{{Short description|Single-species lichen genus}} {{Use dmy dates|date=September 2024}} {{Speciesbox | image = | parent_authority = [[A.Massal.]] (1860) | taxon = Pliariona montagnei | authority = ([[Roelof Benjamin van den Bosch|Bosch]]) A.Massal. (1860) | synonyms_ref = <ref name="Species Fungorum synonymy"/> | synonyms = *''Graphis montagnei'' {{au|Bosch (1855)}} *''Lecanactis montagnei'' {{au|(Bosch) [[Nyl.]] (1858)}} *''Graphina montagnei'' {{au|(Bosch) [[Müll.Arg.]] (1880)}}<ref name="Müller 1880"/> *''Phaeographina montagnei'' {{au|(Bosch) Müll.Arg. (1882)}} *''Thecaria montagnei'' {{au|(Bosch) Staiger (2002)}}<ref name="Staiger 2002"/> }}

'''''Pliariona''''' is a fungal [[genus]] in the family [[Graphidaceae]].<ref name="CoL"/><ref name="Wijayawardene et al. 2022"/> It comprises the single species '''''Pliariona montagnei''''', a [[corticolous lichen|corticolous]] (bark-dwelling) [[script lichen]] widely distributed across various [[tropical]] and [[subtropical]] regions.

==Taxonomy==

''Pliariona'' was originally [[circumscription (taxonomy)|circumscribed]] by the Italian lichenologist [[Abramo Bartolommeo Massalongo]] in 1860,<ref name="Massalongo 1860"/> to contain a [[script lichen]] species that was originally named ''Graphis montagnei'' by the Dutch botanist [[Roelof Benjamin van den Bosch]].<ref name="Miquel 1856"/>

The genus ''Pliariona'' was reinstated in 2013 based on [[molecular phylogenetics]] studies that revealed distinct [[lineage (evolution)|lineages]] within the [[Graphidaceae]]. These studies showed that species previously grouped under the same genus due to [[morphology (biology)|morphological]] similarities were not always closely related.<ref name="Joshi et al. 2013"/> For example, ''Pliariona montagnei'' (formerly known as ''Thecaria montagnei'') and ''[[Thecaria quassicola]]'', despite their superficial resemblance, were found to be [[genetic distance|genetically distant]] and not [[Biological specificity#congener|congeneric]].<ref name="Joshi et al. 2013"/>

The reinstatement of ''Pliariona'' as a separate genus is supported by molecular evidence and key taxonomic characteristics that distinguish it from related genera. These distinguishing features include an {{lichengloss|epruinose}} apothecial {{lichengloss|disc}} (the spore-producing surface lacking a powdery coating) and a dark red [[hymenium]] (the fertile layer of the fungus).<ref name="Joshi et al. 2013"/>

==Description== ''Pliariona montagnei'' has a thallus that ranges in colour from olive to dull green, with a glossy appearance. The surface texture varies from smooth to warty, often reflecting the contours of the underlying bark {{lichengloss|substrate}}. The species is characterised by its numerous [[ascomata]] (fruiting bodies), which are {{lichengloss|sessile}} and {{lichengloss|lirellate}} in form. These reproductive structures are predominantly {{lichengloss|simple}}, though some display occasional branching, and measure up to 0.6&nbsp;mm by 0.5&nbsp;mm. The ascomata feature an open {{lichengloss|disc}} that is red and epruinose, surrounded by a well-developed {{lichengloss|exciple}}. This exciple is fully {{lichengloss|carbonised}} (blackened), measuring 130 to 250&nbsp;[[μm]], and is covered by a thick, unbroken {{lichengloss|thalline margin}}.<ref name="Joshi et al. 2013"/>

The [[hymenium]], which measures 150–200&nbsp;μm in height, is red-pigmented and densely {{lichengloss|inspersed}}. Within this layer, the [[ascus|asci]] (spore-bearing sacs) each contain a single [[ascospore]]. These ascospores are {{lichengloss|muriform}}, [[ellipsoid]]al to oval in shape, and measure 125–155&nbsp;μm by 25–40&nbsp;μm. Initially [[hyaline]] and non-[[amyloid (mycology)|amyloid]], the spores develop a brownish or [[mottling|mottled]] appearance as they mature. The species contains the compound [[isohypocrelline]], though the biological significance of this substance in ''P.&nbsp;montagnei'' remains unclear. The combination of these morphological and chemical characteristics distinguishes ''P.&nbsp;montagnei'' from other superficially similar species within the Graphidaceae.<ref name="Joshi et al. 2013"/>

==Habitat and distribution==

''Pliariona montagnei'' exhibits a wide distribution across various tropical and subtropical regions. The species has been documented in Australia, West Africa, Japan, the Philippines, India, Indonesia, the Solomon Islands, [[Vanuatu]], and [[Norfolk Island]]. It has more recently been recorded in Vietnam, specifically in [[Đắk Lắk province]], within [[Chư Yang Sin National Park]].<ref name="Joshi et al. 2013"/>

The species is [[corticolous lichen|corticolous]], growing on tree bark. In the Vietnamese location where it was observed, ''P.&nbsp;montagnei'' was found at an elevation of about 780 metres above sea level.<ref name="Joshi et al. 2013"/>

==References== {{Reflist|colwidth=30em|refs=

<ref name="CoL">{{Catalogue of Life |id=6SX5 |title=''Pliariona'' |access-date=13 September 2024}}</ref>

<ref name="Species Fungorum synonymy">{{cite web |title=GSD Species Synonymy. Current Name: ''Pliariona montagnei'' (Bosch) A. Massal., Atti Inst. Veneto Sci. lett., ed Arti, Sér. 3 5: 318 (1860) |url=https://www.speciesfungorum.org/GSD/GSDspecies.asp?RecordID=401667 |publisher=[[Species Fungorum]] |access-date=13 September 2024}}</ref>

<ref name="Wijayawardene et al. 2022">{{cite journal |display-authors=6 |last1=Wijayawardene |first1=N.N. |last2=Hyde |first2=K.D. |last3=Dai |first3=D.Q. |last4=Sánchez-García |first4=M. |last5=Goto |first5=B.T. |last6=Saxena |first6=R.K. |last7=Erdoğdu |first7=M. |last8=Selçuk |first8=F. |last9=Rajeshkumar |first9=K.C. |last10=Aptroot |first10=A. |last11=Błaszkowski |first11=J. |last12=Boonyuen |first12=N. |last13=da Silva |first13=G. |last14=de Souza |first14=F.A. |last15=Dong |first15=W. |last16=Ertz |first16=D. |last17=Haelewaters |first17=D. |last18=Jones |first18=E.B. |last19=Karunarathna |first19=S.C. |last20=Kirk |first20=P.M. |last21=Kukwa |first21=M. |last22=Kumla |first22=J. |last23=Leontyev |first23=D.V. |last24=Lumbsch |first24=H.T. |last25=Maharachchikumbura |first25=S.S.N. |last26=Marguno |first26=F. |last27=Martínez-Rodríguez |first27=P. |last28=Mešić |first28=A. |last29=Monteiro |first29=J.S. |last30=Oehl |first30=F. |last31=Pawłowska |first31=J. |last32=Pem |first32=D. |last33=Pfliegler |first33=W.P. |last34=Phillips |first34=A.J.L. |last35=Pošta |first35=A. |last36=He |first36=M.Q. |last37=Li |first37=J.X. |last38=Raza |first38=M. |last39=Sruthi |first39=O.P. |last40=Suetrong |first40=S. |last41=Suwannarach |first41=N. |last42=Tedersoo |first42=L. |last43=Thiyagaraja |first43=V. |last44=Tibpromma |first44=S. |last45=Tkalčec |first45=Z. |last46=Tokarev |first46=Y.S. |last47=Wanasinghe |first47=D.N. |last48=Wijesundara |first48=D.S.A. |last49=Wimalaseana |first49=S.D.M.K. |last50=Madrid |first50=H. |last51=Zhang |first51=G.Q. |last52=Gao |first52=Y. |last53=Sánchez-Castro |first53=I. |last54=Tang |first54=L.Z. |last55=Stadler |first55=M. |last56=Yurkov |first56=A. |last57=Thines |first57=M. |year=2022 |title=Outline of Fungi and fungus-like taxa – 2021 |journal=Mycosphere |volume=13 |issue=1 |pages=53–453 [161] |doi=10.5943/mycosphere/13/1/2 |url=https://www.researchgate.net/publication/358798332|hdl=1854/LU-8754813 |hdl-access=free }}</ref>

<ref name="Joshi et al. 2013">{{cite journal |last1=Joshi |first1=Santosh |last2=Jayalal |first2=Udeni |last3=Oh |first3=Soon-Ok |last4=Koh |first4=Young Jin |last5=Nguyen |first5=Thi Thuy |last6=Dzung |first6=Nguyen Anh |last7=Hur | first7=Jae-Seoun |title=New species and new records in the family Graphidaceae (Ascomycota: Ostropales ) from Vietnam |journal=The Lichenologist |volume=45 |issue=5 |year=2013 |doi=10.1017/S002428291300025X |pages=599–609}}</ref>

<ref name="Massalongo 1860">{{cite journal |last=Massalongo |first=A.B. |year=1860 |title=Esame comparativo di alcune genere di licheni |journal=Atti dell'Istituto Veneto Scienze |volume=5 |pages=313–337 |language=la}}</ref>

<ref name="Miquel 1856">{{cite book |last1=Miquel |first1=F.A.W. |last2=Van den Bosch |first2=R.B. |chapter=Lichenes |year=1856 |title=Plantae Junghuhnianae |volume=4 |pages=427–494 |url=https://www.biodiversitylibrary.org/page/612794 |language=la}}</ref>

<ref name="Müller 1880">{{cite journal |last=Müller |first=J. |year=1880 |title=Lichenologische Beiträge. X |journal=Flora (Regensburg) |volume=63 |issue=3 |pages=40–45 |language=la |url=https://www.biodiversitylibrary.org/page/65506}}</ref>

<ref name="Staiger 2002">{{cite book |last1=Staiger |last2=B. |year=2002 |title=Die Flechtennamilie Graphidaceae: Studien in Richtung einer natürlichen Gliederung |trans-title=The lichen family Graphidaceae: studies towards a natural organization |series=Bibliotheca Lichenologica |volume=85 |page=446 |publisher=J. Cramer |location=Berlin/Stuttgart |language=de |isbn=978-3-443-58064-3}}</ref>

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{{Taxonbar|from1=Q107289447 |from2=Q108263326 |from3=Q59587260 |from4=Q107901571 |from5=Q107708171 |from6=Q105498147 |from7=Q10695946}}

[[Category:Graphidaceae]] [[Category:Graphidales genera]] [[Category:Lichen genera]] [[Category:Taxa described in 1860]] [[Category:Taxa named by Abramo Bartolommeo Massalongo]]