The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets).[1] The edge of the leaf may be regular or irregular, and may be smooth or have hair, bristles, or spines. For more terms describing other aspects of leaves besides their overall morphology, see the leaf article.

The terms listed here all are supported by technical and professional usage, but they cannot be represented as mandatory or undebatable; readers must use their judgement. Authors often use terms arbitrarily, or coin them to taste, possibly in ignorance of established terms, and it is not always clear whether because of ignorance, or personal preference, or because usages change with time or context, or because of variation between specimens, even specimens from the same plant.[citation needed] For example, whether to call leaves on the same tree "acuminate", "lanceolate", or "linear" could depend on individual judgement, or which part of the tree one collected them from. The same cautions might apply to "caudate", "cuspidate", and "mucronate", or to "crenate", "dentate", and "serrate".

Another problem is to establish definitions that meet all cases or satisfy all authorities and readers. For example, it seems altogether reasonable to define a mucro as "a small sharp point as a continuation of the midrib", but it may not be clear how small is small enough, how sharp is sharp enough, how hard the point must be, and what to call the point when one cannot tell whether the leaf has a midrib at all. Various authors or field workers might come to incompatible conclusions, or might try to compromise by qualifying terms so vaguely that a description of a particular plant practically loses its value.

Use of these terms is not restricted to leaves, but may be applied to morphology of other parts of plants, e.g. bracts, bracteoles, stipules, sepals, petals, carpels or scales. Some of these terms are also used for similar-looking anatomical features on animals.

Leaf structure

Leaves of most plants include a flat structure called the blade or lamina supported by a network of veins, a petiole and a leaf base;[1] but not all leaves are flat, some are cylindrical.[citation needed] Leaves may be simple, with a single leaf blade, or compound, with several leaflets. Compound leaves may be pinnate with pinnae (leaflets) on both sides of a rachis (axis), or may be palmate with multiple leaflets arising from a single point.[1] Leaf structure is described by several terms that include:

ImageTermLatinDescription
bifoliolateHaving two leaflets[2]
bigeminateHaving two leaflets, each leaflet being bifoliolate
bipinnatebipinnatusThe leaflets are themselves pinnately-compound; twice pinnate[1] see imparipinnate and paripinnate
biternateWith three components, each with three leaflets[1]
imparipinnateWith an odd number of leaflets, pinnate with a terminal leaflet (the opposite of paripinnate)[1]
paripinnatePinnate with an even number of leaflets, lacking a terminal leaflet (the opposite of imparipinnate)[1]
even-pinnate
palmatepalmatusConsisting of leaflets all radiating from one point
simpleLeaf blade in one continuous section, without leaflets (not compound)[1]
ternateternatusWith three leaflets[1]
trifoliatetrifoliatus
trifoliolatetrifoliolatus
tripinnatetripinnatusPinnately compound in which each leaflet is itself bipinnate[1]

Leaf and leaflet shapes

Being one of the more visible features, leaf shape is commonly used for plant identification. Similar terms are used for other plant parts, such as petals, tepals, and bracts.

ImageTermLatinRefers principally toDescription
aciculariswhole leafSlender and pointed, needle-like
acuminatusleaf tipTapering to a long point in a concave manner
leaf tip or basePointed, having a short sharp apex angled less than 90°
apiculatusleaf tipTapering and ending in a short, slender point
aristatusleaf tipEnding in a stiff, bristle-like point
whole leafWith the blade shape different on each side of the midrib
attenuatusleaf baseHaving leaf tissue taper down the petiole to a narrow base and always having some leaf material on each side of the petiole
auriculatusleaf baseHaving ear-shaped appendages reaching beyond the attachment to the petiole or stem (in case of a seated leaf)
caudatusleaf tipTailed at the apex
,leaf tipHaving a rachis that extends beyond the leaf blade or leaflets into a long whip-like extension or cirrus (common in climbing palms); antonym: ecirrate
,cordatuswhole leaf or baseHeart-shaped, with the petiole or stem attached to the notch
cuneatusleaf baseTriangular, wedge-shaped, stem attaches to point
whole leafNarrowly triangular, widest on the opposite end from the stem, with the corners at that end rounded
cuspidatusleaf tipWith a sharp, elongated, rigid tip; tipped with a cusp
,deltoideuswhole leafShaped like the Greek letter delta; triangular with stem attached to side
digitatuswhole leafA palmately compound leaf with leaflets, similar to palmate[3]
leaf tipWithout a cirrus; antonym: cirrate
ellipticuswhole leafShaped like an ellipse (widest at mid-blade and with similar convex tapering towards apex and base), with a short or no point
emarginatusleaf tipSlightly indented at the tip
ensiformiswhole leafShaped like a sword; long and narrow with a sharp pointed tip
falcatuswhole leafSickle-shaped
fenestratusleaf surface featuresLarge openings through the leaf; see perforate; sometimes used to describe leaf epidermal windows
filiformiswhole leafThread- or filament-shaped
flabellatuswhole leafSemi-circular or fan-like
hastatuswhole leaf or baseSpear-shaped: pointed, with barbs, shaped like a spear point, with flaring pointed lobes at the base
lacinatuswhole leafVery deeply lobed with the lobes being very drawn out and often making the leaf look somewhat like a branch or a pitchfork
3-D shapeFlat (like most leaves)
lanceolatuswhole leafLong, wider in the middle, shaped like a lance tip
lineariswhole leafLong and very narrow like a blade of grass
lobatuswhole leafBeing divided by clefts; may be pinnately lobed or palmately lobed
loratuswhole leafHaving the form of a thong or strap
lyratuswhole leafShaped like a lyre, pinnately lobed leaf with an enlarged terminal lobe and smaller lateral lobes. See also List of lyrate plants.
mucronatusleaf tipEnding abruptly in a small sharp point as a continuation of the midrib[4]
multi + finderewhole leafCleft into many parts or lobes
obcordatuswhole leafHeart-shaped, stem attaches at the tapering end
oblanceolatuswhole leafMuch longer than wide and with the widest portion near the tip; reversed lanceolate
leaf baseAsymmetrical leaf base, with one side lower than the other
oblonguswhole leafHaving an elongated form with slightly parallel sides; roughly rectangular
obovatuswhole leafTeardrop-shaped, stem attaches to the tapering end; reversed ovate
whole leafReversed trullate; the longer sides meet at the base rather than the apex.
obtususleaf tip or baseBlunt, forming an angle > 90°
orbiculariswhole leafCircular
ovatuswhole leafEgg-shaped, with a tapering point and the widest portion near the petiole
palmatuswhole leafPalm-shaped, i.e. with lobes or leaflets stemming from the leaf base[5]
palmatuswhole leafLobes spread radially from a point[6]
palma + finderewhole leafPalm-shaped, having lobes with incisions that extend less than halfway toward the petiole
palma + partiriwhole leafHaving palmate lobes with incisions that extend over halfway toward the petiole
palma + secarewhole leafHaving palmate lobes with incisions that extend almost up, but not quite to the petiole.
panduratuswhole leafFiddle-shaped; obovate with a constriction near the middle.
pedatuswhole leafPalmate, with cleft lobes[7]
peltatusstem attachmentA round leaf where the petiole attaches near the center, e.g. a lotus leaf
perfoliatusstem attachmentWith the leaf blade surrounding the stem such that the stem appears to pass through the leaf
perforatusleaf surface featuresMany holes, or perforations, on leaf surface. Compare with fenestrate.
pinna + lobuswhole leafHaving lobes pinnately arranged on the central axis
pinna + finderewhole leafHaving lobes with incisions that extend less than halfway to the midrib
pinnatus + partiriwhole leafHaving lobes with incisions that extend more than halfway to the midrib
pinnatus + sectuswhole leafHaving lobes with incisions that extend almost to, or up to, the midrib
reniformiswhole leafShaped like a kidney, with an inward curve on one side
leaf tipWith a shallow notch in a round apex
,rhomboidaliswhole leafDiamond-shaped
rotundifoliusleaf tip or baseCircular, no distinct point
semiterete3-D shapeRounded on one side and flat on the other
sagittatuswhole leafArrowhead-shaped with the lower lobes folded, or curled downward
spathulatuswhole leafSpoon-shaped; having a broad flat end which tapers to the base
-shapedhastatuswhole leafSee hastate.
subobtususleaf tip or baseSomewhat blunted; neither blunt nor sharp
subulatusleaf tipAwl-shaped with a tapering point
3-D shapeCylindrical with a circular or distorted circular cross-section and a single surface wrapping around it with no grooves or ridges. Subterete means the leaves are not completely terete, as seen in various lichens and succulents.
whole leafShaped like a masonry trowel
truncatusleaf tip or baseWith a squared-off end
undulatus3-D shapeWave-like
unifoliatuscompound leavesWith a single leaflet; it is distinct from a simple leaf by the presence of two abscission layers and often by petiolules and stipels.

Edge

Leaf margins (edges) are frequently used in visual plant identification because they are usually consistent within a species or group of species, and are an easy characteristic to observe. Edge and margin are interchangeable in the sense that they both refer to the outside perimeter of a leaf.

ImageTermLatinDescription
entireForma
integra
Even; with a smooth margin; without toothing
ciliateciliatusFringed with hairs
crenatecrenatusWavy-toothed; dentate with rounded teeth
crenulatecrenulatusFinely crenate
crispedcrispusCurly
dentatedentatus

Toothed; may be coarsely dentate, having large teeth

or glandular dentate, having teeth which bear glands

denticulatedenticulatusFinely toothed
doubly serrateduplicato-dentatusEach tooth bearing smaller teeth
serrateserratusSaw-toothed; with asymmetrical teeth pointing forward
serrulateserrulatusFinely serrate
sinuatesinuosusWith deep, wave-like indentations; coarsely crenate
lobatelobatusIndented, with the indentations not reaching the center
undulateundulatusWith a wavy edge, shallower than sinuate
spiny or pungentspiculatusWith stiff, sharp points such as thistles

Leaf folding

Leaves may also be folded, sculpted or rolled in various ways. If the leaves are initially folded in the bud, but later unrolls it is called vernation, ptyxis is the folding of an individual leaf in a bud.

ImageTermLatinDescription
or keeledcarinatusWith a longitudinal ridge, keel-shaped
Folded upwards, with the surfaces close to parallel
Forming a hood, margins and tip curved downward
Rolled upwards (towards the adaxial surface)
plicatusWith parallel folds
Folded downwards, with the surfaces close to parallel
Rolled downwards (towards the abaxial surface)
Opposing left and right halves of lamina folded along longitudinal axis, with one half rolled completely within the other

Latin descriptions

The Latin word for 'leaf', folium, is neuter. In descriptions of a single leaf, the neuter singular ending of the adjective is used, e.g. folium lanceolatum 'lanceolate leaf', folium lineare 'linear leaf'. In descriptions of multiple leaves, the neuter plural is used, e.g. folia linearia 'linear leaves'. Descriptions commonly refer to the plant using the ablative singular or plural, e.g. foliis ovatis 'with ovate leaves'.[8]

See also

References

  1. ^ The Ultimate Visual Family Dictionary. New Delhi: DK Pub.. 2012. pp. 136–137. ISBN 978-0-1434-1954-9.
  2. ^ Radford, A.E.; Dickison, W.C.; Massey, J.R.; Bell, C.R. (1976). "Phytography - Morphological Evidence". Vascular Plant Systematics. Harper and Row, New York.
  3. ^ Index of Garden Plants, Mark Griffiths, Timber Press, 1992
  4. ^ Mucronate Archived 2016-03-03 at the Wayback Machine, Answers.com, from Roget's Thesaurus.
  5. ^ "palmate (adj. palmately)". GardenWeb Glossary of Botanical Terms. iVillage GardenWeb. 2006. Archived from the original on 13 February 2009. Retrieved 19 October 2008.
  6. ^ Nelson, Randal C. (2009 [2012]). "Leaf description glossary". University of Rochester. Archived from the original on 1 August 2020. Retrieved 2 November 2017.
  7. ^ Nicholson, George; Garret, John; Trail, JWH (1887–1889). The Illustrated Dictionary of Gardening. London: L. Upcott Gill. ISBN 978-1436572316.
  8. ^ Stearn (2004), pp. 439–440.

Bibliography

  • Stearn, W.T. (2004). Botanical Latin. 4th (p/b) ed. Portland, Oregon: Timber Press. ISBN 978-0-7153-1643-6.
  • "Leaves", in Massey & Murphy (1996)
  • "Shapes", in Massey & Murphy (1996)
  • Massey, Jimmy R. & Murphy, James C. (1996). "Vascular plant systematics". NC Botnet. University of North Carolina at Chapel Hill. Retrieved 19 January 2016.