{{Short description|Genus of ferns}} {{Automatic taxobox | image = Saccoloma inaequale herbarium.jpg | image_caption = ''Saccoloma inaequale''<br/> herbarium specimen | taxon = Saccoloma | authority = Kaulf. | type_species = ''Saccoloma elegans'' | type_species_authority = Kaulf. | subdivision_ranks = Species | subdivision = See text | synonyms_ref = <ref name=POWO_30010104-2/> | synonyms = * ''Neuropteris'' <small>Desv.</small> * ''Orthiopteris'' <small>Copel.</small> }}
'''''Saccoloma''''' is a fern genus in family Saccolomataceae.<ref name=PPGI/><ref name=Phyto>[http://www.mapress.com/phytotaxa/content/2011/f/pt00019p054.pdf Christenhusz ''et al.'' "A linear sequence of extant families and genera of lycophytes and ferns"] ''Phytotaxa'' 19: 7-54. 18 Feb. 2011</ref> It is the only genus in the family in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), but further investigation is needed.<ref name=PPGI/> It is pantropical and its species are found in wet, shaded forest areas.<ref>[http://www.metodosenecologiaysistematica.org/vol5/vol5%281%29/saccoloma5%281%29.pdf Rojas-Alvarado "Novelties in the Saccoloma inaequale complex (Saccolomataceae) from the neotropics"] ''Métodos en Ecología y Sistemática'' 5(1): 1-16. Apr 2010</ref> ''Saccoloma'' species are characterized by an omega-shaped (Ω) vascular bundle in the cross-sections of their petioles.<ref>[http://www.plantsystematics.org/imgs/robbin/r/Dennstaedtiaceae_Saccoloma_moranii_28675.html ''Saccoloma inaequale'' petiole] Image at PlantSystematics.org 01 Feb 2012</ref><ref>[http://www.plantsystematics.org/imgs/robbin/r/Dennstaedtiaceae_Saccoloma_dominguensis_39778.html ''Saccoloma dominguense'' petiole] Image at PlantSystematics.org 01 Feb 2012</ref> The common name '''soralpouch fern''' is used for ''Saccoloma''.<ref>{{PLANTS|id=SACCO|taxon=Saccoloma|accessdate=26 October 2015}}</ref>
==Phylogeny== ''Saccoloma'' includes ''Orthiopteris'' in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), with a total of 18 species.<ref name=PPGI/> {{As of|2019|August}}, Plants of the World Online accepts the following species (which may be based on a wider circumscription of the genus). Synonyms are taken from the ''Checklist of Ferns and Lycophytes of the World''.<ref name=CFLW/>
Phylogeny of ''Saccoloma''<ref>{{cite journal |last1=Nitta |first1=Joel H. |last2=Schuettpelz |first2=Eric |last3=Ramírez-Barahona |first3=Santiago |last4=Iwasaki |first4=Wataru |display-authors=et al. |year=2022 |title=An Open and Continuously Updated Fern Tree of Life |journal=Frontiers in Plant Science |volume=13 |issue= |article-number= 909768| doi=10.3389/fpls.2022.909768 |pmid= 36092417|pmc= 9449725|bibcode= |doi-access=free}}</ref><ref>{{cite web |last1= |first1= |display-authors=et al. |year=2024 |title=Tree viewer: interactive visualization of FTOL |url=https://fernphy.github.io/viewer.html |version=FTOL v1.7.0 (GenBank release 261) |access-date=1 May 2025}}</ref> {{Clade | style=font-size:90%;line-height:80% |1={{clade |label1=''Orthiopteris'' |1={{clade |1=''O. campylura'' <small>(Kunze) Copel.</small> (syn. ''Saccoloma campylurum'' <small>(Kunze) Mett.</small>; ''Saccoloma caudatum'' <small>Copel.</small>; ''Saccoloma kingii'' <small>(Bedd.) G.B.Nair</small>) |2=''O. ferulacea'' <small>(T. Moore) Copel.</small> (syn. ''Saccoloma ferulaceum'' <small>(T.Moore) R.M.Tryon & A.F.Tryon</small>) }} |label2=''Saccoloma'' |2={{clade |1={{clade |1=''S. galeottii'' <small>(Fée) A.Rojas</small> |2=''S. sunduei'' <small>A.Rojas</small> }} |2={{clade |1={{clade |1=''S. domingense'' <small>(Spr.) Prantl</small> |2=''S. elegans'' <small>Kaulf.</small> (syn. ''Saccoloma chartaceum'' <small>G.B.Nair</small>) }} |2={{clade |1=''S. inaequale'' <small>(Kunze) Mett.</small> |2={{clade |1=''S. brasiliense'' <small>(C.Presl) Mett.</small> |2=''S. nigrescens'' <small>(Mett.) A.Rojas</small> }} }} }} }} }} }}
Other species include: *''Saccoloma acuminatum'' <small>(Rosenst.) Christenh.</small> (syn. ''Orthiopteris acuminata''<small>(Rosenst.) Copel.</small>) *''Saccoloma cicutarioides'' <small>(Baker) Christenh.</small> (syn. ''Orthiopteris cicutarioides'' <small>(Baker) Copel.</small>) *''Saccoloma firmum'' <small>(Kuhn) C.Chr.</small> (syn. ''Orthiopteris firma'' <small>(Kuhn) Brownlie</small>) *''Saccoloma henriettae'' <small>(Baker) C.Chr.</small> (syn. ''Orthiopteris henriettae'' <small>(Baker) Copel.</small>) *''Saccoloma laxum'' <small>R.C.Moran & B.Øllg.</small> *''Saccoloma membranaceum'' <small>Mickel</small> *''Saccoloma moranii'' <small>A.Rojas</small> *''Saccoloma quadripinnatum'' <small>A.Rojas</small> *''Saccoloma samoense'' <small>(Hovenkamp & T.T.Luong) Christenh.</small> (syn. ''Orthiopteris samoensis'' <small>Hovenk. & T. T. Luong</small>) *''Saccoloma squamosum'' <small>R.C.Moran</small> *''Saccoloma tenue'' <small>(Brack.) Mett.</small> (syn. ''Orthiopteris tenuis'' <small>(Brack.) Brownlie</small>) *''Saccoloma trichophyllum'' <small>(Copel.) G.B.Nair</small> (syn. ''Orthiopteris trichophylla'' <small>Copel.</small>)
==References== {{Reflist|refs= <ref name=CFLW>{{Citation |mode=cs1|last1=Hassler |first1=Michael |last2=Schmitt |first2=Bernd |date=June 2019 |title=Checklist of Ferns and Lycophytes of the World |volume=8 |issue=2 |url=https://worldplants.webarchiv.kit.edu/ferns/ |access-date=2019-08-14 |name-list-style=amp }}</ref>
<ref name=POWO_30010104-2>{{cite web |title=''Saccoloma'' Kaulf. |work=Plants of the World Online |publisher=Royal Botanic Gardens, Kew|url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30010104-2 |access-date=2019-08-14 }}</ref>
<ref name=PPGI>{{citation |mode=cs1|author=PPG I |year=2016 |title=A community-derived classification for extant lycophytes and ferns |journal=Journal of Systematics and Evolution |volume=54 |issue=6 |pages=563–603 |doi=10.1111/jse.12229|s2cid=39980610 |doi-access=free }}</ref> }}
{{Commons}} {{Wikispecies}} {{Plant classification}} {{Fern classification}} {{Taxonbar|from=Q310724}} Category:Polypodiales Category:Fern genera Category:Pantropical flora Category:Taxonomy articles created by Polbot