Columbellidae

{{Short description|Family of gastropods}} {{Automatic taxobox | taxon = Columbellidae | image = Amphissa columbiana 545049496.jpg | image_caption = Live individual of the columbellid species ''Amphissa columbiana'' | authority = Swainson, 1840<ref>Swainson (1840). ''Treat. Malacol.'' 152, 313.</ref> | synonyms_ref = <ref name="WoRMS" /> | synonyms = * Anachidae Golikov & Starobogatov, 1972 * Pyrenidae Suter, 1909 | subdivision_ranks = Genera | subdivision = See text | display_parents = 3 }}

The '''Columbellidae''', the dove snails or dove shells, are a family of small sea snails, marine gastropod mollusks in the order Neogastropoda.<ref name ="WoRMS">Bouchet, P.; Gofas, S. (2010). Columbellidae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=23009 on 2010-11-04</ref><ref name="Bouchet">Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. ''Classification and nomenclator of gastropod families''. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. {{ISBN|3-925919-72-4}}. {{ISSN|0076-2997}}. 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278</ref> [[File:Pardalinops testudinaria 001.jpg|thumb|''Pardalinops testudinarius'']]

These snails are characterized by their elongated, spiral shells and their carnivorous feeding habits. The family Columbellidae comprises numerous genera and species, many of which share similar shell morphologies and ecological niches.

==Distribution== Species in this family are found worldwide,<ref>{{Cite web |url=http://shells.tricity.wsu.edu/ArcherdShellCollection/Gastropoda/Columbariidae.html |title=Columbariidae (Pagoda Shells) |access-date=2007-12-25 |archive-date=2007-12-22 |archive-url=https://web.archive.org/web/20071222083507/http://shells.tricity.wsu.edu/archerdshellcollection/Gastropoda/Columbariidae.html |url-status=dead }}</ref> but are most abundant in shallow tropical water.

== Subfamilies == According to the taxonomy of the Gastropoda by Bouchet & Rocroi (2005), the family Columbellidae consists of two subfamilies: * Atiliinae Cossmann, 1901 - synonyms: Pyrenidae Suter, 1909; Anachidae Golikov & Starobogatov, 1972 * Columbellinae Swainson, 1840 * Columbellidae incertae sedis (temporary name) ** ''Columbellidae incertae sedis mariato'' (Pilsbry & H. N. Lowe, 1932)

==Genera== The family Columbellidae contains these genera: ;subfamily Atiliinae ;subfamily Columbellinae {{div col|colwidth=20em}} * † ''Ademitrella'' Ludbrook, 1941 * ''Aesopus'' Gould, 1860 * ''Alcira'' H. Adams, 1861 * ''Alia'' H. Adams & A. Adams, 1853 * ''Amphissa'' H. Adams & A. Adams, 1853 * ''Anachis'' H. Adams & A. Adams, 1853 * ''Antimitrella'' Powell, 1937 * † ''Antizafra'' H. J. Finlay, 1926 * ''Aoteatilia'' Powell, 1939 * ''Ascalista'' Drivas & Jay, 1990 * ''Astyris'' H. Adams and A. Adams, 1853 * † ''Auingeria'' Harzhauser & Landau, 2021 * ''Bathyglypta'' Pelorce, 2017 * † ''Bellacolumbella'' Harzhauser & Landau, 2021 * ''Bifurcium'' P. Fischer, 1884 * † ''Cigclirina'' Woodring, 1928 * ''Cilara'' Thiele, 1924 * ''Clathranachis'' Kuroda & Habe, 1954 * ''Clavistrombina'' Jung, 1989 * † ''Clinurella'' Sacco, 1890 * ''Columbella'' Lamarck, 1799 - type genus * † ''Columbellisipho'' Cossmann, 1889 * ''Conella'' Swainson, 1840 * ''Cosmioconcha'' Dall, 1913 * ''Costoanachis'' Sacco, 1890 * ''Cotonopsis'' Olsson, 1942 * ''Decipifus'' Olsson and McGinty, 1958 * † ''Defensina'' Harzhauser & Landau, 2021 * ''Euplica'' Dall, 1889 * ''Eurypyrene'' Woodring, 1928 * ''Euspiralta'' K. Monsecour & Pelorce, 2013 * ''Exaesopus'' de Maintenon, 2019 * ''Exomilopsis'' Powell, 1964 * ''Falsuszafrona'' Pelorce, 2020 * ''Gatliffena'' Iredale, 1929 * ''Glyptanachis'' Pilsbry & Lowe, 1932 * ''Graphicomassa'' Iredale, 1929 * ''Indomitrella'' Oostingh, 1940 * ''Ithiaesopus'' Olsson & Harbison, 1953 * † ''Juliamitrella'' L. D. Campbell, 1993 * ''Liratilia'' Finlay, 1927 * ''Macrozafra'' Finlay, 1927 * ''Maintenonia'' K. Monsecour & D. Monsecour, 2024 * † ''Martaia'' Harzhauser & Landau, 2021 * ''Mazatlania'' Dall, 1913 * ''Metanachis'' Thiele, 1924 * ''Metulella'' Gabb, 1873 * ''Microcithara'' P. Fischer, 1884 * ''Minimanachis'' Pelorce, 2020 * ''Mitrella'' Risso, 1826 * ''Mitropsis'' Pease, 1868 * ''Mokumea'' Habe, 1991 * ''Nassarina'' Dall, 1889 * ''Nitidella'' Swainson, 1840 * ''Nodochila'' Rehder, 1980 * † ''Orthurella'' Sacco, 1890 * ''Parametaria'' Dall, 1916 * ''Pardalinops'' De Maintenon, 2008 * ''Parvanachis'' Radwin, 1968 * ''Parviterebra'' Pilsbry, 1904 * ''Paxula'' Finlay, 1927 * ''Pictocolumbella'' Habe, 1945 * ''Pleurifera'' Drivas & Jay, 1997 * ''Pseudamycla'' Pace, 1902 * ''Pyrene'' Röding, 1798 * ''Pyreneola'' Iredale, 1918 * ''Radwinia'' Shasky, 1970 * ''Ramoliva'' Cotton & Godfrey, 1932 * ''Redfernella'' Espinosa & Ortea, 2020 * ''Retizafra'' Hedley, 1913 * ''Rhombinella'' Radwin, 1968 * ''Ruthia'' Shasky, 1970 * ''Salitra'' Marincovich, 1973 * † ''Scabrella'' Sacco, 1890 * ''Seminella'' Pease, 1867 * ''Sincola'' Olsson & Harbison, 1953 * ''Smithena'' Boyer, Pelorce & Gori, 2022 * ''Speirazafra'' K. Monsecour & Raines, 2024 * ''Steironepion'' Pilsbry and Lowe, 1932 * ''Strombina'' Mörch, 1852 * † ''Strombinella'' Dall, 1896 * ''Sulcomitrella'' Kuroda, Habe & Oyama, 1971 * ''Suturoglypta'' Radwin, 1968 * † ''Thiarella'' Sacco, 1890 * † ''Trahaldia'' Pacaud, 2017 * † ''Turricolumbus'' Cossmann, 1901 * ''Turrijaumelia'' Sarasúa, 1975 * ''Urpiella'' Arias Ávila, 2021 * ''Zafra'' A. Adams, 1860 * ''Zafrona'' Iredale, 1916 * ''Zanassarina'' Pilsbry & H. N. Lowe, 1932 * ''Zella'' Iredale, 1924 * ''Zemitrella'' Finlay, 1927 * ''Zetekia'' Dall, 1918 {{div col end}}

==Synonyms== * ''Atilia'' H. Adams & A. Adams, 1853 - type genus of the subfamily Atiliinae: synonym of ''Anachis'' H. Adams & A. Adams, 1853 * ''Columbellopsis'' Bucquoy, Dautzenberg and Dollfus, 1882: synonym of ''Mitrella'' Risso, 1826 * ''Minipyrene'' Coomans, 1967: synonym of ''Mitromorpha'' P. P. Carpenter, 1865

==Gallery== <gallery> Zemitrella choava 482327614.jpg|Live ''Zemitrella choava'' Alia carinata 465040954.jpg|Live ''Alia carinata'' Alia carinata foot.png|Foot of live ''A. carinata'' Pardalinops testudinarius 457375157.jpg|Live ''Pardalinops testudinarius'' Mitrella ocellata 434014002.jpg|Two live ''Mitrella ocellata'' </gallery>

==References== {{Reflist}}

== External links == {{Commons category|Columbellidae}} * [https://pubs.usgs.gov/pp/0642/report.pdf Miocene Gastropods and Biostratigraphy of the Kern River Area, California; United States Geological Survey Professional Paper 642] {{PD-notice}} * [http://sciencepress.mnhn.fr/sites/default/files/articles/hd/z2017n2a2_hd.pdf Russini V., Fassio G., Modica M. V., deMaintenon M. J. & Oliverio M. 2017. ''An assessment of the genus Columbella Lamarck, 1799 (Gastropoda: Columbellidae) from eastern Atlantic''. Zoosystema 39 (2): 197-212] * [https://peerj.com/articles/13996.pdf deMaintenon, Marta, and Ellen E. Strong. "Molecular phylogeny of Columbellidae (Gastropoda: Neogastropoda)." PeerJ 10 (2022): e13996.]

Category:Columbellidae