{{short description|Species of plant}} {{Speciesbox | image = Holcus lanatus Gestreepte witbol (2).jpg | genus = Holcus | species = lanatus | authority = L. }}

'''''Holcus lanatus''''' is a perennial flowering plant in the grass family Poaceae. The specific epithet ''{{lang|la|lanatus}}'' is Latin for 'woolly' which describes the plant's hairy texture. Common names include '''Yorkshire fog''', '''tufted grass''', and '''meadow soft grass'''. In North America, where it is an invasive species,<ref>[http://www.cal-ipc.org/ip/definitions/index.php CAL-IPC Invasive Plant Definitions.] California Invasive Plant Council.</ref> names include '''velvet grass''' and '''common velvet grass'''.<ref name=hub>Hubbard, C. E. ''Grasses''. Harmondsworth: Penguin Books. 1976. {{ISBN|978-0-14-013227-4}}</ref><ref name=go>[http://www.gardenorganic.org.uk/organicweeds/weed_information/weed.php?id=81 Yorkshire Fog.] {{Webarchive|url=https://web.archive.org/web/20120722031359/http://www.gardenorganic.org.uk/organicweeds/weed_information/weed.php?id=81 |date=2012-07-22 }} Garden Organic. Henry Doubleday Research Association (HDRA).</ref>

In parts of northern Europe the grass is a common native species and a hardy pasture grass.{{not verified in body|date=May 2024}}

==Characteristics and hybrids== [[File:Holcus.lanatus.3.jpg|right|thumb|Flower showing anthers]] ''Holcus lanatus'' has velvety grey-green leaves. The stems are round. The bases of the stems are white with pink stripes or veins; this character has been called the "stripy pyjamas".<ref name=nc>[http://www.naturescalendar.org.uk/wildlife/factfiles/grasses/yorkshirefog.htm ''Holcus lanatus''.] {{Webarchive|url=https://web.archive.org/web/20130929235007/http://www.naturescalendar.org.uk/wildlife/factfiles/grasses/yorkshirefog.htm |date=2013-09-29 }} The Nature's Calendar Survey.</ref> The inflorescence is robust and often tinged purple. It produces a large amount of seed and is a rapid coloniser of disturbed ground. It prefers wetter ground; it is often seen around drainage ditches. The ligule is {{convert|1|-|4|mm}} long, blunt, and hairy.

This species can be distinguished from ''H.&nbsp;mollis'' by the beardless nodes on its culm, the absence of rhizomes, and the awn becoming hooked when dry and not projecting beyond the tips of the glumes.<ref name=hub/> It has been known to hybridize with ''H.&nbsp;mollis'', producing a male sterile hybrid with 2''n'' = 21 chromosomes.<ref name=hub/> Hybrids tend to resemble ''H.&nbsp;lanatus'' in their morphology.<ref name=go/>

It spreads vegetatively by developing new shoots and roots at its nodes. Plants form a blanket of runners on the soil surface. Semi-prostrate rosettes of shoots called 'mops' may form at the end of the runners. These mops root readily in contact with moist soil.<ref name=go/>

==Invasive species and habitat preferences== right|thumb|Mature flowers In a European survey of weed contamination in cereal seed in 1970, ''Holcus lanatus'' seed was found in 1% of samples. ''H.&nbsp;lanatus'' is an indicator of poor soil, low grazing levels, and poor drainage. It is tolerant of a range of soil pH, but grows best between 5.0 and 7.5. It exhibits climatic tolerance over a wide altitude range, but severe frosts can kill it. It does not survive trampling and puddling. It can be controlled in some European locations by increasing available potassium and phosphorus, increasing stock, and improving drainage. These remedies are not as effective in North America.<ref name=go/>

===Noxious weed=== ''Holcus lanatus'' is a significant pest weed in Australia, as it is a winter-growing C<sub>3</sub> grass and survives droughts and hot summers as seed. It is distasteful to stock unless it is young and little other plant material is available. The flowers are wind-pollinated and usually out-crossing. The first seeds become viable 5 to 9 days after flowering and all are viable after 20 days. Seeds are shed from in summer and early autumn. One panicle has 100 to 380 seeds, with 177,000 to 240,000 seeds per plant, depending on time of emergence.<ref name=go/>

===Invasive species=== In North America, ''Holcus lanatus'' is an invasive species in native grasslands and other ecosystems. In Yosemite National Park it is one of nine priority noxious weeds to control for habitat restoration and regenerating native plant balances.<ref>[https://web.archive.org/web/20090515050702/http://www.nps.gov/yose/naturescience/invasive-plant-management.htm Invasive Plant Management.] Yosemite National Park. National Park Service. Retrieved June 20, 2013.</ref> It forms dense stands that can exclude other plants. {{Commons}} It is also established in Chile and Australia. In the former it has been found as an invasive species in ''Sphagnum'' peatlands disturbed by peat extraction in southern Patagonia.<ref name=anales>{{Cite journal |title=Efectos de la extracción de turba sobre la composición y estructura de una turbera de Sphagnum explotada y abandonada hace 20 años, Chile |journal=Anales del Instituto de la Patagonia |url=https://www.scielo.cl/scielo.php?script=sci_arttext&pid=S0718-686X2012000200003 |last1=Domínguez |first1=Erwin |issue=2 |volume=40 |pages=37–45 |last2=Bahamonde |first2=Nelson |language=Spanish |trans-title=Effects of peat extraction on the structure and composition of Sphagnum peatland exploited and abandones for 20 years, Chile |last3=Muñoz-Escobar |first3=Christian |date=2012 |doi=10.4067/S0718-686X2012000200003 |doi-access=free }}</ref>

==Ecology== ''Holcus lanatus'' in its natural habitat is a food source for butterflies such as the speckled wood, the wall, and especially the small skipper. It is rarely utilized by the Essex skipper. In its native range it may occur in plant associations such as the ''Juncus subnodulosus''–''Cirsium palustre'' fen-meadow habitat.

==See also== *Invasive grasses of North America

==References== {{Reflist}}

{{Taxonbar|from=Q158110}}

Category:Pooideae Category:Flora of Africa Category:Flora of Asia Category:Flora of Europe Category:Plants described in 1753 Category:Botanical taxa named by Carl Linnaeus