{{Short description|Extinct genus of arthropod}} {{Speciesbox | fossil_range = Cambrian Stage 3, {{fossil range|518|ref=<ref>{{Cite journal|last1=Yang|first1=C.|last2=Li|first2=X.-H.|last3=Zhu|first3=M.|last4=Condon|first4=D. J.|last5=Chen|first5=J.|date=2018|title=Geochronological constraint on the Cambrian Chengjiang biota, South China|journal=Journal of the Geological Society|language=en|volume=175|issue=4|pages=659–666|doi=10.1144/jgs2017-103|bibcode=2018JGSoc.175..659Y |s2cid=135091168 |issn=0016-7649|url=http://nora.nerc.ac.uk/id/eprint/521412/1/2018-JGS-Chuan%20Yang%20et%20al.pdf}}</ref>}} | image = Retifacies fossil (cropped).png | image_caption = Fossil of the dorsal carapace | image_upright = 0.7 | parent_authority = Hou et al. 1989 | display_parents = 3 | genus = Retifacies | species = abnormalis | authority = Hou et al. 1989 }}
'''''Retifacies abnormalis''''' is an extinct arthropod that lived in the lower Cambrian (about 518 million years ago). Its fossil remains have been found in the Maotianshan Shales of Yunnan, China. It is a member of the Artiopoda, and closely related to ''Pygmaclypeatus''.
== Description == left|thumb|Reconstruction in various views. E=uniramous cephalic appendages F=biramous appendages Key: exopod (exo) exite (exi) protopodite (pt) podomere (pd) spine/setae (s) terminal claw (tc) left|thumb|Two life restorations showing variation in carapace ornamentation ''Retifacies'' was relatively large sized, reaching a carapace length of {{Convert|35|cm|ft}}, and a total length of {{Convert|55|cm|ft}}, including the extended antennae and tailspine. The head shield was broad and short, and appears to have lacked eyes. The antennae had 17 segments, which telescoped into each other, with spines present on the underside of the antennae at the boundary between the segments. Also present on the head were four closely spaced pairs of appendages, the first three of which were uniramous (single branched), while the last was biramous (two branched). The three uniramous appendages had six segments (podomeres), each ending with a terminal claw, with the endopod (lower leg-like branch) of the fourth biramous limb having a similar morphology. The trunk was divided up into 10 segments (tergites), all of approximately equal length and width, each of which were associated with pairs of biramous appendages. The endopods of these limbs had 5/6 podomeres with terminal claws, while the exopod (upper limb branch) comprised a semicircular lobe that bore over 12 paddle-shaped lamellae, the last of which bore setae (hair-like structures) at its edge. These limbs also bore exites, consisting of lamellae borne from the basal segment (basipod) of the appendage. The pygidium segment at the end of the body was proportionally large, and bore 5-6 pairs of biramous appendages. The segmented tailspine emerged from the underside of the pygidium.<ref name=":0" /> The carapace was covered in polygonal reticulated (net-like) ornamentation, which varied between specimens, corresponding to two morphotypes. Due to one morphotype only being found among the largest specimens the differences may be due to ontogeny.<ref>{{Cite journal |last1=Lin |first1=Weiliang |last2=Pates |first2=Stephen |last3=Losso |first3=Sarah R. |last4=Fu |first4=Dongjing |date=2024-02-19 |title=Intraspecific variation of early Cambrian (stage 3) arthropod Retifacies abnormalis revealed by morphometric analyses |journal=Frontiers in Ecology and Evolution |volume=12 |doi=10.3389/fevo.2024.1336365 |doi-access=free |issn=2296-701X}}</ref>
== Lifestyle == thumb|Life restoration Due to a lack of information about the gut morphology or contents, the reconstruction of the ecology of ''Retifacies'' is speculative. The authors of the 2022 redescription suggested that ''Retifacies'' was a benthic organism (living on the seafloor) that scavenged on the remains of soft-bodied organisms or organic matter.<ref name=":0" />
== Classification == The vaguely similar appearance to that of a trilobite originally led scholars to think that ''Retifacies'' was a possible relative of ''Helmetia'' or ''Naraoia'', two other primitive arthropods related to trilobites (Delle Cave and Simonetta 1991, Hou and Bergstrom 1997). Subsequently, another study on showed how ''Retifacies'' was related to other basal arthropods, as ''Emeraldella'' and ''Sidneyia,'' from the Burgess Shale in Canada, as part of Artiopoda, a placement that has been confirmed in subsequent studies. Hou and Bergstrom suggested a close relationship to the genus ''Squamacula.'' However, after a redescription of both ''Retifacies'' and ''Pygmaclypeatus'' from the same deposit in 2022, the latter is considered to be ''Retifacies'' closest known relative, based on the fact that both taxa share a multi-segmented tailspine, among other morphological similarities, with a close relation to ''Squamacula'' being rejected. The clade containing ''Retifacies''+''Pygmaclypeatus'' was either classified in Trilobitomorpha, or in a more basal position within Artiopoda.<ref>{{Cite journal |last1=Schmidt |first1=Michel |last2=Hou |first2=Xianguang |last3=Zhai |first3=Dayou |last4=Mai |first4=Huijuan |last5=Belojević |first5=Jelena |last6=Chen |first6=Xiaohan |last7=Melzer |first7=Roland R. |last8=Ortega-Hernández |first8=Javier |last9=Liu |first9=Yu |date=2022-03-28 |title=Before trilobite legs: Pygmaclypeatus daziensis reconsidered and the ancestral appendicular organization of Cambrian artiopods |journal=Philosophical Transactions of the Royal Society B: Biological Sciences |volume=377 |issue=1847 |article-number=20210030 |doi=10.1098/rstb.2021.0030 |pmc=8819370 |pmid=35125003}}</ref><ref name=":0">{{Cite journal |last1=Zhang |first1=Maoyin |last2=Liu |first2=Yu |last3=Hou |first3=Xianguang |last4=Ortega-Hernández |first4=Javier |last5=Mai |first5=Huijuan |last6=Schmidt |first6=Michel |last7=Melzer |first7=Roland R. |last8=Guo |first8=Jin |date=2022-08-19 |title=Ventral Morphology of the Non-Trilobite Artiopod Retifacies abnormalis Hou, Chen & Lu, 1989, from the Early Cambrian Chengjiang Biota, China |journal=Biology |language=en |volume=11 |issue=8 |page=1235 |doi=10.3390/biology11081235 |issn=2079-7737|doi-access=free |pmid=36009864 |pmc=9405172 }}</ref>
Implied weights parsimony phylogeny after Berks et al. 2023.<ref name=":02">{{Cite journal |last1=Berks |first1=H. O. |last2=Nielsen |first2=M. L. |last3=Flannery-Sutherland |first3=J. |last4=Nielsen |first4=A. T. |last5=Park |first5=T.-Y. S. |last6=Vinther |first6=J. |year=2023 |title=A possibly deep branching artiopodan arthropod from the lower Cambrian Sirius Passet Lagerstätte (North Greenland) |journal=Papers in Palaeontology |volume=9 |issue=3 |at=e1495 |doi=10.1002/spp2.1495 |s2cid=259253098 |doi-access=free}}</ref>{{clade|{{clade |1={{clade |1=''Thulaspis'' 100px |2=''Squamacula'' 80px }} |2={{clade |label1=Protosutura |1={{clade |1=''Zhiwenia'' 50px |2=''Australimicola'' |3=''Acanthomeridion'' 100px }} |2={{clade |label1=Vicissicaudata |1={{clade |1=''Sidneyia'' 100px |2={{clade |1=Cheloniellida 70px |2={{clade |1=''Emeraldella'' 70px |2={{clade |1=''Kodymirus'' 70px |2={{clade |1=''Eozetetes'' |2=Aglaspidida 100px }}}}}}}}}} |label2=Trilobitomorpha |2={{clade |1={{clade |1=''Retifacies'' 100px |2=''Pygmaclypeatus'' 100px }} |2={{clade |1=Xandarellida 70px |2={{clade |1=Nektaspida 70px |2={{clade |1=Conciliterga 100px |2={{clade |1=''Kwanyinaspis'' 70px |2={{clade |1=''Phytophilaspis'' 70px |2=Trilobita 100px }}}}}}}}}}}}}}}}}}|label1=Artiopoda}}
== References == {{reflist}}
==Sources== * Hou, Chen & Lu, 1989. Early Cambrian new arthropods from Chengjiang, Yunnan. Acta Palaeontologica Sinica, 28, 42–57. * Chen & Zhou, 1997. Biology of the Chengjiang fauna. Bulletin of the Natural Museum of Natural Science, 10, 11 – 106. * Hou & Bergstrom, 1997. Arthropods of the Lower Cambrian Chengjiang fauna, southwest China. Fossils and strata, 45, 116 pp. * Edgecombe & Ramskold, 1999b. Relationships of Cambrian Arachnata and the systematic position of Trilobita. Journal of Paleontology, 73, 263 – 287. * Hou, Bergstrom, Wang, Feng & Chen, 1999. The Chengjiang fauna. Exceptionally well-preserved animals from 530 million years ago. 170 pp. Yunnan Science and Technology Press, Kunming, Yunnan Province, China.
{{Artiopoda}} {{Taxonbar|from=Q3933748}}
Category:Artiopoda Category:Cambrian genus extinctions Category:Monotypic artiopod genera