Sematurinae

{{Short description|Subfamily of moths}} {{Automatic taxobox | image = Acylasisis.JPG | image_caption = ''Mania'' species from the collection of Felix Stumpe | taxon = Sematurinae | authority = | subdivision_ranks = Genera | subdivision = *''Coronidia'' Westwood, 1879 **=''Conoris'' Berthold, 1827 [rejected name] **=''Larunda'' Hübner, 1823 [junior homonym] **=''Coronis'' Latreille, 1829 [junior homonym] *''Homidiana'' Strand, 1914 **=''Homidia'' Strand, 1911 [junior homonym] *''Mania'' Hübner, 1821 **=''Nothus'' Billberg, 1820 [junior homonym of ''Nothus'' Ziegler in Oliver, 1811] **=''Sematura'' Dalman, 1825 **=''Manidia'' Westwood, 1879 [unnecessary replacement name] *''Lonchotura'' Hampson, 1918 *''Anurapteryx'' Hampson, 1918 }}

'''Sematurinae''' is a subfamily of moths in the family Sematuridae represented by at least 29 species in the Neotropics.

==Taxonomy, classification and identification== Sematurinae has three principal genera, and possibly another two. The hindwing of ''Coronidia'' is distinguished by an iridescent blue band,<ref>{{cite web|author=Leptree.net |url=http://www.leptree.net/Coronidia_orithea |title=leptree.net |publisher=leptree.net |date=2007-01-27 |accessdate=2012-06-12 |url-status=dead |archiveurl=https://web.archive.org/web/20160120010033/http://www.leptree.net/Coronidia_orithea |archivedate=2016-01-20 }}</ref> but ''Homidiana'' characteristically has pink, orange or yellow markings,<ref>{{cite web|url=http://www.sil.si.edu/imagegalaxy/imageGalaxy_enlarge.cfm?id_image=5972 |title=sil.si.edu |publisher=sil.si.edu |date= |accessdate=2012-06-12}}</ref> especially in females (Minet and Scoble, 1999). The genus ''Mania'' is large, brown and tailed, resembling to some extent the genus ''Lyssa''. The approximately twelve other sematurid species in the genera ''Anurapteryx'' and ''Lonchotura'' are placed here for now (making about 41 species) but may not belong in this subfamily; one species, ''Anurapteryx crenulata'' reaches the subtropics of Arizona<ref>{{cite web |url=http://nitro.biosci.arizona.edu/zeeb/butterflies/figs/moths/Sematuridae/A_crenulata.jpg |archive-url=https://web.archive.org/web/20020816064109/http://nitro.biosci.arizona.edu/zeeb/butterflies/figs/moths/Sematuridae/A_crenulata.jpg |url-status=dead |archive-date=2002-08-16 |title=nitro.biosci.arizona.edu |accessdate=2012-06-12 }}</ref> (Minet and Scoble, 1999). The relationships within Sematuridae clearly need deeper investigation.

*Note: the name ''Nothus'' is preoccupied by a genus of Coleoptera<ref>{{cite web |url=http://uio.mbl.edu/pleary/Animalium/index.php?search=page&id=4418 |title=uio.mbl.edu |publisher=uio.mbl.edu |accessdate=2012-06-12 |url-status=dead |archiveurl=https://web.archive.org/web/20110929204810/http://uio.mbl.edu/pleary/Animalium/index.php?search=page&id=4418 |archivedate=2011-09-29 }}</ref> (Minet and Scoble, 1999). The genus name ''Sematura'' was also in use for this genus, but it was younger (being published in 1825) than Hübner's name ''Mania'' from 1821; a petition to conserve the junior name was rejected by the ICZN in Opinion 2352 (2015),<ref>[http://iczn.org/node/40515 Opinion 2352 (Case 3531): ''Sematura'' Dalman, 1825 (Insecta, Lepidoptera, Sematuridae): precedence over ''Mania'' Hübner, 1821 not given]</ref> so Hübner's name is officially now the valid name for the genus.

==Behaviour== The adult moths of Sematurinae are either day or night flying, depending on the genus and have the wings outspread at rest and the genus ''Sematura'' displays impressive eyespots at the hindwing tails.

==Biology== The biology of Sematurinae is poorly known. The larva of ''Homidiana'' was illustrated by Westwood (1879) who incorrectly placed it among ''Coronidia'', and was described by Anton Hermann Fassl (1910). From 1995 to 1998, caterpillars were found in Costa Rica.<ref>{{cite web|url=http://janzen.sas.upenn.edu |title=janzen.sas.upenn.edu |publisher=janzen.sas.upenn.edu |date= |accessdate=2012-06-12}}</ref> Sematurine eggs have been described by Joël Minet and Malcolm J. Scoble (1999:302), and are upright, high, with vertical and finer transverse ribs. The caterpillars of ''Homidiana subpicta'' have rows of black spines either side of the dorsal midline and five pairs of prolegs.<ref>[http://janzen.sas.upenn.edu/2700ARCHIVES/Sematuridae/Coronidia%20subpicta/06-SRNP-4964_DHJ410813.jpg janzen.sas.upenn.edu]{{dead link|date=June 2012}}</ref> The larva of ''Coronidia orithea'' lacks such spines, and is well camouflaged like a lichen-covered twig<ref>[http://janzen-db.bio.upenn.edu:16080/2700ARCHIVES/Sematuridae/Coronidia%20orithea/95-SRNP-8509_DHJ25681_f.jpg janzen-db.bio.upenn.edu]{{dead link|date=December 2016}}</ref> and has realistic-looking false eyes making it resemble a predatory spider.<ref>[http://janzen-db.bio.upenn.edu:16080/2700ARCHIVES/Sematuridae/Coronidia%20orithea/95-SRNP-8509_DHJ25677_f.jpg janzen-db.bio.upenn.edu]{{dead link|date=December 2016}}</ref> The larvae of ''Mania'' were reared in Costa Rica in 2001 and are a bit similar to those of ''Coronidia'' being well camouflaged but bearing small projections rather than long spines one the dorsal surface.<ref>[http://www.tulane.edu/~ldyer/lsacat/species/uraniidae/ura1/17327g.gif tulane.edu]{{dead link|date=June 2012}}</ref> Like some Uraniidae, pupation occurs on the ground among debris. The pupa of ''Homidiana'' has a long projecting proboscis "sheath"<ref>[http://janzen.sas.upenn.edu/2700ARCHIVES/Sematuridae/Coronidia%20subpicta/00-SRNP-9565_DHJ53585_f.jpg janzen.sas.upenn.edu]{{dead link|date=June 2012}}</ref> with eight hooklets at the cremaster tip (Minet and Scoble, 1999).

==Larval host plants== The caterpillar of ''Coronidia orithea'' is so far only known to feed on mistletoes (Viscaceae: ''Phoradendron quadrangulare''). ''Homidiana subpicta'' feeds on a more diverse range of plants, including the families Araliaceae (''Oreopanax''), Gesneriaceae, Piperaceae (''Piper''), Primulaceae (''Ardisia'', ''Myrsine''), Solanaceae (''Solanum''), Onagraceae (''Ludwigia'') and Theophrastaceae (''Clavija'').<ref>{{cite web|url=http://janzen.sas.upenn.edu |title=janzen.sas.upenn.edu |publisher=janzen.sas.upenn.edu |date= |accessdate=2012-06-12}}</ref> ''"Sematura luna"''<ref>[http://www.tulane.edu/~ldyer/lsacat/species/uraniidae/ura1/ura1.htm tulane.edu] {{webarchive |url=https://web.archive.org/web/20041105153344/http://www.tulane.edu/~ldyer/lsacat/species/uraniidae/ura1/ura1.htm |date=November 5, 2004 }}</ref> reared in 2001 whose well-camouflaged larva is reported to feed on the legume ''Pentaclethra macroloba'' (Mimosoideae) and on ''Syzygium longifolium'' (Myrtaceae).

==Provisional list of species (based on LepIndex)== *''Coronidia erecthea'' Westwood, 1879 [type locality Espiritu Santu, Brazil]<ref>{{LepIndex |id=69147 |name=Coronidia erecthea |accessdate=May 14, 2018}}</ref> *''Coronidia hyphasis'' (Hopffer, 1856) (originally in ''Coronis'') [type locality Mexico]<ref>{{LepIndex |id=69148 |name=Coronidia hyphasis |accessdate=May 14, 2018}}</ref> *''Coronidia orithea'' (Cramer, ''Papill. Exot.'' 3:121) (originally in ''Phalaena'') [type locality Suriname]<ref>{{cite web |last=Savela |first=Markku |title=''Coronidia'' Westwood, 1879 |work=Lepidoptera and some other life forms |url=https://nic.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/geometroidea/sematuridae/coronidia/#orithea |access-date=25 May 2025}}</ref> **=''? durvillii'' Guenée, 1857<ref>{{LepIndex |id=69150 |name=Coronidia orithea |accessdate=May 14, 2018}}</ref> **=''Coronidia hysudrus'' Hopffer, 1857<ref>{{LepIndex |id=69151 |name=Coronidia orithea |accessdate=May 14, 2018}}</ref> **=''Coronidia boreada'' Westwood, 1879<ref>{{LepIndex |id=69152 |name=Coronidia orithea |accessdate=May 14, 2018}}</ref> *''Coronidia difficilis'' Strand, 1911 [type locality Chanchamayo, Peru]<ref>{{LepIndex |id=69153 |name=Coronidia difficilis |accessdate=May 14, 2018}}</ref> **=''Coronidia ecuadorensis'' Strand, 1911<ref>{{LepIndex |id=69154 |name=Coronidia difficilis |accessdate=May 14, 2018}}</ref> *''Homidiana aeneophlebia'' (Hampson, ''Novitates Zoologia'' 25:374) (originally in ''Homidia'')<ref>{{LepIndex |id=69157 |name=Homidiana aeneophlebia |accessdate=May 14, 2018}}</ref> *''Homidiana anilina'' Bryk, 1930 [type locality Bolivia]<ref>{{LepIndex |id=69158 |name=Homidiana anilina |accessdate=May 14, 2018}}</ref> *''Homidiana brachyura'' Hampson, ''Novit. Zool.'' 25:373 (originally in ''Coronidia'' subg. (''Homidiana''))<ref>{{LepIndex |id=69159 |name=Homidiana brachyura |accessdate=May 14, 2018}}</ref> *''Homidiana briseis'' (Westwood, 1879) (originally in ''Coronidia'')<ref>{{LepIndex |id=69160 |name=Homidiana briseis |accessdate=May 14, 2018}}</ref> *''Homidiana canace'' Hopffer, 1856 [https://web.archive.org/web/20060907224229/http://data.acnatsci.org/peale/peale.php?mode=specimen&collection=CMNH&specimen=855] [http://www.sil.si.edu/imagegalaxy/imageGalaxy_enlarge.cfm?id_image=5972]<ref>{{LepIndex |id=69161 |name=Homidiana canace |accessdate=May 14, 2018}}</ref> **=''Coronidia aeola'' Westwood, 1879<ref>{{LepIndex |id=69162 |name=Homidiana canace |accessdate=May 14, 2018}}</ref> **=''Coronidia paulina'' Westwood, 1879<ref>{{LepIndex |id=69165 |name=Homidiana egina |accessdate=May 14, 2018}}</ref> **=''Coronis cana'' Hopffer, 1856<ref>{{LepIndex |id=69163 |name=Homidiana canace |accessdate=May 14, 2018}}</ref> *''Homidiana egina'' (Blanchard, 1849) (originally in ''Coronidia'')<ref>{{LepIndex |id=69164 |name=Homidiana egina |accessdate=May 14, 2018}}</ref> **=''Homidiana nicaraguana'' Westwood, 1879<ref>{{LepIndex |id=69166 |name=Homidiana egina |accessdate=May 14, 2018}}</ref> *''Homidiana evenus'' Boisduval, 1849 **=''Coronis ducatrix'' Schaufuss, 1870<ref>{{LepIndex |id=69168 |name=Homidiana evenus |accessdate=May 14, 2018}}</ref> **=''Homidia subevenus'' Strand, 1911<ref>{{LepIndex |id=69169 |name=Homidiana evenus |accessdate=May 14, 2018}}</ref> *''Homidiana gueneei'' (Druce, 1891) (originally in ''Coronidia'') [type locality Panama, Mexico]<ref>{{LepIndex |id=69170 |name=Homidiana gueneei |accessdate=May 14, 2018}}</ref> *''Homidiana hoppi'' (Bryk, 1930) (originally in ''Coronidia'') [type locality Colombia]<ref>{{LepIndex |id=69171 |name=Homidiana hoppi |accessdate=May 14, 2018}}</ref> *''Homidiana leachi'' (Godart, 1819) (originally in ''Ag[u]arista'') [type locality Brazil]<ref>{{LepIndex |id=69172 |name=Homidiana leachi |accessdate=May 14, 2018}}</ref> **=''Homidiana japet'' (Blanchard, 1849) (originally in ''Coronis'') [type locality Brazil]<ref>{{LepIndex |id=69173 |name=Homidiana leachi |accessdate=May 14, 2018}}</ref> *''Homidiana lederi'' Pfeiffer, 1925 [type locality Macas, Ecuador]<ref>{{LepIndex |id=69174 |name=Homidiana lederi |accessdate=May 14, 2018}}</ref> *''Homidiana leucosticta'' Hampson, 1918 (originally in ''Coronidia'' subg. ''Homidia'') [type locality Peru]<ref>{{LepIndex |id=69175 |name=Homidiana leucosticta |accessdate=May 14, 2018}}</ref> *''Homidiana strandi'' Pfeiffer, 1916 [type locality Peru]<ref>{{LepIndex |id=69176 |name=Homidiana strandi |accessdate=May 14, 2018}}</ref> *''Homidiana monotona'' (Hampson, 1918) (originally in ''Coronidia'') [type locality Huaylas, Peru]<ref>{{LepIndex |id=69177 |name=Homidiana monotona |accessdate=May 14, 2018}}</ref> *''Homidiana restincta'' Strand, 1911 [type locality Macas, Ecuador] (originally in ''Homidia'')<ref>{{LepIndex |id=69178 |name=Homidiana restincta |accessdate=May 14, 2018}}</ref> *''Homidiana rosina'' (Felder & Rogenhofer, 1875) (originally in ''Coronis'') [type locality Bogota, Colombia])<ref>{{LepIndex |id=69179 |name=Homidiana rosina |accessdate=May 14, 2018}}</ref> **=''Coronidia columbiana'' Westwood, 1879<ref>{{LepIndex |id=69180 |name=Homidiana rosina |accessdate=May 14, 2018}}</ref> *''Homidiana rubrivena'' (Dognin, 1919) (originally in ''Coronidia)'' [type locality Colombia]<ref>{{LepIndex |id=69181 |name=Homidiana rubrivena |accessdate=May 14, 2018}}</ref> *''Homidiana subpicta'' Walker, 1854<ref>{{LepIndex |id=69182 |name=Homidiana subpicta |accessdate=May 14, 2018}}</ref> **=''Coronidia biblina'' Westwood, 1879<ref>{{LepIndex |id=69184 |name=Homidiana subpicta |accessdate=May 14, 2018}}</ref> **=''Homidiana echenais'' Hopffer, 1856 [http://www.sil.si.edu/imagegalaxy/imageGalaxy_enlarge.cfm?id_image=5972]<ref>{{LepIndex |id=69184 |name=Homidiana subpicta |accessdate=May 14, 2018}}</ref> **=''Coronidia granadina'' Westwood, 1879<ref>{{LepIndex |id=69185 |name=Homidiana subpicta |accessdate=May 14, 2018}}</ref> *''Homidiana tangens'' (Strand, 1911) (originally in ''Homidia'') [type locality Macas, Ecuador]<ref>{{LepIndex |id=69186 |name=Homidiana tangens |accessdate=May 14, 2018}}</ref> *''Homidiana traducta'' (Strand, 1911) (originally in ''Homidia'') [type locality 2–3000 m. Chaco, Bolivia]<ref>{{LepIndex |id=69187 |name=Homidiana traducta |accessdate=May 14, 2018}}</ref> *''Homidiana westwoodi'' Oberthür, 1881 (originally in ''Coronis'') [type locality Manazales, Colombia]<ref>{{LepIndex |id=69188 |name=Homidiana westwoodi |accessdate=May 14, 2018}}</ref> *''Mania aegisthus'' Fabricius, 1781 [Jamaica/Hispaniola/Cuba] [http://www.mbarnes.force9.co.uk/jamaicamoths/images2/notaeg.htm] [http://www.mbarnes.force9.co.uk/jamaicamoths/images2/notind1.htm "Nothus" species]<ref>{{LepIndex |id=69199 |name=Nothus aegisthus |accessdate=May 14, 2018}}</ref> **=''Mania lunigeraria'' Hübner, 1823 [1825]<ref>{{LepIndex |id=69200 |name=Nothus aegisthus |accessdate=May 14, 2018}}</ref> **=''Sematura excavatus'' Walker, 1854<ref>{{LepIndex |id=69201 |name=Nothus aegisthus |accessdate=May 14, 2018}}</ref> **=''Sematura phoebe'' Guenée, 1857<ref>{{LepIndex |id=69202 |name=Nothus aegisthus |accessdate=May 14, 2018}}</ref> *''Mania diana'' Guenée, 1857 [Rio de Janeiro]<ref>{{LepIndex |id=69203 |name=Nothus diana |accessdate=May 14, 2018}}</ref> *''Mania lunus'' Linnaeus, 1758 [http://www.mbarnes.force9.co.uk/belizemoths/images2/nothind1.htm]<ref>{{LepIndex |id=69205 |name=Nothus lunus |accessdate=May 14, 2018}}</ref> **=''Sematura actaeon'' Felder & Rogenhofer, 1874<ref>{{LepIndex |id=69205 |name=Nothus lunus |accessdate=May 14, 2018}}</ref> **=''Mania caudilunaria'' Hübner, 1823 [1925]<ref>{{LepIndex |id=69206 |name=Nothus lunus |accessdate=May 14, 2018}}</ref> *''Mania empedocles'' Cramer, 1782 [type locality Suriname] [http://www.ogphoto.com/images/bfly2/treebark.jpg]<ref>{{LepIndex |id=69207 |name=Nothus empedocles |accessdate=May 14, 2018}}</ref> **=''Sematura selene'' Guenée, 1857 [https://web.archive.org/web/20041105153344/http://www.tulane.edu/~ldyer/lsacat/species/uraniidae/ura1/ura1.htm Dyer/Gentry/''"Sematura luna"''] [http://www.papillon-poitou-charentes.org/Lepido_Uraniidae/pages/Papillon44.htm]<ref>{{LepIndex |id=69208 |name=Nothus empedocles |accessdate=May 14, 2018}}</ref> **=''Mania empedoclaria'' Hübner, 1823<ref>{{LepIndex |id=69209 |name=Nothus empedocles |accessdate=May 14, 2018}}</ref> *''Anurapteryx beckeri'' (Druce, 1897) (originally in ''Coronidia'') [type locality Mexico, near Durangs City]<ref>{{LepIndex |id=69133 |name=Anurapteryx beckeri |accessdate=May 14, 2018}}</ref> *''Anurapteryx brueckneri'' Hering, 1928<ref>{{LepIndex |id=69134 |name=Anurapteryx brueckneri |accessdate=May 14, 2018}}</ref> *''Anurapteryx crenulata'' Barnes & Lindsey, 1919 [type locality Paradise, Arizona]<ref>{{LepIndex |id=69135 |name=Anurapteryx crenulata |accessdate=May 14, 2018}}</ref> [https://web.archive.org/web/20020816064109/http://nitro.biosci.arizona.edu/zeeb/butterflies/figs/moths/Sematuridae/A_crenulata.jpg Image] *''Anurapteryx flavidorsata'' (Hampson, 1918) (originally in ''Homidia'') [no card] *''Anurapteryx gephyra'' Hering, 1928 [type locality Guatemala]<ref>{{LepIndex |id=69136 |name=Anurapteryx gephyra |accessdate=May 14, 2018}}</ref> *''Anurapteryx interlineata'' (Walker, 1854) [Central and South America, no type locality given] (originally in ''Coronis'')<ref>{{LepIndex |id=69137 |name=Anurapteryx interlineata |accessdate=May 14, 2018}}</ref> **=''Anurapteryx abbreviata'' (Maassen, 1890) (originally in ''Coronis'') [type locality Banos, Ecuador, 1800 m.]<ref>{{LepIndex |id=69138 |name=Anurapteryx interlineata |accessdate=May 14, 2018}}</ref> **=''Anurapteryx meticulosa'' Pief, ''Ent. Zt.'' 31:7 (originally in ?(type locality West Colombia, Rio Aguacabal, 2000 m., by Fassl)<ref>{{LepIndex |id=69139 |name=Anurapteryx interlineata |accessdate=May 14, 2018}}</ref> *''Anurapteryx insolita'' (Strand, 1911) (originally in ''Homidia'') [no type locality]<ref>{{LepIndex |id=69140 |name=Anurapteryx insolita |accessdate=May 14, 2018}}</ref> *''Anurapteryx montana'' Beutelspacher, 1984 [type locality Mexico]<ref>{{LepIndex |id=69141 |name=Anurapteryx montana |accessdate=May 14, 2018}}</ref> *''Anurapteryx ribbei'' (Druce, 1891) (originally in ''Coronidia'') [type locality Chiriqui, Panama]<ref>{{LepIndex |id=69142 |name=Anurapteryx ribbei |accessdate=May 14, 2018}}</ref> *''Lonchotura fassli'' Pfeiffer, ''Ent. Zt.'' 31:7, Pl. (originally in ?) [type locality West Colombia, Rio Aguacabal, by Fassl]<ref>{{LepIndex |id=69190 |name=Lonchotura fassli |accessdate=May 14, 2018}}</ref> *''Lonchotura ocylus'' (Boisduval, 1870) (originally in ''Coronis'') [type locality Guatemala]<ref>{{LepIndex |id=69191 |name=Lonchotura ocylus |accessdate=May 14, 2018}}</ref> **=''Lonchotura dutreuxi'' Deyrolle, 1874 (originally in ''Coronis'') [type locality Costa Rica]<ref>{{LepIndex |id=69192 |name=Lonchotura ocylus |accessdate=May 14, 2018}}</ref> **=''Lonchotura peruviana'' Hering, 1928 (originally in ?) [type locality Chachamayo, Peru]<ref>{{LepIndex |id=69194 |name=Lonchotura ocylus |accessdate=May 14, 2018}}</ref> *''Lonchotura genevana'' Westwood, 1879 (originally in ''Coronidia'') [type locality Mexico]<ref>{{LepIndex |id=69193 |name=Lonchotura genevana |accessdate=May 14, 2018}}</ref>

==References== {{Reflist}} *Fassl, A.H. (1910). Die Raupe einer Uranide. ''Z. wiss. InsektBiol'', '''6'''(10): 355. *Minet, J. and Scoble, M. J. (1999) [1998] The Drepanoid/Geometroid Assemblage. Pp. 301–320 in Kristensen, N.P. (Ed.), 1999 [1998]. ''Lepidoptera, Moths and Butterflies. Volume 1, Evolution, Systematics, and Biogeography''. Handbook of Zoology, vol. IV, Arthropoda: Insecta, Part 35: 491 pp. Walter de Gruyter, Berlin and New York. *Westwood, J.O. 1879. Observations on the Uraniidae, a family of Lepidopterous Insects, with a Synopsis of the Family and a Monograph of ''Coronidia'', one of the genera of which it is composed. ''Transactions of the Zoological Society'', '''10'''(12): 1–35, 3 plates.

==Other sources== *[http://www.nhm.ac.uk/research-curation/projects/lepindex/ NHM Lepindex] *Dyer, L.A. and G.L. Gentry. 2002. Caterpillars and parasitoids of a tropical lowland wet forest, [http://www.caterpillars.org caterpillars.org], Accessed, March 2007.

==External links== *[http://www.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/geometroidea/sematuridae/anurapteryx/index.html Markku Savela's website: ''Anurapteryx'' and ''Lonchotura''] *[http://www.sil.si.edu/imagegalaxy/imageGalaxy_enlarge.cfm?id_image=5972 Insecta. Lepidoptera-Heterocera . Volume III (1881-1900) by Herbert Druce] *[https://web.archive.org/web/20160120010033/http://www.leptree.net/Coronidia_orithea LepTree ''Coronidia orithea''] *[http://www.barcodinglife.org/views/taxbrowser.php?taxid=58866 Barcoding progress] *[https://web.archive.org/web/20070918014718/http://www.leptree.net/status_matrix LepTree nuclear gene sequencing progress] *[https://web.archive.org/web/20070429003038/http://nitro.biosci.arizona.edu/zeeb/butterflies/doid.html Moths of South-East Arizona (''Anurapteryx crenulata'')]

Category:Sematuridae