{{Short description|Genus of amphibians}} {{Automatic taxobox | image = uperoleia_tyleri.jpg | image_caption = Tyler's toadlet, ''Uperoleia tyleri'' | taxon = Uperoleia | authority = Gray, 1841 | subdivision_ranks = Species | subdivision = See text. | synonyms = ''Hyperolia'' — unjustified emendation<br /> ''Glauertia'' <small>Loveridge, 1933</small><br /> ''Hosmeria'' <small>Wells and Wellington, 1985</small><br /> ''Prohartia'' <small>Wells and Wellington, 1985</small> | synonyms_ref = <ref name=Frost/> }}
'''''Uperoleia''''' is a genus of frogs in the family Myobatrachidae. They are native to northern and eastern Australia and southern lowlands of New Guinea.<ref name=Frost/> These are small squat frogs, more commonly known as "toadlets". They have glandular skin, often with a pair of raised glands behind each eye, or on the flanks.
They have bumpy, rough skin giving them the appearance of a small toad, hence the name "toadlet", although they are often called "gungans" in Queensland. There are two distinct types of calls—''Uperoleia'' species make either a "click" or a "squelch". Generally, the "clicking" ''Uperoleia'' have long thin inguinal glands that run along the dorsal surface, while the "squelching" ''Uperoleia'' have round inguinal glands restricted to the posterior half of the dorsal surface. It is unusual to find more than one species of "clickers" or "squelchers" in the same location, although finding one of each is quite frequent in northern Australia. The species in this genus show great similarities in body shape and colouration making them difficult to tell apart. Call analysis is often required to confirm identification.{{citation needed|date=February 2017}} This genus is the largest of any in the family Myobatrachidae.<ref name="Catullo et al. 2011"/><ref name=Frost-Myobatrachidae/>
Clicking ''Uperoleia'' include ''U. glandulosa, U. aspera, U. minima, U. trachyderma, U. lithomoda, U. littlejohni, U. altissima, U. mimula,'' and ''U. rugosa''. The squelching ''Uperoleia'' include ''U. russelli, U. saxatilis, U. talpa, U. borealis, U. crassa'', and ''U. inundata''. The eastern species ''U. laevigata, U. fusca, U. tyleri,'' and ''U. martini'' are also squelchers, but are distantly related. The species ''U. mjobergi, U. micromeles, U. micra,'' and ''U. daviesae'' are distinct from these other groups.<ref name="Catullo et al. 2011">{{cite journal |last1= Catullo |first1=R.A. |author2=Doughty, P. |author3=Roberts, J.D. |author4=Keogh, J.S. |year=2011 |title=Multi-locus phylogeny and taxonomic revision of ''Uperoleia'' toadlets (Anura: Myobatrachidae) from the western arid zone of Australia, with a description of a new species |journal=Zootaxa |volume=2902 |pages=1–43 |doi=10.11646/zootaxa.2902.1.1 |url=http://www.mapress.com/zootaxa/2011/f/zt02902p043.pdf|doi-access=free }}</ref>
==Species== Following the Amphibian Species of the World, there are 29 species:<ref>{{Cite web |title=Uperoleia Gray, 1841 {{!}} Amphibian Species of the World |url=https://amphibiansoftheworld.amnh.org/Amphibia/Anura/Myobatrachoidea/Myobatrachidae/Uperoleia |access-date=2022-04-26 |website=amphibiansoftheworld.amnh.org}}</ref> {|style="text-align:left; border:1px solid #999999; " |-style="background:#CCCC99; text-align: center; " !Common name!!Binomial name |-style="background:#EEEEEE;" |Montane toadlet||''Uperoleia altissima'' <br /> {{Small|Davies, Watson, McDonald, Trenerry & Werren, 1993}} |-style="background:#FFFDFF;" |Jabiru toadlet||''Uperoleia arenicola'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Derby toadlet||''Uperoleia aspera'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#FFFDFF;" |Northern toadlet||''Uperoleia borealis'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Fat toadlet||''Uperoleia crassa'' {{Small|Tyler, Davies & Martin, 1981}} |-style=background:#FFFDFF;" |Howard Springs toadlet||''Uperoleia daviesae'' <small>Young, Tyler & Kent, 2005</small> |-style="background:#EEEEEE;" |Dusky toadlet||''Uperoleia fusca'' {{Small|Davies, McDonald & Corben, 1986}} |-style="background:#FFFDFF;" |Glandular toadlet||''Uperoleia glandulosa'' {{Small|Davies, Mahony & Roberts, 1985}} |-style="background:#EEEEEE;" |Gurrumul's toadlet |''Uperoleia gurrumuli'' {{Small|Catullo & Keogh, 2021}} |-style="background:#FFFDFF;" |Flood plain toadlet||''Uperoleia inundata'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Smooth toadlet||''Uperoleia laevigata'' {{Small|Keferstein, 1867}} |-style="background:#FFFDFF;" |Stonemason's toadlet||''Uperoleia lithomoda'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Littlejohn's toadlet||''Uperoleia littlejohni'' {{Small|Davies, McDonald & Corben, 1986}} |-style="background:#FFFDFF;" |Mahony's toadlet||''Uperoleia mahonyi'' {{Small|Clulow, Anstis, Keogh, and Catullo, 2016}} |-style="background:#EEEEEE;" |Marbled toadlet, yellow-spotted toadlet||''Uperoleia marmorata'' {{Small|Gray, 1841}} |-style="background:#FFFDFF;" |Martin's toadlet||''Uperoleia martini'' {{Small|Davies & Littlejohn, 1986}} |-style="background:#EEEEEE;" |Tiny toadlet||''Uperoleia micra'' {{Small|Doughty and Roberts, 2008}} |-style="background:#FFFDFF;" |Tanami toadlet||''Uperoleia micromeles'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Mimic toadlet||''Uperoleia mimula'' {{Small|Davies, McDonald & Corben, 1986}} |-style="background:#FFFDFF;" |Small toadlet||''Uperoleia minima'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Mjoberg's toadlet||''Uperoleia mjobergi'' {{Small|(Andersson, 1913)}} |-style="background:#FFFDFF;" |Alexandria toadlet||''Uperoleia orientalis'' {{Small|(Parker, 1940)}} |-style="background:#EEEEEE;" |Wrinkled toadlet, red-groined toadlet||''Uperoleia rugosa'' {{Small|(Andersson, 1916)}} |-style="background:#FFFDFF;" |Russell's toadlet||''Uperoleia russelli'' {{Small|(Loveridge, 1933)}} |-style="background:#EEEEEE;" |Pilbara toadlet||''Uperoleia saxatilis'' {{Small|Catullo ''et al.'', 2011}}<ref name="Catullo et al. 2011"/> |-style="background:#FFFDFF;" |Ratcheting toadlet||''Uperoleia stridera'' {{Small|Catullo, Doughty, and Keogh, 2014}} |-style="background:#EEEEEE;" |Mole toadlet||''Uperoleia talpa'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#FFFDFF;" |Blacksoil toadlet||''Uperoleia trachyderma'' {{Small|Tyler, Davies & Martin, 1981}} |-style="background:#EEEEEE;" |Tyler's toadlet||''Uperoleia tyleri'' {{Small|Davies & Littlejohn, 1986}} |- |}
Note that the AmphibiaWeb recognizes the small-headed toadlet (''Uperoleia capitulata'' {{Small|(Davies, McDonald & Corben, 1986)}}),<ref name=AmphibiaWeb/> a species that the Amphibian Species of the World, following Catullo and Keogh (2014),<ref name="Catullo and Keogh 2014"/> treats as a synonym of the wrinkled toadlet (''Uperoleia rugosa'').<ref name=Frost/>
==References== {{Reflist|30em|refs= <ref name=AmphibiaWeb>{{cite web |url=http://www.amphibiaweb.org/lists/Myobatrachidae.shtml|title=Myobatrachidae |year=2017 |work=AmphibiaWeb |publisher=University of California, Berkeley |access-date=6 February 2017}}</ref>
<ref name="Catullo and Keogh 2014">{{cite journal|last1=Catullo|first1=R.A.|last2=Keogh|first2=J.S. |title=Aridification drove repeated episodes of diversification between Australian biomes: Evidence from a multi-locus phylogeny of Australian toadlets (''Uperoleia'': Myobatrachidae) |journal=Molecular Phylogenetics and Evolution |date=2014 |volume=79 |pages=106–117 |doi=10.1016/j.ympev.2014.06.012|pmid=24971737}}</ref>
<ref name=Frost>{{cite web |url=http://research.amnh.org/vz/herpetology/amphibia/Amphibia/Anura/Myobatrachidae/Uperoleia |title=''Uperoleia'' Gray, 1841 |author=Frost, Darrel R. |year=2016 |work=Amphibian Species of the World: an Online Reference. Version 6.0 |publisher=American Museum of Natural History |access-date=6 February 2017}}</ref>
<ref name=Frost-Myobatrachidae>{{cite web |url=http://research.amnh.org/vz/herpetology/amphibia/Amphibia/Anura/Myobatrachidae |title=Myobatrachidae Schlegel, 1850 |author=Frost, Darrel R. |year=2016 |work=Amphibian Species of the World: an Online Reference. Version 6.0 |publisher=American Museum of Natural History |access-date=6 February 2017}}</ref> }}
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Category:Uperoleia Category:Myobatrachidae Category:Amphibian genera Category:Taxa named by John Edward Gray