{{Short description|Extinct clade of dinosaurs}} {{Redirect|Cerapoda |the genus of moths |Cerapoda (moth)}} {{Automatic taxobox | name = Neornithischians | fossil_range = Middle Jurassic–Late Cretaceous, {{fossilrange|170|66|earliest=199.6}} (possible Early Jurassic record) | image = Neornithischia Infobox Panoply.png | image_caption = Six neornithischians (top left to bottom right): ''Psittacosaurus'', ''Styracosaurus'', ''Thescelosaurus'', ''Kulindadromeus'', ''Hypacrosaurus'', ''Pachycephalosaurus'' | image_upright = 1.2 | taxon = Neornithischia | authority = Cooper, 1985 | subdivision_ranks = Subgroups | subdivision_ref = <ref name="fonseca2024"/> | subdivision = *{{extinct}}''Agilisaurus'' *{{extinct}}''Enigmacursor'' *{{extinct}}''Gongbusaurus? wucaiwanensis'' *{{extinct}}''Hexinlusaurus'' *{{extinct}}''Lesothosaurus''? *{{extinct}}''Minimocursor'' *{{extinct}}''Nanosaurus'' *{{extinct}}''Phyllodon'' *{{extinct}}''Pulaosaurus'' *{{extinct}}''Sanxiasaurus'' *{{extinct}}''Xiaosaurus'' *{{extinct}}''Yandusaurus'' *{{extinct}}'''Pyrodontia''' <small>Fonseca et al., 2024<ref name="fonseca2024">{{cite journal|last1=Fonseca|first1=A.O.|last2=Reid|first2=I.J.|last3=Venner|first3=A.|last4=Duncan|first4=R.J.|last5=Garcia|first5=M.S.|last6=Müller|first6=R.T.|year=2024|title=A comprehensive phylogenetic analysis on early ornithischian evolution|journal=Journal of Systematic Palaeontology|volume=22|issue=1|article-number=2346577|doi=10.1080/14772019.2024.2346577|bibcode=2024JSPal..2246577F }}</ref></small> **{{extinct}}''Kulindadromeus'' **{{extinct}}''Laosaurus? minimus'' **{{extinct}}'''Thescelosauridae''' **{{extinct}}'''Cerapoda''' <small>Sereno, 1986</small> ***{{extinct}}''Siluosaurus'' ***{{extinct}}'''Marginocephalia''' ***{{extinct}}'''Ornithopoda''' }}
'''Neornithischia''' ("new ornithischians") is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs.
Neornithischians include a variety of basal forms historically known as "hypsilophodonts", including the Thescelosauridae. As these taxa do not all form a monophyletic clade, the term 'small-bodied early diverging ornithischian' (SBEDO) has been used to refer to these as a collective group.<ref name="Fona">{{Cite journal |last1=Avrahami |first1=Haviv M. |last2=Makovicky |first2=Peter J. |last3=Tucker |first3=Ryan T. |last4=Zanno |first4=Lindsay E. |date=2024-07-09 |title=A new semi-fossorial thescelosaurine dinosaur from the Cenomanian-age Mussentuchit Member of the Cedar Mountain Formation, Utah |journal=The Anatomical Record |volume=307 |issue=12 |pages=3717–3781 |language=en |doi=10.1002/ar.25505 |issn=1932-8486|doi-access=free |pmid=38979930 }}</ref> In addition, there are derived forms classified in the groups Marginocephalia and Ornithopoda. The former includes clades Pachycephalosauria and Ceratopsia, while the latter typically includes ''Hypsilophodon'' and the more derived Iguanodontia.
==Classification== Neornithischia was first named by Cooper in 1985 and defined as "all genasaurians more closely related to ''Parasaurolophus walkeri'' than to ''Ankylosaurus magniventris'' or ''Stegosaurus stenops''".<ref name=Butler08>{{Cite journal |author1=Richard J. Butler |author2=Paul Upchurch |author3=David B. Norman |year=2008 |title=The phylogeny of the ornithischian dinosaurs |journal=Journal of Systematic Palaeontology |volume=6 |issue=1 |pages=1–40 |doi=10.1017/S1477201907002271 |bibcode=2008JSPal...6....1B |s2cid=86728076 }}</ref> In 2021, Neornithischia was given a formal definition under the PhyloCode: "The largest clade containing ''Iguanodon bernissartensis'' and ''Triceratops horridus'' but not ''Ankylosaurus magniventris'' and ''Stegosaurus stenops''."<ref name="madziaetal2021">Madzia D, Arbour VM, Boyd CA, Farke AA, Cruzado-Caballero P, Evans DC. 2021. The phylogenetic nomenclature of ornithischian dinosaurs. ''PeerJ'' '''9''':e12362 [https://doi.org/10.7717/peerj.12362%7Chttps://doi.org/10.7717/peerj.12362]</ref>
A 2017 study by Matthew G. Baron, David B. Norman, and Paul M. Barrett recovered the Early Jurassic taxon ''Lesothosaurus diagnosticus'' from Southern Africa as the most basal known member of Neornithischia – a position previously held by ''Stormbergia dangershoeki'' (a taxon considered by the authors to be an adult form of ''Lesothosaurus'' and therefore a junior subjective synonym). However, Baron et al. go on to state that this result is only poorly supported and that future studies will be needed in order to better resolve the base of the ornithischian tree.<ref>{{Cite journal |author1=Matthew G. Baron |author2=David B. Norman |author3=Paul M. Barrett |date=2016 |title=Postcranial anatomy of ''Lesothosaurus diagnosticus'' (Dinosauria: Ornithischia) from the Lower Jurassic of southern Africa: implications for basal ornithischian taxonomy and systematics |journal=Zoological Journal of the Linnean Society |doi=10.1111/zoj.12434|doi-access=free }}</ref>
'''Pyrodontia''' was named by André O. Fonseca and colleagues in 2024 to unite the members of '''Thescelosauridae''' and '''Cerapoda'''. Its name translates to "fire teeth" in reference to the rapid diversification of this clade during the Middle Jurassic.<ref name="fonseca2024"/> The clade is formally defined in the ''PhyloCode'' as "the smallest clade containing ''Ceratops montanus'', ''Iguanodon bernissartensis'', ''Pachycephalosaurus wyomingensis'', and ''Thescelosaurus neglectus''.<ref name="fonseca2024"/>
'''Cerapoda''' is the most diverse clade within Neornithischia. The name "Cerapoda" is a portmanteaux of "Ceratopsia" and "Ornithopoda". As the name suggests, the clade is divided into two groups: Ornithopoda ("bird-foot") and Marginocephalia ("fringed heads"). The latter group includes the Pachycephalosauria ("thick-headed lizards") and Ceratopsia ("horned faces"). The following taxonomy follows Richard J. Butler, Paul Upchurch and David B. Norman, 2008 (and Butler et al., 2011) unless otherwise noted.<ref name=Butler08 /><ref name=ButlerEtAl>{{Cite journal |author1=Richard J. Butler |author2=Jin Liyong |author3=Chen Jun |author4=Pascal Godefroit | year = 2011 | title = The postcranial osteology and phylogenetic position of the small ornithischian dinosaur ''Changchunsaurus parvus'' from the Quantou Formation (Cretaceous: Aptian–Cenomanian) of Jilin Province, north-eastern China | journal = Palaeontology | volume = 54 | issue = 3 | pages = 667–683 | doi = 10.1111/j.1475-4983.2011.01046.x | doi-access = |bibcode=2011Palgy..54..667B }}</ref>
Cerapoda was first named by Sereno in 1986 and defined by him as "''Parasaurolophus walkeri'' Parks, 1922, ''Triceratops horridus'' Marsh, 1889, their most recent common ancestor and all descendants".<ref name="Butler08"/> A similar clade Neornithopoda was tentatively proposed by David B. Norman to unite ceratopsians with advanced ornithopods in a 1984 paper.<ref name="norman84a">{{cite book |last1=Norman |first1=D.B. |editor1-last=Reif |editor1-first=W.E. |editor2-last=Westphal |editor2-first=F. |title=Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers |date=1984 |publisher=ATTEMPTO Verlag |pages=157–162 |chapter=A systematic reappraisal of the reptile order Ornithischia}}</ref> In 2021, Cerapoda was given a formal definition under the PhyloCode: "The smallest clade containing ''Iguanodon bernissartensis'' Boulenger in Beneden, 1881, ''Pachycephalosaurus wyomingensis'' (Gilmore, 1931), and ''Triceratops horridus'' Marsh, 1889."<ref name="madziaetal2021" />
The phylogenetic study of Fonseca and colleagues in 2024 recovered results similar to the previous analyses of Boyd and Herne and colleagues,<ref name="boyd2015">{{cite journal | title=The systematic relationships and biogeographic history of ornithischian dinosaurs | author=Clint A. Boyd | journal=PeerJ | year=2015 |volume=3 | article-number=e1523 | doi=10.7717/peerj.1523| pmc=4690359 | pmid=26713260 | doi-access=free }}</ref><ref name="herne2019">{{cite journal | title=New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999 | last1=Herne | first1=Matthew C. | last2=Nair | first2=Jay P. | last3=Evans | first3=Alistair R. | last4=Tait | first4=Alan M. | journal=Journal of Paleontology | year=2019 | volume=93 | issue=3 | pages=543–584 | doi=10.1017/jpa.2018.95| doi-access=free | bibcode=2019JPal...93..543H }}</ref> with thescelosaurids outside Ornithopoda and heterodontosaurids outside Neornithischia, while other aspects of relationships, like the placement of ''Changmiania'' or the grouping within Ornithopoda were novel results. Their equal-weights results are below.<ref name="fonseca2024"/>
{{clade|style=font-size:85%;line-height:85%; |label1=Ornithischia |1={{clade |1=''Asilisaurus'' |2={{clade |1=''Lutungutali'' |2=''Silesaurus'' |3={{clade |1=''Technosaurus'' |2={{clade |1=''Kwanasaurus'' |2={{clade |1=''Eucoelophysis'' |2={{clade |1=''Pisanosaurus'' |2={{clade |label1=Saphornithischia |1={{clade |1=Heterodontosauridae |2={{clade |1=''Eocursor'' |2={{clade |label1=Genasauria |1={{clade |label1=Thyreophora |1={{clade |1={{clade |1=''Laquintasaura'' |2=''Lesothosaurus'' }} |2={{clade |1={{clade |1=''Emausaurus'' |2=''Scutellosaurus'' }} |2={{clade |1=''Scelidosaurus'' |2={{clade |1=''Yuxisaurus'' |2={{clade |1=''Jakapil'' |2=Eurypoda }} }} }} }} }} |label2='''Neornithischia''' |2={{clade |1=''Agilisaurus'' |2={{clade |1=''Hexinlusaurus'' |2=''Sanxiasaurus'' |3={{clade |1=''Minimocursor'' |2=''Yandusaurus'' |3={{clade |1=''Nanosaurus'' |2={{clade |label1='''Pyrodontia''' |1={{clade |1={{clade |label1=Thescelosauridae |1={{clade |label1=Orodrominae |1={{clade |1=''Zephyrosaurus'' |2={{clade |1={{clade |1=''Orodromeus'' |2=''Oryctodromeus'' }} |2={{clade |1=''Albertadromeus'' |2=''Koreanosaurus'' }} }} }} |label2=Thescelosaurinae |2={{clade |1=''Yueosaurus'' |2={{clade |1=''Changmiania'' |2={{clade |1={{clade |1=''Haya griva'' |2=RTMP 2008.045.0002 }} |2={{clade |1={{clade |1=''Changchunsaurus'' |2=''Jeholosaurus'' }} |2={{clade |1=''Parksosaurus'' |2=''Thescelosaurus'' }} }} }} }} }} }} }} |2={{clade |1=''Kulindadromeus'' |2={{clade |label1='''Cerapoda''' |1={{clade |label1=Marginocephalia |1={{clade |1=Pachycephalosauria |2=Ceratopsia }} |label2=Ornithopoda |2={{clade |1=Hypsilophodontidae |2={{clade |label1=Iguanodontia |1={{clade |1=Rhabdodontomorpha |2={{clade |label1=Euiguanodontia |1={{clade |1={{clade |1=''Anabisetia'' |2=''Diluvicursor'' }} |2={{clade |1=''Gasparinisaura'' |2=''Notohypsilophodon'' }} |3=Elasmaria |4={{clade |1=''Iyuku'' |2=Dryomorpha }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }} }}
==References== {{Reflist}} * {{cite journal |author=Butler, R.J. |year=2005 |title=The 'fabrosaurid' ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho |journal=Zoological Journal of the Linnean Society |volume=145 |issue=2 |page=175-18 |doi=10.1111/j.1096-3642.2005.00182.x|doi-access=free }} * {{cite journal |author=Sereno, P.C. |year=1986 |title=Phylogeny of the bird-hipped dinosaurs (order Ornithischia) |journal=National Geographic Research |volume=2 |issue=2 |pages=234–56}}
==External links== * [http://www.britannica.com/eb/article-225959 Encyclopædia Britannica] * [https://web.archive.org/web/20060313160201/http://www.palaeos.com/Vertebrates/Units/320Ornithischia/500.html Palæos]
{{Ornithischia|O.}} {{Portal bar|Dinosaurs}} {{Taxonbar|from=Q379167}}
Category:Neornithischia Category:Dinosaur clades