{{Short description|Genus of lichens}} {{Use dmy dates|date=December 2024}} {{Automatic taxobox | image = | taxon = Gilbertaria | authority = M.Svensson & Fryday (2022) | type_species = Gilbertaria contristans | type_species_authority = (Nyl.) M.Svensson & Fryday (2022) | subdivision_ranks = Species | subdivision = ''G. astrapeana''<br /> ''G. contristans''<br /> ''G. holomeloides''<br /> ''G. squalescens'' }}
'''''Gilbertaria''''' is a genus of lichen-forming fungi in the family Sphaerophoraceae,<ref name="CoL"/><ref name="2024 Outline"/> comprising four species of crustose lichens found primarily in arctic and alpine environments. The genus is characterised by its thin, crust-like growth on rocks and dead plant material, black button-like reproductive structures (apothecia), and frequent association with blue-green algae of the genus ''Stigonema''. Molecular studies published in 2022 revealed that several previously misunderstood species, formerly classified in different genera, formed a distinct evolutionary lineage. The genus is unique within its family as the only member with a crustose growth form, and is named in honour of the British lichenologist Oliver Gilbert. Most species occur in the Northern Hemisphere, particularly in areas where snow persists late into the growing season, with one species known from the Falkland Islands.
==Taxonomy==
The genus ''Gilbertaria'' was circumscribed in 2022 by Måns Svensson and Alan Fryday, who demonstrated through molecular phylogenetics analysis that several previously misunderstood lichen species formed a distinct evolutionary lineage within the family Sphaerophoraceae. The type species of the genus is ''Gilbertaria contristans'',<ref name="Svensson & Fryday 2022"/> which was originally described as ''Lecidea contristans'' by William Nylander in 1865.<ref name="Nylander 1865"/> Prior to the establishment of ''Gilbertaria'', the species now placed in this genus had been variously classified under different genera including ''Catillaria'', ''Lecidea'', and ''Toninia'', reflecting the historical uncertainty about their relationships.<ref name="Svensson & Fryday 2022"/>
The placement of ''Gilbertaria'' within the Sphaerophoraceae was initially surprising to researchers, as it is the only crustose genus in a family otherwise characterised by more complex growth forms. While other members of the family typically produce spores in specialised {{lichengloss|mazaedia}} (spore masses) and have different types of reproductive structures, ''Gilbertaria'' species have evolved a simpler form while retaining their evolutionary relationship to the family. The genus name combines "Gilbert-" (honouring the British lichenologist Oliver Gilbert) with "-aria" from ''Catillaria'', one of the genera where some of these species were previously classified.<ref name="Svensson & Fryday 2022"/>
==Description==
''Gilbertaria'' is a genus of crustose lichens that forms thin, crust-like growths on rocks and dead plant material. The lichen's main body (thallus) varies between species, from barely visible films to well-developed grey crusts made up of small {{lichengloss|granules}} or warty bumps. In rare cases, some species may develop small, round structures called soralia that help with reproduction.<ref name="Svensson & Fryday 2022"/>
A distinctive feature of all ''Gilbertaria'' species is their black, button-like reproductive structures (apothecia) that range from 0.2 to 1.5 mm in diameter. These structures lack any visible border or rim around their edges. When examined under a microscope, they reveal several unique characteristics: *long, relatively thick supporting filaments (paraphyses) that are 1.5–4 μm wide *specialised spore-producing cells (asci) that contain eight spores each *clear, two-celled spores that range from 5–18 μm in length.<ref name="Svensson & Fryday 2022"/>
''Gilbertaria'' species frequently grow alongside blue-green algae of the genus ''Stigonema'', which can be found between the lichen's granules or near its reproductive structures. While most species lack any distinct chemical compounds that would cause colour reactions with standard lichen spot tests, one species (''G. squalescens'') sometimes shows a faint grey or white glow under ultraviolet light.<ref name="Svensson & Fryday 2022"/>
==Species==
* ''Gilbertaria astrapeana'' {{au|Fryday & M.Svensson (2022)}} – Falkland Islands * ''Gilbertaria contristans'' {{au|(Nyl.) M.Svensson & Fryday (2022)}} – Northern and Central Europe and Greenland * ''Gilbertaria holomeloides'' {{au|(Nyl.) M.Svensson & Fryday (2022)}} – UK, Sweden, and Canada * ''Gilbertaria squalescens'' {{au|(Nyl.) M.Svensson & Fryday (2022)}} – North America, far eastern Russia, and Central and Northern Europe
Molecular studies suggest there may be additional undescribed species within the genus, as some specimens show genetic differences that do not correspond to currently recognised species. In particular, some specimens previously identified as ''G. contristans'' may represent a distinct, currently undescribed species.<ref name="Svensson & Fryday 2022"/>
==Habitat and ecology==
''Gilbertaria'' species are specialised lichens adapted to arctic and apline environments. They primarily inhabit areas where snow persists late into the growing season, particularly on rock walls, outcrops, and alpine tundra. These lichens show a strong preference for growing on dying or dead moss colonies, especially species of ''Andreaea'' and ''Gymnomitrion'', and are often found in seepage areas where water periodically flows over rocks.<ref name="Svensson & Fryday 2022"/>
The genus plays an important ecological role as a pioneer organism in these harsh environments. Their ability to colonise dying bryophytes suggests they may be adapted to utilising nutrients released by decomposing moss tissues. The lichens commonly form associations with filamentous cyanobacteria, particularly species of ''Stigonema'', which can be found growing between the lichen's structures or near its reproductive bodies. This association may provide additional nitrogen to support the lichen's growth in nutrient-poor environments.<ref name="Svensson & Fryday 2022"/>
Individual species show subtle variations in their habitat preferences. ''G. contristans'' typically occurs in relatively dry situations on dying bryophytes on sloping rock walls, while ''G. holomeloides'' favours more humid conditions and is often found growing directly on the ground. ''G. squalescens'' frequently grows alongside other alpine lichens such as ''Frutidella caesioatra'' and ''Lecanora leptacina''. Despite their specialised habitat requirements, these lichens can be easily overlooked due to their small size and inconspicuous appearance, suggesting they may be more widely distributed than current records indicate.<ref name="Svensson & Fryday 2022"/>
==References== {{Reflist|refs=
<ref name="CoL">{{Catalogue of Life |id=B2X33 |title=''Gilbertaria'' |access-date=29 December 2024}}</ref>
<ref name="Nylander 1865">{{cite journal |last=Nylander |first=W. |year=1865 |title=Adhuc novitiae quaedam Lichenum Europae variarum tribuum |journal=Flora (Regensburg) |volume=48 |pages=353–358 |language=la |url=https://www.biodiversitylibrary.org/page/56467}}</ref>
<ref name="Svensson & Fryday 2022">{{cite journal |last1=Svensson |first1=Måns |last2=Fryday | first2=Alan M. |title=''Gilbertaria'', a first crustose genus in the Sphaerophoraceae (Lecanoromycetes, Ascomycota) for ''Catillaria contristans'', ''Toninia squalescens'' and related species |journal=Mycological Progress |volume=21 |issue=10 |year=2022 |doi=10.1007/s11557-022-01838-5 |doi-access=free |page=e90|bibcode=2022MycPr..21...90S }}</ref>
<ref name="2024 Outline">{{cite journal |display-authors=6 |last1=Hyde |first1=K.D. |last2=Noorabadi |first2=M.T. |last3=Thiyagaraja |first3=V. |last4=He |first4=M.Q. |last5=Johnston |first5=P.R. |last6=Wijesinghe |first6=S.N. |last7=Armand |first7=A. |last8=Biketova |first8=A.Y. |last9=Chethana |first9=K.W.T. |last10=Erdoğdu |first10=M. |last11=Ge |first11=Z.W. |last12=Groenewald |first12=J.Z. |last13=Hongsanan |first13=S. |last14=Kušan |first14=I. |last15=Leontyev |first15=D.V. |last16=Li |first16=D.W. |last17=Lin |first17=C.G. |last18=Liu |first18=N.G. |last19=Maharachchikumbura |first19=S.S.N. |last20=Matočec |first20=N. |last21=May |first21=T.W. |last22=McKenzie |first22=E.H.C. |last23=Mešić |first23=A. |last24=Perera |first24=R.H. |last25=Phukhamsakda |first25=C. |last26=Piątek |first26=M. |last27=Samarakoon |first27=M.C. |last28=Selcuk |first28=F. |last29=Senanayake |first29=I.C. |last30=Tanney |first30=J.B. |last31=Tian |first31=Q. |last32=Vizzini |first32=A. |last33=Wanasinghe |first33=D.N. |last34=Wannasawang |first34=N. |last35=Wijayawardene |first35=N.N. |last36=Zhao |first36=R.L. |last37=Abdel-Wahab |first37=M.A. |last38=Abdollahzadeh |first38=J. |last39=Abeywickrama |first39=P.D. |last40=Abhinav |last41=Absalan |first41=S. |last42=Acharya |first42=K. |last43=Afshari |first43=N. |last44=Afshan |first44=N.S. |last45=Afzalinia |first45=S. |last46=Ahmadpour |first46=S.A. |last47=Akulov |first47=O. |last48=Alizadeh |first48=A. |last49=Alizadeh |first49=M. |last50=Al-Sadi |first50=A.M. |last51=Alves |first51=A. |last52=Alves |first52=V.C.S. |last53=Alves-Silva |first53=G. |last54=Antonín |first54=V. |last55=Aouali |first55=S. |last56=Aptroot |first56=A. |last57=Apurillo |first57=C.C.S. |last58=Arias |first58=R.M. |last59=Asgari |first59=B. |last60=Asghari |first60=R. |last61=Assis |first61=D.M.A. |last62=Assyov |first62=B. |last63=Atienza |first63=V. |last64=Aumentado |first64=H.D.R. |last65=Avasthi |first65=S. |last66=Azevedo |first66=E. |last67=Bakhshi |first67=M. |last68=Bao |first68=D.F. |last69=Baral |first69=H.O. |last70=Barata |first70=M. |last71=Barbosa |first71=K.D. |last72=Barbosa |first72=R.N. |last73=Barbosa |first73=F.R. |last74=Baroncelli |first74=R. |last75=Barreto |first75=G.G. |last76=Baschien |first76=C. |last77=Bennett |first77=R.M. |last78=Bera |first78=I. |last79=Bezerra |first79=J.D.P. |last80=Bhunjun |first80=C.S. |last81=Bianchinotti |first81=M.V. |last82=Błaszkowski |first82=J. |last83=Boekhout |first83=T. |last84=Bonito |first84=G.M. |last85=Boonmee |first85=S. |last86=Boonyuen |first86=N. |last87=Bortnikov |first87=F.M. |last88=Bregant |first88=C. |last89=Bundhun |first89=D. |last90=Burgaud |first90=G. |last91=Buyck |first91=B. |last92=Caeiro |first92=M.F. |last93=Cabarroi-Hernández |first93=M. |last94=Cai |first94=M.F. |last95=Cai |first95=L. |last96=Calabon |first96=M.S. |last97=Calaça |first97=F.J.S. |last98=Callalli |first98=M. |last99=Câmara |first99=M.P.S. |last100=Cano-Lira |first100=J. |last101=Cao |first101=B. |last102=Carlavilla |first102=J.R. |last103=Carvalho |first103=A. |last104=Carvalho |first104=T.G. |last105=Castañeda-Ruiz |first105=R.F. |last106=Catania |first106=M.D.V. |last107=Cazabonne |first107=J. |last108=Cedeño-Sanchez |first108=M. |last109=Chaharmiri-Dokhaharani |first109=S. |last110=Chaiwan |first110=N. |last111=Chakraborty |first111=N. |last112=Cheewankoon |first112=R. |last113=Chen |first113=C. |last114=Chen |first114=J. |last115=Chen |first115=Q. |last116=Chen |first116=Y.P. |last117=Chinaglia |first117=S. |last118=Coelho-Nascimento |first118=C.C. |last119=Coleine |first119=C. |last120=Costa-Rezende |first120=D.H. |last121=Cortés-Pérez |first121=A. |last122=Crouch |first122=J.A. |last123=Crous |first123=P.W. |last124=Cruz |first124=R.H.S.F. |last125=Czachura |first125=P. |last126=Damm |first126=U. |last127=Darmostuk |first127=V. |last128=Daroodi |first128=Z. |last129=Das |first129=K. |last130=Das |first130=K. |last131=Davoodian |first131=N. |last132=Davydov |first132=E.A. |last133=da Silva |first133=G.A. |last134=da Silva |first134=I.R. |last135=da Silva |first135=R.M.F. |last136=da Silva Santos |first136=A.C. |last137=Dai |first137=D.Q. |last138=Dai |first138=Y.C. |last139=de Groot |first139=M.D. |last140=De Kesel |first140=A. |last141=De Lange |first141=R. |last142=de Medeiros |first142=E.V. |last143=de Souza |first143=C.F.A. |last144=de Souza |first144=F.A. |last145=dela Cruz |first145=T.E.E. |last146=Decock |first146=C. |last147=Delgado |first147=G. |last148=Denchev |first148=C.M. |last149=Denchev |first149=T.T. |last150=Deng |first150=Y.L. |last151=Dentinger |first151=B.T.M. |last152=Devadatha |first152=B. |last153=Dianese |first153=J.C. |last154=Dima |first154=B. |last155=Doilom |first155=M. |last156=Dissanayake |first156=A.J. |last157=Dissanayake |first157=D.M.L.S. |last158=Dissanayake |first158=L.S. |last159=Diniz |first159=A.G. |last160=Dolatabadi |first160=S. |last161=Dong |first161=J.H. |last162=Dong |first162=W. |last163=Dong |first163=Z.Y. |last164=Drechsler-Santos |first164=E.R. |last165=Druzhinina |first165=I.S. |last166=Du |first166=T.Y. |last167=Dubey |first167=M.K. |last168=Dutta |first168=A.K. |last169=Elliott |first169=T.F. |last170=Elshahed |first170=M.S. |last171=Egidi |first171=E. |last172=Eisvand |first172=P. |last173=Fan |first173=L. |last174=Fan |first174=X. |last175=Fan |first175=X.L. |last176=Fedosova |first176=A.G. |last177=Ferro |first177=L.O. |last178=Fiuza |first178=P.O. |last179=Flakus |first179=A. |last180=Fonseca |first180=E.O.W. |last181=Fryar |first181=S.C. |last182=Gabaldón |first182=T. |last183=Gajanayake |first183=A.J. |last184=Gannibal |first184=P.B. |last185=Gao |first185=F. |last186=García-Sánchez |first186=D. |last187=García-Sandoval |first187=R. |last188=Garrido-Benavent |first188=I. |last189=Garzoli |first189=L. |last190=Gasca-Pineda |first190=J. |last191=Gautam |first191=A.K. |last192=Gené |first192=J. |last193=Ghobad-Nejhad |first193=M. |last194=Ghosh |first194=A. |last195=Giachini |first195=A.J. |last196=Gibertoni |first196=T.B. |last197=Gentekaki |first197=E. |last198=Gmoshinskiy |first198=V.I. |last199=Góes-Neto |first199=A. |last200=Gomdola |first200=D. |last201=Gorjón |first201=S.P. |last202=Goto |first202=B.T. |last203=Granados-Montero |first203=M.M. |last204=Griffith |first204=G.W. |last205=Groenewald |first205=M. |last206=Grossart |first206=H.-P. |last207=Gu |first207=Z.R. |last208=Gueidan |first208=C. |last209=Gunarathne |first209=A. |last210=Gunaseelan |first210=S. |last211=Guo |first211=S.L. |last212=Gusmão |first212=L.F.P. |last213=Gutierrez |first213=A.C. |last214=Guzmán-Dávalos |first214=L. |last215=Haelewaters |first215=D. |last216=Haituk |first216=H. |last217=Halling |first217=R.E. |last218=He |first218=S.C. |last219=Heredia |first219=G. |last220=Hernández-Restrepo |first220=M. |last221=Hosoya |first221=T. |last222=Hoog |first222=S.D. |last223=Horak |first223=E. |last224=Hou |first224=C.L. |last225=Houbraken |first225=J. |last226=Htet |first226=Z.H. |last227=Huang |first227=S.K. |last228=Huang |first228=W.J. |last229=Hurdeal |first229=V.G. |last230=Hustad |first230=V.P. |last231=Inácio |first231=C.A. |last232=Janik |first232=P. |last233=Jayalal |first233=R.G.U. |last234=Jayasiri |first234=S.C. |last235=Jayawardena |first235=R.S. |last236=Jeewon |first236=R. |last237=Jerônimo |first237=G.H. |last238=Jin |first238=J. |last239=Jones |first239=E.B.G. |last240=Joshi |first240=Y. |last241=Jurjević |first241=Ž. |last242=Justo |first242=A. |last243=Kakishima |first243=M. |last244=Kaliyaperumal |first244=M. |last245=Kang |first245=G.P. |last246=Kang |first246=J.C. |last247=Karimi |first247=O. |last248=Karunarathna |first248=S.C. |last249=Karpov |first249=S.A. |last250=Kezo |first250=K. |last251=Khalid |first251=A.N. |last252=Khan |first252=M.K. |last253=Khuna |first253=S. |last254=Khyaju |first254=S. |last255=Kirchmair |first255=M. |last256=Klawonn |first256=I. |last257=Kraisitudomsook |first257=N. |last258=Kukwa |first258=M. |last259=Kularathnage |first259=N.D. |last260=Kumar |first260=S. |last261=Lachance |first261=M.A. |last262=Lado |first262=C. |last263=Latha |first263=K.P.D. |last264=Lee |first264=H.B. |last265=Leonardi |first265=M. |last266=Lestari |first266=A.S. |last267=Li |first267=C. |last268=Li |first268=H. |last269=Li |first269=J. |last270=Li |first270=Q. |last271=Li |first271=Y. |last272=Li |first272=Y.C. |last273=Li |first273=Y.X. |last274=Liao |first274=C.F. |last275=Lima |first275=J.L.R. |last276=Lima |first276=J.M.S. |last277=Lima |first277=N.B. |last278=Lin |first278=L. |last279=Linaldeddu |first279=B.T. |last280=Linn |first280=M.M. |last281=Liu |first281=F. |last282=Liu |first282=J.K. |last283=Liu |first283=J.W. |last284=Liu |first284=S. |last285=Liu |first285=S.L. |last286=Liu |first286=X.F. |last287=Liu |first287=X.Y. |last288=Longcore |first288=J.E. |last289=Luangharn |first289=T. |last290=Luangsa-ard |first290=J.J. |last291=Lu |first291=L. |last292=Lu |first292=Y.Z. |last293=Lumbsch |first293=H.T. |last294=Luo |first294=L. |last295=Luo |first295=M. |last296=Luo |first296=Z.L. |last297=Ma |first297=J. |last298=Madagammana |first298=A.D. |last299=Madhushan |first299=A. |last300=Madrid |first300=H. |last301=Magurno |first301=F. |last302=Magyar |first302=D. |last303=Mahadevakumar |first303=S. |last304=Malosso |first304=E. |last305=Malysh |first305=J.M. |last306=Mamarabadi |first306=M. |last307=Manawasinghe |first307=I.S. |last308=Manfrino |first308=R.G. |last309=Manimohan |first309=P. |last310=Mao |first310=N. |last311=Mapook |first311=A. |last312=Marchese |first312=P. |last313=Marasinghe |first313=D.S. |last314=Mardones |first314=M. |last315=Marin-Felix |first315=Y. |last316=Masigol |first316=H. |last317=Mehrabi |first317=M. |last318=Mehrabi-Koushki |first318=M. |last319=Meiras-Ottoni |first319=A.de |last320=Melo |first320=R.F.R. |last321=Mendes-Alvarenga |first321=R.L. |last322=Mendieta |first322=S. |last323=Meng |first323=Q.F. |last324=Menkis |first324=A. |last325=Menolli Jr |first325=N. |last326=Mikšík |first326=M. |last327=Miller |first327=S.L. |last328=Moncada |first328=B. |last329=Moncalvo |first329=J.M. |last330=Monteiro |first330=J.S. |last331=Monteiro |first331=M. |last332=Mora-Montes |first332=H.M. |last333=Moroz |first333=E.L. |last334=Moura |first334=J.C. |last335=Muhammad |first335=U. |last336=Mukhopadhyay |first336=S. |last337=Nagy |first337=G.L. |last338=Najam ul Sehar |first338=A. |last339=Najafiniya |first339=M. |last340=Nanayakkara |first340=C.M. |last341=Naseer |first341=A. |last342=Nascimento |first342=E.C.R. |last343=Nascimento |first343=S.S. |last344=Neuhauser |first344=S. |last345=Neves |first345=M.A. |last346=Niazi |first346=A.R. |last347=Nie |first347=Y. |last348=Nilsson |first348=R.H. |last349=Nogueira |first349=P.T.S. |last350=Novozhilov |first350=Y.K. |last351=Noordeloos |first351=M. |last352=Norphanphoun |first352=C. |last353=Nuñez Otaño |first353=N. |last354=O'Donnell |first354=R.P. |last355=Oehl |first355=F. |last356=Oliveira |first356=J.A. |last357=Oliveira Junior |first357=I. |last358=Oliveira |first358=N.V.L. |last359=Oliveira |first359=P.H.F. |last360=Orihara |first360=T. |last361=Oset |first361=M. |last362=Pang |first362=K.L. |last363=Papp |first363=V. |last364=Pathirana |first364=L.S. |last365=Peintner |first365=U. |last366=Pem |first366=D. |last367=Pereira |first367=O.L. |last368=Pérez-Moreno |first368=J. |last369=Pérez-Ortega |first369=S. |last370=Péter |first370=G. |last371=Pires-Zottarelli |first371=C.L.A. |last372=Phonemany |first372=M. |last373=Phongeun |first373=S. |last374=Pošta |first374=A. |last375=Prazeres |first375=J.F.S.A. |last376=Quan |first376=Y. |last377=Quandt |first377=C.A. |last378=Queiroz |first378=M.B. |last379=Radek |first379=R. |last380=Rahnama |first380=K. |last381=Raj |first381=K.N.A. |last382=Rajeshkumar |first382=K.C. |last383=Rajwar |first383=S. |last384=Ralaiveloarisoa |first384=A.B. |last385=Rämä |first385=T. |last386=Ramírez-Cruz |first386=V. |last387=Rambold |first387=G. |last388=Rathnayaka |first388=A.R. |last389=Raza |first389=M. |last390=Ren |first390=G.C. |last391=Rinaldi |first391=A.C. |last392=Rivas-Ferreiro |first392=M. |last393=Robledo |first393=G.L. |last394=Ronikier |first394=A. |last395=Rossi |first395=W. |last396=Rusevska |first396=K. |last397=Ryberg |first397=M. |last398=Safi |first398=A. |last399=Salimi |first399=F. |last400=Salvador-Montoya |first400=C.A. |last401=Samant |first401=B. |last402=Samaradiwakara |first402=N.P. |last403=Sánchez-Castro |first403=I. |last404=Sandoval-Denis |first404=M. |last405=Santiago |first405=A.L.C.M.A. |last406=Santos |first406=A.C.D.S. |last407=Santos |first407=L.A. |last408=Sarma |first408=V.V. |last409=Sarwar |first409=S. |last410=Savchenko |first410=A. |last411=Savchenko |first411=K. |last412=Saxena |first412=R.K. |last413=Schoutteten |first413=N. |last414=Selbmann |first414=L. |last415=Ševčíková |first415=H. |last416=Sharma |first416=A. |last417=Shen |first417=H.W. |last418=Shen |first418=Y.M. |last419=Shu |first419=Y.X. |last420=Silva |first420=H.F. |last421=Silva-Filho |first421=A.G.S. |last422=Silva |first422=V.S.H. |last423=Simmons |first423=D.R. |last424=Singh |first424=R. |last425=Sir |first425=E.B. |last426=Sohrabi |first426=M. |last427=Souza |first427=F.A. |last428=Souza-Motta |first428=C.M. |last429=Sri-indrasutdhi |first429=V. |last430=Sruthi |first430=O.P. |last431=Stadler |first431=M. |last432=Stemler |first432=J. |last433=Stephenson |first433=S.L. |last434=Stoyneva-Gaertner |first434=M.P. |last435=Strassert |first435=J.F.H. |last436=Stryjak-Bogacka |first436=M. |last437=Su |first437=H. |last438=Sun |first438=Y.R. |last439=Svantesson |first439=S. |last440=Sysouphanthong |first440=P. |last441=Takamatsu |first441=S. |last442=Tan |first442=T.H. |last443=Tanaka |first443=K. |last444=Tang |first444=C. |last445=Tang |first445=X. |last446=Taylor |first446=J.E. |last447=Taylor |first447=P.W.J. |last448=Tennakoon |first448=D.S. |last449=Thakshila |first449=S.A.D. |last450=Thambugala |first450=K.M. |last451=Thamodini |first451=G.K. |last452=Thilanga |first452=D. |last453=Thines |first453=M. |last454=Tiago |first454=P.V. |last455=Tian |first455=X.G. |last456=Tian |first456=W.H. |last457=Tibpromma |first457=S. |last458=Tkalčec |first458=Z. |last459=Tokarev |first459=Y.S. |last460=Tomšovský |first460=M. |last461=Torruella |first461=G. |last462=Tsurykau |first462=A. |last463=Udayanga |first463=D. |last464=Ulukapı |first464=M. |last465=Untereiner |first465=W.A. |last466=Usman |first466=M. |last467=Uzunov |first467=B.A. |last468=Vadthanarat |first468=S. |last469=Valenzuela |first469=R. |last470=Van den Wyngaert |first470=S. |last471=Van Vooren |first471=N. |last472=Velez |first472=P. |last473=Verma |first473=R.K. |last474=Vieira |first474=L.C. |last475=Vieira |first475=W.A.S. |last476=Vinzelj |first476=J.M. |last477=Tang |first477=A.M.C. |last478=Walker |first478=A. |last479=Walker |first479=A.K. |last480=Wang |first480=Q.M. |last481=Wang |first481=Y. |last482=Wang |first482=X.Y. |last483=Wang |first483=Z.Y. |last484=Wannathes |first484=N. |last485=Wartchow |first485=F. |last486=Weerakoon |first486=G. |last487=Wei |first487=D.P. |last488=Wei |first488=X. |last489=White |first489=J.F. |last490=Wijesundara |first490=D.S.A. |last491=Wisitrassameewong |first491=K. |last492=Worobiec |first492=G. |last493=Wu |first493=H.X. |last494=Wu |first494=N. |last495=Xiong |first495=Y.R. |last496=Xu |first496=B. |last497=Xu |first497=J.P. |last498=Xu |first498=R. |last499=Xu |first499=R.F. |last500=Xu |first500=R.J. |last501=Yadav |first501=S. |last502=Yakovchenko |first502=L.S. |last503=Yang |first503=H.D. |last504=Yang |first504=X. |last505=Yang |first505=Y.H. |last506=Yang |first506=Y. |last507=Yang |first507=Y.Y. |last508=Yoshioka |first508=R. |last509=Youssef |first509=N.H. |last510=Yu |first510=F.M. |last511=Yu |first511=Z.F. |last512=Yuan |first512=L.L. |last513=Yuan |first513=Q. |last514=Zabin |first514=D.A. |last515=Zamora |first515=J.C. |last516=Zapata |first516=C.V. |last517=Zare |first517=R. |last518=Zeng |first518=M. |last519=Zeng |first519=X.Y. |last520=Zhang |first520=J.F. |last521=Zhang |first521=J.Y. |last522=Zhang |first522=S. |last523=Zhang |first523=X.C. |last524=Zhao |first524=C.L. |last525=Zhao |first525=H. |last526=Zhao |first526=Q. |last527=Zhao |first527=H. |last528=Zhao |first528=H.J. |last529=Zhou |first529=H.M. |last530=Zhu |first530=X.Y. |last531=Zmitrovich |first531=I.V. |last532=Zucconi |first532=L. |last533=Zvyagina |first533=E. |title=The 2024 Outline of Fungi and fungus-like taxa |journal=Mycosphere |volume=15 |issue=1 |pages=5146–6239 [5255] |year=2024 |doi=10.5943/mycosphere/15/1/25 |url=https://www.researchgate.net/publication/385939154|hdl=1854/LU-8660838 |hdl-access=free }}</ref>
}}
{{Taxonbar|from=Q117771945}}
Category:Lecanorales Category:Lecanorales genera Category:Lichen genera Category:Taxa described in 2022 Category:Taxa named by Alan Fryday