{{Short description|Family of gastropods}} {{Automatic taxobox | image = Holospira elizabethae shell.jpg | image_caption = Apertural view of a shell of ''Holospira elizabethae'' | taxon = Urocoptidae | authority = Pilsbry, 1898<ref>Pilsbry H. A. (1898). ''The Nautilus'' [https://archive.org/details/nautilus11amer '''11'''](9): [https://archive.org/stream/nautilus11amer#page/107/mode/1up 107].</ref> (1868) | synonyms = }}
'''Urocoptidae''' is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea.<ref name="Uit de Weerd 2008"/>
== Taxonomy == === 2005 taxonomy === The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).<ref name="Bouchet 2005">{{Bouchet 2005}}</ref>
The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868 * Apomatinae Paul, 1982 * Brachypodellinae H. B. Baker, 1956 * Eucalodiinae P. Fischer & Crosse, 1873 * Holospirinae Pilsbry, 1946 * Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972 * Tetrentodontinae Bartsch, 1943
=== 2008 taxonomy === Uit de Weerd (2008)<ref name="Uit de Weerd 2008"/> moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.<ref name="Uit de Weerd 2008">{{cite journal |pages=317–29 |doi=10.1093/mollus/eyn023 |title=Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion |year=2008 |last1=Uit De Weerd |first1=D. R. |journal=Journal of Molluscan Studies |volume=74 |issue=4|doi-access= }}</ref>
The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.<ref name="Uit de Weerd 2008"/> There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.<ref name="Uit de Weerd 2008"/>
{{clade |label1=Urocoptoidea |1={{clade |1={{clade |1={{clade |1={{clade |1={{clade |1={{clade |1=''Macroceramus microdon'' |2=''Microceramus pontificus'' }} |2={{clade |1=''Torrecoptis holguinensis'' |2=''Autocoptis menkeana'' }} }} |2={{clade |1=''Simplicervix inornata'' |2=''Brachypodella dominicensis gabbi'' }} }} |2={{clade |1={{clade |1={{clade |1=''Dissotropis'' sp. nov. [https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=513533&lvl=3&p=mapview&p=has_linkout&p=blast_url&p=genome_blast&lin=f&keep=1&srchmode=1&unlock] |2=''Anisospira lebmanni'' }} |2=''Archegocoptis crenata'' }} |2=''Eucalodium'' sp. }} }} |label2={{Background color|Khaki|Cerionidae}} |2={{Background color|Khaki|''Cerion striatellum''}} }} |2={{clade |1=''Coelostemma'' sp. nov. [https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=513532&lvl=3&p=mapview&p=has_linkout&p=blast_url&p=genome_blast&keep=1&srchmode=1&unlock&mod=1#modif] |2=''Hendersoniella lux lux'' |3=''Holospira fortisculpta'' }} |3=''Epirobia polygyra'' }} }}
=== 2012 taxonomy === Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (''Epirobia'', ''Propilsbrya'', ''Pectinistemma'', ''Gyrocion'', ''Prionoloplax''), that were previously placed within Urocoptidae.<ref name="Thompson 2012">Thompson F. G. (2012). "The land snail genus ''Epirobia'' and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". ''Bulletin of the Florida Museum of Natural History'' '''51''': 167-215. [http://www.flmnh.ufl.edu/bulletin/vol51no3.pdf PDF].</ref> Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.<ref name="Thompson 2012"/>
==Genera == Genera in the family Urocoptidae include:
Urocoptinae * ''Autocoptis'' Pilsbry, 1902<ref name="Uit de Weerd 2008"/> * ''Brachypodella'' L. Pfeiffer, 1840 (as ''Cylindrella'') * ''Urocoptis'' Beck, 1837 - type genus of the family Urocoptidae<ref name="Bouchet 2005"/>
Apominae * ''Apoma'' Beck, 1837 - type genus of the subfamily Apomatinae<ref name="Bouchet 2005"/> * ''Simplicervix'' Pilsbry, 1903<ref name="Uit de Weerd 2008"/>
Brachypodellinae * ''Brachypodella'' Beck, 1837 - type genus of the subfamily Brachypodellinae<ref name="Bouchet 2005"/>
Eucalodiinae * ''Anisospira'' Strebel, 1880<ref name="Uit de Weerd 2008"/> * ''Archegocoptis'' Pilsbry, 1903<ref name="Uit de Weerd 2008"/> *†''Condonella'' <small>McLellan, 1927</small><ref name="Roth2000">{{cite journal |last1=Roth |first1=B. |year=2000 |title=Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California |journal=Journal of Molluscan Studies |volume=66 |issue=3 |pages=373–381 |url=http://mollus.oxfordjournals.org/content/66/3/373.full.pdf+html |doi=10.1093/mollus/66.3.373|doi-access=free |url-access=subscription }}</ref> * ''Dissotropis'' Bartsch, 1906<ref name="Uit de Weerd 2008"/> * ''Eucalodium'' Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae<ref name="Bouchet 2005"/>
Holospirinae * ''Coelostemma'' Dall, 1895<ref name="Uit de Weerd 2008"/> <!-- ''Epirobia'' Strebel & Pfeffer, 1880<ref name="Uit de Weerd 2008"/> IT IS WITHIN Epirobiidae --> * ''Hendersoniella'' Dall, 1905<ref name="Uit de Weerd 2008"/> * ''Holospira'' Martens, 1860 - type genus of the subfamily Holospirinae<ref name="Bouchet 2005"/>
Microceraminae * ''Microceramus'' Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae<ref name="Bouchet 2005"/> * ''Macroceramus'' Guilding, 1828<ref name="Uit de Weerd 2008"/>
Tetrentodontinae * ''Tetrentodon'' Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae<ref name="Bouchet 2005"/> * ''Torrecoptis'' Bartsch, 1943<ref name="Uit de Weerd 2008"/>
Other genera include:<ref>{{cite web| url=https://www.discoverlife.org/mp/20q?search=Urocoptidae | title=Urocoptidae - MICROCERAMIDAE | access-date=2024-01-22 | url-status=live | archive-url=https://web.archive.org/web/20120716235158/https://www.discoverlife.org/mp/20q?search=Urocoptidae | archive-date=2012-07-16}}</ref> * ''Allocoptis'' Thompson & Franz, 1976 * ''Amphicosmia'' Pilsbry & Vanatta, 1898 * ''Anoma'' * ''Arangia'' * ''Bactrocoptis'' * ''Badiofax'' * ''Bostrichocentrum'' * ''Bostrychocentrum'' * ''Callonia'' * ''Centralia'' * ''Cochlodinella'' * ''Coelocentrum'' * ''Geoscala'' * ''Gongylostoma'' * ''Gyraxis'' * ''Haplocion'' * ''Heterocoptis'' * ''Idiostemma'' * ''Liocallonia'' * ''Metastoma'' * ''Mychostoma'' * ''Nesocoptis'' * ''Nodulia'' * ''Oligostylus'' * ''Organocoptis'' * ''Pfeiffericoptis'' * ''Pineria'' * ''Pleurostemma'' * ''Poeycoptis'' * ''Pseudopineria'' * ''Pycnoptychia'' Pilsbry & Vanatta, 1898 * ''Sagracoptis'' * ''Spiroceramus'' * ''Spirostemma'' * ''Strophina'' * ''Tomelasmus'' * ''Trilamellaxis'' * ''Uncinicoptis''
==References== {{Reflist|30em}}
== Further reading == * {{cite journal | last1 = Bartsch | first1 = P. | author-link = Paul Bartsch | year = 1943 | title = Notes on Mexican Urocoptid mollusks | journal = Journal of the Washington Academy of Sciences | volume = 33 | issue = 2| pages = 54–59 }}; 7 figs. * Carlos de la Torre. 1912. ''New Cuban Urocoptis of the U. cinerea group''. The Nautilus, [https://archive.org/details/nautilus26amer volume 26], number 4, pages [https://archive.org/stream/nautilus26amer#page/54/mode/2up 55]-58, [https://archive.org/stream/nautilus26amer#page/n71/mode/2up plate 6]. * Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". ''Zootaxa'' '''3362''': 43–53. [http://www.mapress.com/zootaxa/2012/f/zt03362p053.pdf PDF]. * Pilsbry H. A. & Henderson J. B. jr. (February 1913). ''Two new Cuban Urocoptidae''. [https://archive.org/details/nautilus26amer The Nautilus, volume 26], number 10, pages [https://archive.org/stream/nautilus26amer#page/n137/mode/2up 109]-110. * {{in lang|es}} Miguel L. Jaume & Alfredo de la Torre. 1976. ''Los Urocoptidae de Cuba (Mollusca-Pulmonata)''. Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp. *{{cite journal |pmid=6763741 |year=1982 |last1=Mesa Garcia |first1=R |last2=Jaume |first2=ML |title=Zoogeography, ecology, and protection of the Urocoptidae family in Cuba (Gastropoda-Pulmonata). II |volume=34 |issue=2 |pages=172–92 |journal=Revista Cubana de Medicina Tropical}} * Richardson C. L. 1991. ''Urocoptidae: catalogue of species''. Tryonia 22: 1–245. * Tryon G. W. (1903) ''Manual of Conchology. Second series: Pulmonata. [https://archive.org/details/manualofconcholo15tryorich Volume 15]. <!-- part 57 -->Urocoptidae''. Continued by H. A. Pilsbry.
== External links == {{Commons category|Urocoptidae}} * [http://elrincon-continentales3.iespana.es/Urocoptidae.htm photos] * [http://shell.kwansei.ac.jp/~shell/pic_book/family/3548.html photos] * [http://www.ashbreure.nl/snailblog/files/tag-urocoptidae.html blog Bram's Snail Site with photos]
{{Taxonbar|from=Q3074668}} {{Authority control}}
Category:Urocoptidae Category:Taxa named by Henry Augustus Pilsbry