{{Short description|Genus of anhanguerid pterosaur from the Early Cretaceous}} {{Automatic taxobox | taxon = Mythunga | fossil_range = {{fossilrange|Albian|Albian|Albian}} | image = Mythunga.jpg | image_caption = Holotype skull and mandible of ''Mythunga'' (QM F18896) | authority = Molnar and Thulborn, 2008 | type_species = {{extinct}}'''''Mythunga camara''''' | type_species_authority = Molnar and Thulborn, 2008 }}
'''''Mythunga''''' is a potentially dubious genus of anhanguerid pterosaur from the late Early Cretaceous of Australia. Fossil remains of ''Mythunga'' dated back to the Albian stage of the Early Cretaceous, and the animal itself was found to be a close relative of another Australian anhanguerid called ''Ferrodraco''.
==Discovery and naming== ''Mythunga'' is known from a partial skull, holotype '''QM F18896''' found in April 1991 by Philip Gilmore in marine rocks of the late Albian-age Toolebuc Formation at Dunluce Station west of Hughenden, Queensland. Only the middle snout and corresponding parts of the lower jaws are known, including the rear of a left premaxilla, the lower parts of both maxillae, the rear dentaries and a right splenial. They were three-dimensionally preserved, associated in a chalk nodule. It represents a subadult individual.<ref name=MT08/> The fossil was prepared by Angela Hatch of the Queensland Museum, both by mechanical means and by an acid bath.<ref name="Pentland2018"/>
The type species ''Mythunga camara'' was named and described by Ralph Molnar and R. A. Thulborn in 2007/2008. The generic name is that of the constellation Orion in the local aboriginal language. The specific name, from Latin ''camera'', "room", refers to the camerate air spaces in the bone.<ref name=MT08>{{cite journal |last=Molnar |first=Ralph E. |author2=Thulborn, R.A. |year=2008 |title=An incomplete pterosaur skull from the Cretaceous of north-central Queensland, Australia |journal=Arquivos do Museu Nacional, Rio de Janeiro |volume=65 |issue=4 |pages=461–470 }}</ref>
''Mythunga'' was redescribed by Adele Pentland and Stephen Poropat in 2018, benefiting from further preparation of the specimen. At that time it was still the most completely known pterosaur of Australia. No more than twenty pterosaur fossils were known from that continent, most of them teeth and bone fragments.<ref name="Pentland2018">Adele H. Pentland; Stephen F. Poropat, 2018, "Reappraisal of ''Mythunga camara'' Molnar & Thulborn, 2007 (Pterosauria, Pterodactyloidea, Anhangueria) from the upper Albian Toolebuc Formation of Queensland, Australia". ''Cretaceous Research'' {{doi| 10.1016/j.cretres.2018.09.011}}</ref>
==Description== The wingspan of ''Mythunga'' was in 2007 estimated at {{convert|4.7|m|ft|sp=us}}.<ref name=MT08/> However, that was done under the assumption that the first preserved teeth were located on the left premaxilla which would imply that ''Mythunga'' was a relatively short-snouted form with a skull length of {{convert|50|to|80|cm|ft|sp=us}}. In 2018, it was concluded that they had been confused with a maxillary tooth and replacement tooth and that the skull length was likely between {{convert|80|cm|ft|sp=us}} and {{convert|1|m|ft|sp=us}}, with a corresponding wingspan of between {{convert|4|and|6|m|ft|sp=us}}.<ref name="Pentland2018"/> In 2022, Gregory S. Paul estimated its wingspan at {{cvt|5|m|ft}} and body mass at {{cvt|20|kg|lbs}}.<ref>{{Cite book|last=Paul|first=Gregory S.|title=The Princeton Field Guide to Pterosaurs|year=2022|publisher=Princeton University Press|pages=165|doi=10.1515/9780691232218|isbn=9780691232218|s2cid=249332375 }}</ref>
In 2007, two autapomorphies were suggested, unique derived traits. Firstly, the teeth in the rear dentary of the lower jaw were relatively tall (half the depth of the supporting bone at that point). Secondly, the three hindmost maxillary teeth were widely spaced.<ref name=MT08/> In 2018 however, the first trait was rejected, as it could not be reliably established where the dentary ended. Another autapomorphy was now proposed: the outer sides of the jaws are undulating because of the bulging tooth sockets.<ref name="Pentland2018"/>
The entire piece has a preserved length of {{convert|263|mm|in|sp=us}}. As with most pterosaurs, the snout was hollow, with a supporting internal boxwork of bone, as could be observed at breaks. The jugal bone apparently extended to a point below the front of the large skull opening, the nasoantorbital fenestra. There are at least eight teeth in each maxilla. That is also the minimal number in the dentary. The teeth are relatively short, conical and moderately recurved. Their cross-section is oval. They are widely spaced at equal distances.<ref name="Pentland2018"/>
==Classification== In 2007, ''Mythunga'' was provisionally thought to belong to a group of plesiomorphic, and thus possibly basal, pterodactyloids: the Archaeopterodactyloidea.<ref name=MT08/> In 2010, Kellner et al. placed ''Mythunga'' within the Anhangueridae.<ref>Kellner, A.W., Rich, T.H., Costa, F.R., Vickers-Rich, P., Kear, B.P., Walters, M. & Kool, L. 2010. "New isolated pterodactyloid bones from the Albian Toolebuc Formation (western Queensland, Australia) with comments on the Australian pterosaur fauna". ''Alcheringa'' '''34''': 219–230</ref> In 2018, it was recovered by a cladistic analysis in the Anhangueria.<ref name="Pentland2018"/>
The results of a phylogenetic analysis by Pentland et al. in 2019 have recovered ''Mythunga'' as a member of the family Ornithocheiridae, more precisely within the subfamily Ornithocheirinae as the sister taxon of ''Ferrodraco''.<ref name="Pentland2019">{{Cite journal|last1=Pentland|first1=Adele H.|last2=Poropat|first2=Stephen F.|last3=Tischler|first3=Travis R.|last4=Sloan|first4=Trish|last5=Elliott|first5=Robert A.|last6=Elliott|first6=Harry A.|last7=Elliott|first7=Judy A.|last8=Elliott|first8=David A.|date=December 2019|title=''Ferrodraco lentoni'' gen. et sp. nov., a new ornithocheirid pterosaur from the Winton Formation (Cenomanian–lower Turonian) of Queensland, Australia|journal=Scientific Reports|volume=9|issue=1|pages=13454|doi=10.1038/s41598-019-49789-4|pmid=31582757|pmc=6776501|bibcode=2019NatSR...913454P |issn=2045-2322}}</ref> However, a study made by Borja Holgado and Rubi Pêgas in 2020 had again recovered ''Mythunga'' within the family Anhangueridae, more precisely within the subfamily Tropeognathinae.<ref name="holgado2020">{{cite journal |last1=Holgado |first1=B. |last2=Pêgas |first2=R.V. |year=2020 |title=A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae |journal=Acta Palaeontologica Polonica |volume=65 |doi=10.4202/app.00751.2020|doi-access=free |hdl=2072/444907 |hdl-access=free }}</ref> In 2023, Richards et al. published a revised diagnosis of the related ''Thapunngaka'', where they also erected the clade Mythungini. The clade comprises all Australian tropeognathines, including ''Mythunga'', the type genus of said clade.<ref name="Richards2023">{{cite journal |last1=Richards |first1=Timothy |last2=Stumkat |first2=Paul |last3=Salisbury |first3=Steven |title=A second specimen of the pterosaur Thapunngaka shawi from the Lower Cretaceous (upper Albian) Toolebuc Formation of North West Queensland, Australia |journal=Cretaceous Research |date=6 October 2023 |volume=154 |doi=10.1016/j.cretres.2023.105740 |doi-access=free }}</ref>
{{col-begin|width=95%}} {{col-2}} '''Topology 1:''' First analysis by Pentland et al. (2019). {{clade| style=font-size:90%;line-height:90% |label1=Anhangueria |1={{clade |1=''Guidraco'' |2={{clade |1={{clade |1={{clade |1=''Brasileodactylus'' |2=''Ludodactylus'' }} |2={{clade |1=''Cearadactylus'' |label2=Ornithocheirae |2={{clade |label1=Anhangueridae |1={{clade |1=''Liaoningopterus'' |2=''Anhanguera'' }} |label2=Ornithocheiridae |2={{clade |1=''Tropeognathus'' |label2=Ornithocheirinae |2={{clade |1=''Coloborhynchus'' |2={{clade |1=''Ornithocheirus'' |2={{clade |1=''Ferrodraco'' |2='''''Mythunga''''' }} }} }} }} }} }} }} }} }} }} {{col-2}} '''Topology 2:''' Richards et al. (2023). {{clade|style=font-size:90%; line-height:90% |label1=Anhangueria |1={{clade |1=Hamipteridae |label2=Anhangueridae |2={{clade |1={{clade |1=Coloborhynchinae |2=Anhanguerinae }} |label2=Tropeognathinae |2={{clade |label1=Tropeognathini |1={{clade |1=''Siroccopteryx'' |2=''Tropeognathus'' |3=''Amblydectes'' }} |label2=Mythungini |2={{clade |1=''Ferrodraco'' |2=''Thapunngaka'' |3='''''Mythunga''''' }} }} }} }} }} {{col-end}}
In 2025, Pêgas suggested that ''Mythunga'' should be classified as a ''nomen dubium'', as he found none of the proposed diagnostic characters to be unique to this genus, and disagreed with using the tribe Mythungini. He therefore regarded the holotype specimen as an indeterminate tropeognathine. His phylogenetic analysis recovered ''Haliskia'' and ''Ferrodraco'' within Tropeognathinae, but ''Thapunngaka'' as the closest relative of ''Ornithocheirus'' outside Anhangueroidea, comprising the clade Targaryendraconia and the families Anhangueridae and Hamipteridae.<ref>{{cite journal|last1=Pêgas|first1=R.V.|year=2025|title=On the systematics and phylogenetic nomenclature of the Ornithocheiriformes (Pterosauria, Pteranodontoidea)|journal=Palaeontologia Electronica|volume=28|issue=2|at=a25|doi=10.26879/20|doi-access=free}}</ref>
==See also== * List of pterosaur genera * Timeline of pterosaur research
==References== {{Reflist}} {{Pterosauria|P.}} {{Taxonbar|from=Q2601308}} {{Portal bar|Paleontology|Australia}}
Category:Early Cretaceous pterosaurs Category:Early Cretaceous reptiles of Australia Category:Fossil taxa described in 2008 Category:Pterosaurs of Australia Category:Taxa named by Ralph Molnar Category:Pteranodontoidea Category:Pterosaur genera