{{Short description|Family of spiders}} {{Automatic taxobox | name = | image = Cybaeus charlesi 308353247.jpg | image_caption = ''[[Cybaeus charlesi]]'' | image2 = Dirksia cinctipes - Ellyne Geurts - 166669008.jpeg | image2_caption = ''[[Dirksia cinctipes]]'' in Canada | fossil_range = {{Fossil range|Palaeogene|present}} | taxon = Cybaeidae | authority = [[Nathan Banks|Banks]], 1892 | range_map = | range_map_caption = | diversity = [[#Genera|24 genera]], 303 species }}
'''Cybaeidae''' is a [[Family (taxonomy)|family]] of [[spider]]s first described by [[Nathan Banks]] in 1892.<ref name="banks1892" /> There are small to large sized [[Entelegynae|entelegyne]] spiders, which are [[ecribellate]].<ref name="hedin2025" /> The [[diving bell spider]] or water spider ''Argyroneta aquatica'' was previously included in this family, but is now in the family [[Dictynidae]].<ref name="WSC_s8306" /><ref name="wheeler2016" /> The distribution of this species resembles closely a typical [[Holarctic realm|Holarctic]] biogeography, with all of their species being found in the northern hemisphere. Where they have a tendency to live beneath rocks or woody debris in shaded and cool forest habitats, but they can also be found in caves, ant nests, and moss on tree trunks.<ref name="hedin2025" /> They are morphologically defined as spiders which have three claws, a single row of tarsal [[trichobothria]], and posterior lateral [[Spinneret|spinnerets]] which lack a [[cribellum]].<ref name="hedin2025" />
Most of the cybaeid [[Genus|genera]] are very species rich, as they have a large number of very short-range endemic species.<ref name="hedin2025" />
==Genera== {{as of|2026|1}}, this family includes 24 genera and 303 species:<ref name="wsc" /> {{Div col}} * ''[[Allocybaeina]]'' <small>Bennett, 2020</small> – [[Japan]], [[United States]] * ''[[Blabomma]]'' <small>[[Ralph Vary Chamberlin|Chamberlin]] & Ivie, 1937</small> – [[Korea]], North America * ''[[Calymmaria]]'' <small>Chamberlin & Ivie, 1937</small> – North America * ''[[Cryphoeca]]'' <small>[[Tamerlan Thorell|Thorell]], 1870</small> – Japan, [[Turkey]], Europe, North America * ''[[Cryphoecina]]'' <small>Deltshev, 1997</small> – [[Montenegro]] * ''[[Cybaeina]]'' <small>Chamberlin & Ivie, 1932</small> – Korea, North America * ''[[Cybaeota]]'' <small>Chamberlin & Ivie, 1933</small> – [[Russia]], North America * ''[[Cybaeozyga]]'' <small>Chamberlin & Ivie, 1937</small> – United States * ''[[Cybaeus]]'' <small>[[Ludwig Carl Christian Koch|L. Koch]], 1868</small> – Asia, Europe, North America, [[Peru]] * ''[[Dirksia]]'' <small>Chamberlin & Ivie, 1942</small> – [[France]], North America * ''[[Ethobuella]]'' <small>Chamberlin & Ivie, 1937</small> – North America * ''[[Guicybaeus]]'' <small>Wang, Chen, Yang & Zhang, 2023</small> – [[China]] * ''[[Mastigusa]]'' <small>Menge, 1854</small> – [[Algeria]], [[Georgia (country)|Georgia]], [[Iran]], Turkey, Europe, Britain * ''[[Neocryphoeca]]'' <small>Roth, 1970</small> – United States * ''[[Neocybaeina]]'' <small>Bennett, 2023</small> – United States * ''[[Pseudocybaeota]]'' <small>Bennett, 2022</small> – United States * ''[[Rothaeina]]'' <small>Bennett, 2023</small> – United States * ''[[Sincybaeus]]'' <small>Wang & Zhang, 2022</small> – China, Japan * ''[[Siskiyu]]'' <small>[[:species:Marshal Hedin|Hedin]], Ramírez & Monjaraz-Ruedas, 2025</small> – United States * ''[[Symposia]]'' <small>[[Eugène Simon|Simon]], 1898</small> – [[Colombia]], [[Venezuela]] * ''[[Tuberta]]'' <small>Simon, 1884</small> – [[Azerbaijan]], Turkey, Europe, Britain * ''[[Vagellia]]'' <small>Simon, 1899</small> – [[Indonesia]] * ''[[Willisus]]'' <small>Roth, 1981</small> – United States * ''[[Yorima]]'' <small>Chamberlin & Ivie, 1942</small> – [[Cuba]], United States {{Div col end}}
A fossil genus is also placed in this family:<ref name="selden2001" /> * †''[[Vectaraneus]]'' <small>Selden, 2003</small> (Isle of Wight)
==References== <references> <ref name="wsc">{{cite web | url=https://wsc.nmbe.ch/lsid/urn:lsid:nmbe.ch:spiderfam:000022 | title=Family Cybaeidae Banks, 1892 | publisher=World Spider Catalog | doi=10.24436/2 | accessdate=2026-01-22}}</ref> <ref name="banks1892">{{cite journal| last=Banks| first=N.| year=1892| title=A classification of North American spiders| journal=The Canadian Entomologist| volume=24| issue=4| pages=88–97| doi=10.4039/Ent2488-4| url=https://zenodo.org/record/2460991}}</ref> <ref name="hedin2025">{{Cite journal |last1=Hedin |first1=Marshal |last2=Ramírez |first2=Martín J. |last3=Monjaraz-Ruedas |first3=Rodrigo |date=2025-02-06 |title=Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province |journal=ZooKeys |language=en |issue=1226 |pages=47–75 |bibcode=2025ZooK.1226...47H |doi=10.3897/zookeys.1226.140204 |issn=1313-2970 |pmc=11826229 |pmid=39959859 |doi-access=free}}</ref> <ref name="selden2001">{{cite journal |last1=Selden |first1=Paul A. |title=Eocene Spiders from the Isle of Wight With Preserved Respiratory Structures |journal=Palaeontology |date=July 2001 |volume=44 |issue=4 |pages=695–729 |doi=10.1111/1475-4983.00199 |bibcode=2001Palgy..44..695S |url=https://kuscholarworks.ku.edu/bitstream/handle/1808/8359/Selden_2001_Eocene%20spiders%20Isle%20of%20Wight.pdf?sequence=1}}</ref> <ref name="wheeler2016">{{Cite journal |last1=Wheeler |first1=Ward C. |last2=Coddington |first2=Jonathan A. |last3=Crowley |first3=Louise M. |last4=Dimitrov |first4=Dimitar |last5=Goloboff |first5=Pablo A. |last6=Griswold |first6=Charles E. |last7=Hormiga |first7=Gustavo |last8=Prendini |first8=Lorenzo |last9=Ramírez |first9=Martín J. |last10=Sierwald |first10=Petra |last11=Almeida-Silva |first11=Lina |last12=Alvarez-Padilla |first12=Fernando |last13=Arnedo |first13=Miquel A. |last14=Benavides Silva |first14=Ligia R. |last15=Benjamin |first15=Suresh P. |last16=Bond |first16=Jason E. |last17=Grismado |first17=Cristian J. |last18=Hasan |first18=Emile |last19=Hedin |first19=Marshal |last20=Izquierdo |first20=Matías A. |last21=Labarque |first21=Facundo M. |last22=Ledford |first22=Joel |last23=Lopardo |first23=Lara |last24=Maddison |first24=Wayne P. |last25=Miller |first25=Jeremy A. |last26=Piacentini |first26=Luis N. |last27=Platnick |first27=Norman I. |last28=Polotow |first28=Daniele |last29=Silva-Dávila |first29=Diana |last30=Scharff |first30=Nikolaj |last31=Szűts |first31=Tamás |last32=Ubick |first32=Darrell |last33=Vink |first33=Cor J. |last34=Wood |first34=Hannah M. |last35=Zhang |first35=Junxia |date=2016 |title=The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling |journal=Cladistics |volume=33 |issue=6 |pages=574–616 |doi=10.1111/cla.12182 |pmid=34724759 |s2cid=35535038 |name-list-style=amp |doi-access=free }}</ref> <ref name="WSC_s8306">{{citation |title=Taxon details ''Argyroneta aquatica'' (Clerck, 1757) |work=World Spider Catalog |publisher=Natural History Museum Bern |url=https://www.wsc.nmbe.ch/species/8306 |access-date=2020-12-21 }}</ref> </references>
==External links== * {{inaturalist taxon}}
{{Araneae}} {{Taxonbar|from=Q10028}}
[[Category:Cybaeidae| ]] [[Category:Araneomorphae families]]
{{Cybaeidae-stub}}