus australis'' }} }} |label2=Baurusuchinae |2={{clade |1=''Aphaurosuchus escharafacies'' |2={{clade |1=''Aplestosuchus sordidus'' |2={{clade |1=''Stratiotosuchus maxhechti'' |2={{clade |1=''Baurusuchus albertoi'' |2={{clade |1=''Baurusuchus pachecoi'' |2=''Baurusuchus salgadoensis'' }} }} }} }} }} }} }} }}|label1='''Baurusuchidae'''}}
==Paleobiology== In 2011, fossilized eggs were described from the Late Cretaceous Adamantina Formation of Brazil that may have been laid by a baurusuchid, most probably ''Baurusuchus''. A new oospecies called ''Bauruoolithus fragilis'' was named on the basis of these remains. The eggs are about twice as long as they are wide and have blunt ends. At about a quarter of a millimeter in thickness, the shells are relatively thin. Some eggs may have already hatched by the time they were buried, but none show extensive degradation. In living crocodilians (the closest living relatives of baurusuchids), eggs undergo extrinsic degradation to allow hatchlings to easily break through their shells. The fossils indicate that baurusuchid hatchlings probably broke through thin egg shells rather than shells that had been degraded over their incubation period.<ref name=Oetal11>{{cite journal |last=Oliveira |first=C.E.M. |author2=Santucci, R.M. |author3=Andrade, M.B. |author4=Fulfaro, V.J. |author5=Basílo, J.A.F. |author6= Benton, M.J. |year=2011 |title=Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil |journal=Palaeontology |volume=54 |issue=2 |pages=309–321 |doi=10.1111/j.1475-4983.2010.01028.x|bibcode=2011Palgy..54..309O |doi-access=free }}</ref>
==References== {{Reflist}}
{{Wikispecies|Baurusuchidae}}
{{Notosuchia|S.}} {{Taxonbar|from=Q140104}}
Category:Late Cretaceous crocodylomorphs Category:Terrestrial crocodylomorphs Category:Turonian first appearances Category:Maastrichtian extinctions Category:Baurusuchidae Category:Pseudosuchian families