{{Short description|Genus of haplolepideous mosses}} {{Automatic taxobox | image = Fissidens cristatus0.jpg | image_caption = ''Fissidens dubius'' | taxon = Fissidens | authority = Hedw.<ref name=bryonames/> | synonyms_ref = <ref name=bryonames>{{Cite bryonames|genus=Fissidens|access-date=6 December 2024}}</ref> | synonyms = {{Genus list | Conomitrium sect. Polypodiopsis | Müll.Hal. | Heterodon | Raf. | Fissidens sect. Pachyfissidens | Müll.Hal. | Fissidentella | Cardot | Moenkemeyera | Müll.Hal. | Nanobryum | Dixon | Octodiceras | Brid. | Osmundula | Rabenh. | Pachyfissidens | (Müll.Hal.) Limpr. | Polypodiopsis | (Müll.Hal.) A.Jaeger, nom. illeg. | Sainsburia | Dixon | Schistophyllum | Lindb., nom. illeg. | Skitophyllum | Bach.Pyl., nom. illeg. | Simplicidens | Herzog }} | subdivision_ranks = Species | subdivision = See text }}
'''''Fissidens''''' is the only genus of haplolepideous mosses (Dicranidae) in the family Fissidentaceae.<ref name=bryonames/> It contains over 480 species.
==Morphology== ''Fissidens'' is an acrocarpous genus characterized by the presence of vaginal lamina on their leaves, which may be palmately or pinnately arranged. Each leaf consists of one dorsal lamina, which is the farthest away from the central strand, and two vaginal lamina which clasp the central strand. They are connected via a single costa, though this costa is infrequently nearly absent.<ref name="WFO">{{cite web |title=Fissidens Hedw. |url=https://www.worldfloraonline.org/taxon/wfo-4000014805 |website=World Flora Online |access-date=24 February 2026}}</ref>
==Subordinate taxa== ''Fissidens'' is generally accepted to include multiple subgenera, though these are disputed from publication to publication. A 2004 publication on the infrageneric taxonomy included four: ''Aloma,'' ''Fissidens'', ''Octodiceras'', and ''Pachyfissidens''.<ref name="Pursell2004">{{cite journal |last1=Pursell |first1=Ronald |last2=Bruggeman-Nannenga |first2=Maria |title=A Revision of the Infrageneric Taxa of Fissidens |journal=The Bryologist |date=Spring 2004 |volume=107 |issue=1 |pages=1-20 |url=https://www.jstor.org/stable/3244563 |access-date=24 February 2026}}</ref> In 2018, the clade ''Fissidens'' was confirmed to be monophyletic and only three subgenera were recognized: ''Fissidens,'' ''Pachyfissidens,'' and the novel ''Neoamblyothallia''.<ref name="YMPEV 6171">{{cite journal |last1=Suzuki |first1=Tadashi |last2=Inoue |first2=Yuya |last3=Tsubota |first3=Hiromi |title=Molecular phylogeny of the genus Fissidens (Fissidentaceae, Bryophyta) and a refinement of the infrageneric classification |journal=Molecular Phylogenetics and Evolution |date=May 2018 |volume=127 |pages=190-202 |doi=10.1016/j.ympev.2018.05.020}}</ref>
Subsequent analysis of these clades found that the subgenera ''Aloma'' and ''Octodiceras'' as described by Pursell and Bruggeman-Nannenga in 2004, were strongly indicated to be monophyletic. Three sections described in 2018 were also indicated to be monophyletic, but none of the subgenera were.{{efn|name="monophyletic sections"|The monophyletic sections are ''Neoamblyothallia'' sect. ''Neoamblyothallia'', ''Pachyfissidens'' sect. ''Pachyfissidens'', and ''Fissidens'' sect. ''Aerofissidens''.}} This 2022 infrageneric analysis determined that all other subgenera and sections were nonmonophyletic.<ref name="Budke2022">{{cite journal |last1=Budke |first1=Jessica |last2=Patel |first2=Nikisha |last3=GoFlag Consortium |last4=Wienhold |first4=Mark |last5=Bruggeman-Nannenga |first5=Maria |title=Exploring morphological evolution in relation to habitat moisture in the moss genus Fissidens using molecular data generated from herbarium specimens |journal=Journal of Systematics and Evolution |date=October 2022 |volume=61 |issue=5 |pages=868-889 |doi=10.1111/jse.12926 |url=https://www.researchgate.net/publication/365021554_Exploring_morphological_evolution_in_relation_to_habitat_moisture_in_the_moss_genus_Fissidens_using_molecular_data_generated_from_herbarium_specimens |access-date=24 February 2026}}</ref>
==Included species== Selected species:<ref name=bryonames/> {{Div col}} {{Linked species list | Fissidens acacioides | Schrad. | Fissidens aciphyllus | Dixon | Fissidens acreanus | Irmsch. | Fissidens acrophilus | Brugg.-Nann. | Fissidens adianthoides | Hedw. | Fissidens afropapillosus | P.de la Varde | Fissidens aggestus | (Besch.) Paris | Fissidens ah-pengae | Brugg.-Nann. | Fissidens albolimbatus | Dixon | Fissidens allionii | Broth. | Fissidens allorgei | P.de la Varde | Fissidens alomoides | Müll.Hal. ex Dusén | Fissidens altisetus | Dixon | Fissidens amazonicus | Pursell | Fissidens amoenus | Müll.Hal. | Fissidens amplifolius | Dixon & P.de la Varde | Fissidens andicola | (Herzog) Brugg.-Nann. | Fissidens androgynus | Bruch | Fissidens angustelimbatus | Mitt. | Fissidens angustiusculus | Dixon & P.de la Varde | Fissidens angustus | Thwaites & Mitt. | Fissidens anisophyllus | Dixon | Fissidens annamensis | Paris & Broth. | Fissidens anomalus | Mont. | †Fissidens antiquus | Saporta | Fissidens antrophyi | Müll.Hal. | Fissidens aoraiensis | H.Whittier & H.A.Mill. | Fissidens aphelotaxifolius | Pursell | Fissidens appalachensis | R.H.Zander | Fissidens arboricola | Brugg.-Nann. | Fissidens arcticus | Bryhn | Fissidens argyroloma | P.de la Varde | Fissidens aristifer | Brugg.-Nann. | Fissidens arnigadhensis | Broth. ex S.S.Kumar | Fissidens arnoldi | R.Ruthe | Fissidens artsii | Brugg.-Nann. | Fissidens arunii | J.P.Srivast. & Nork. | Fissidens asplenioides | Hedw. | Fissidens asselii | Bizot | Fissidens autoicus | Thér. & Dixon | Fissidens axilliflorus | Thwaites & Mitt. | Fissidens azoricus | (P.de la Varde) Bizot | Fissidens badyinbarus | I.G.Stone | Fissidens bassilae | Alia, Brugg.-Nann. & Reeb | Fissidens beccarii | (Hampe) Broth. | Fissidens beckettii | Mitt. | Fissidens benitotanii | Z.Iwats., K.T.Yong & Tad.Suzuki | Fissidens berteroi | (Mont.) Müll.Hal. | Fissidens bessouensis | Corb. | Fissidens biformis | Mitt. | Fissidens bifrons | Schimp. ex Müll.Hal. | Fissidens bilaspurensis | Gangulee | Fissidens bilewskyi | P.de la Varde | Fissidens bistratosus | Brugg.-Nann. | Fissidens blechnoides | J.E.Beever | Fissidens bogoriensis | M.Fleisch. | Fissidens bogosicus | Müll.Hal. | Fissidens boninensis | Z.Iwats. | Fissidens borgenii | Hampe | Fissidens bourgaeanus | Besch. | Fissidens brachypus | Mitt. | Fissidens brassii | E.B.Bartram | Fissidens braunii | (Müll.Hal.) Dozy & Molk. | Fissidens brevidorsus | Dixon | Fissidens brevifrons | Mitt. | Fissidens brevinervis | Broth. | Fissidens brevipes | Besch. | Fissidens brindae | Pursell & B.H.Allen | Fissidens bryoides | Hedw. | Fissidens bushii | (Cardot & Thér.) Cardot & Thér. | Fissidens cagoui | Frank Müll., Pursell & Brugg.-Nann. | Fissidens canalae | Broth. & Paris | Fissidens capitulatus | Nog. | Fissidens capriviensis | Magill | Fissidens carnosus | Broth. | Fissidens celticus | Paton | Fissidens ceylonensis | Dozy & Molk. | Fissidens chioneurus | Müll.Hal. | Fissidens chiricahuensis | Pursell & B.H.Allen | Fissidens chitralensis | P.de la Varde | Fissidens circinalis | Mitt. | Fissidens closteri | Austin | Fissidens coacervatus | Brugg.-Nann. | Fissidens compienei | Broth. & Paris | Fissidens consociatus | Thér. | Fissidens costivelatus | Brugg.-Nann. <!--Fissidens crassinervis Sande Lac. Fissidens crassipes Wilson ex Bruch & Schimp. Fissidens crenulatus Mitt. Fissidens crispulus Brid. Fissidens crispus Mont. Fissidens cucullatus I.G.Stone Fissidens curticostatus Brugg.-Nann., K.Hyl. & Pursell Fissidens curvatoinvolutus Dixon Fissidens curvatus Hornsch. Fissidens cuspidifer Dixon Fissidens cuynetii Bizot Fissidens cyatheicola Brugg.-Nann. Fissidens cylindraceus Mitt. Fissidens cylindrothecus Pursell & J.Aguirre Fissidens dalamair H.Akiyama Fissidens daltoniifolius Müll.Hal. Fissidens danckelmannii Müll.Hal. Fissidens darutyi Schimp. ex Besch. Fissidens darwinianus Catches. & I.G.Stone Fissidens dasyphus Welw. & Duby Fissidens dealbatus Hook.f. & Wilson Fissidens decursivus Müll.Hal. Fissidens delicatulus Ångstr. Fissidens dendeliensis Paris & Broth. Fissidens dendrophilus Brugg.-Nann. & Pursell Fissidens dentatolimbatus Dixon Fissidens diaphanodontus (P.de la Varde) Bizot Fissidens diaphanus Bizot Fissidens dietrichiae Müll.Hal. Fissidens discolor Wilson ex Mitt. Fissidens dissitifolius Sull. Fissidens diversifolius Mitt. Fissidens dongensis Besch. --> | Fissidens dubius | P.Beauv. <!--Fissidens duttonii H.A.Mill. Fissidens ecuadorensis Pursell & Brugg.-Nann. --> | Fissidens elegans | Brid. <!-- Fissidens elizbrowniae B.C.Tan & Wongkuna Fissidens ellipticoides Brugg.-Nann. & Kürschner Fissidens elongatus Mitt. Fissidens enervis Sim Fissidens erectothorsbornei Tad.Suzuki Fissidens eremicus J.Guerra & J.A.Jiménez Fissidens ernestii Irmsch. Fissidens erosulus (Müll.Hal.) Paris Fissidens euryloma P.de la Varde Fissidens excurrentinervis R.S.Williams Fissidens exiguus Sull. --> | Fissidens exilis | Hedw. <!-- Fissidens extremiorientalis Ignatov & Ignatova Fissidens ezukanmae Brugg.-Nann. Fissidens faniensis (Besch.) Paris Fissidens fasciculatus Hornsch. Fissidens fautauae H.Whittier & H.A.Mill. Fissidens filiformis Z.Iwats. Fissidens firmus Mitt. Fissidens fissicaulis Müll.Hal. Fissidens flabellatus Hornsch. Fissidens flabellulus Thwaites & Mitt. Fissidens flaccidus Mitt. Fissidens fluitans Dixon Fissidens fontanus (Bach.Pyl.) Steud. Fissidens formosanus Nog. Fissidens fujiensis Tad.Suzuki & Z.Iwats. Fissidens fuscoviridis Thwaites & Mitt. Fissidens ganguleei Nork. Fissidens gardneri Mitt. Fissidens gaultieri Paris & Broth. Fissidens gedehensis M.Fleisch. Fissidens geijskesii Florsch. Fissidens geminiflorus Dozy & Molk. Fissidens georgianus Irmsch. Fissidens geppii M.Fleisch. Fissidens ghatensis K.M.Manjula, Manju & K.P.Rajesh Fissidens giraldii Broth. Fissidens gladiolus Mitt. Fissidens glaucissimus Welw. & Duby Fissidens gomae P.de la Varde & V.Leroy bis Fissidens goyazensis Broth. Fissidens gracilifolius Brugg.-Nann. & Nyholm Fissidens grainvillei P.de la Varde Fissidens grandifolius Broth. & P.de la Varde --> | Fissidens grandifrons | Brid. <!-- Fissidens granulatus (Geh. & Hampe) Paris Fissidens granulidens Brugg.-Nann. Fissidens griffithii Gangulee Fissidens grossiretis Müll.Hal. Fissidens guangdongensis Z.Iwats. & Z.H.Li Fissidens guianensis Mont. Fissidens gymnandrus Buse Fissidens gymnocarpus I.G.Stone Fissidens gymnogynus Besch. Fissidens gymnostomus Brugg.-Nann. Fissidens haconicus Müll.Hal. Fissidens hadii Banu-Fattah Fissidens hallianus (Sull. & Lesq.) Mitt. Fissidens harringtonii Brugg.-Nann. Fissidens henryae I.G.Stone Fissidens hildebrandtii Müll.Hal. Fissidens hirsutus Bizot Fissidens hoei Pursell Fissidens holleanus Dozy & Molk. Fissidens horizonticarpus Müll.Hal. Fissidens hornschuchii Mont. Fissidens hyalinus Wilson & Hook. --> | Fissidens hydropogon | Spruce ex Mitt. <!-- Fissidens hylogenes Dixon Fissidens hylophilus Irmsch. Fissidens hymenodon Besch. Fissidens imbricatus E.Britton & E.B.Bartram Fissidens inaequalilimbatus Paris & Broth. Fissidens inaequalis Mitt. Fissidens inaequiretis I.G.Stone Fissidens incertus Thér. & P.de la Varde Fissidens incisus Herzog Fissidens inclinis Müll.Hal. Fissidens inclusus Bizot & Dury Fissidens incognitus Gangulee --> | Fissidens inconspicuus | Mitt. | Fissidens incurvus | Starke ex Röhl. | Fissidens integerrimus | Mitt. | Fissidens intramarginatus | (Hampe) A.Jaeger | Fissidens intromarginatulus | E.B.Bartram | Fissidens involutus | Wilson ex Mitt. | Fissidens iwatsukii | Brugg.-Nann. | Fissidens jaiorum | H.Whittier & H.A.Mill. | Fissidens jansenii | Sérgio & Pursell | Fissidens javanicus | Dozy & Molk. | Fissidens jeffreyi | Bizot | Fissidens jonesii | Bizot | Fissidens jungermannioides | Griff. | Fissidens jungermanniopsis | Müll.Hal. | Fissidens juruensis | Broth. | Fissidens kalimpongensis | Gangulee | Fissidens kamiyamae | Tad.Suzuki & Z.Iwats. | Fissidens kammadensis | Manju, K.P.Rajesh & Madhus. | Fissidens karataviensis | Samsel | Fissidens karwarensis | Dixon | Fissidens kiguchii | Tad.Suzuki | Fissidens kilaueae | Hoe & H.A.Crum | Fissidens kinabaluensis | Z.Iwats. | Fissidens kondoi | Sakurai | Fissidens kosaninii | Latzel | Fissidens kurzii | Müll.Hal. | Fissidens lachmannii | Bizot | Fissidens lagenarius | Mitt. | Fissidens lancifolius | E.B.Bartram | Fissidens laosianus | Dixon | Fissidens latelimbatus | Brugg.-Nann. | Fissidens lateriticus | K.M.Manjula, Manju & K.P.Rajesh | Fissidens lautokensis | Dixon | †Fissidens microstictus | Dixon & Luisier | Fissidens pokhrensis | Nork. ex S. S. Kumar | Fissidens polypodioides | Hedw. | Fissidens pusillus | (Wilson) Milde | Fissidens serratus | C.Müller | Fissidens taxifolius | Hedw. | Fissidens usambaricus | Broth. }} {{Div col end}}
==Notes== {{notelist}}
==References== {{Reflist}}
{{Taxonbar|from=Q140981}} {{Authority control}}
Category:Fissidentaceae Category:Moss genera Category:Taxonomy articles created by Polbot Category:Taxa named by Johann Hedwig