Cochliopidae

{{Short description|Family of gastropods}} {{Automatic taxobox |taxon=Cochliopidae |image = Antrobia culveri.jpg |image_caption = A live individual of ''Antrobia culveri'' |authority = Tryon, 1866 |synonyms= Semisalsinae Giusti & Pezzoli, 1980 |display_parents= 3 }}

'''Cochliopidae''' is a family of small freshwater snails with gills and an operculum, aquatic gastropod mollusks.

Paludestrina d'Orbigny, 1840 is an archaic synonym,<ref name=":0">{{Cite book|title = Iconographie malacologique des animaux mollusques fluviatiles du Lac Tanganika|url = https://books.google.com/books?id=XEwYAAAAYAAJ|publisher = Impr. Crété|date = 1888-01-01|language = fr|first = Jules René|last = Bourguignat}}</ref> and has been placed on the Official Index of Rejected and Invalid Names by ICZN Opinion 2202.<ref name="WoRMS">MolluscaBase eds. (2021). MolluscaBase. Cochliopidae Tryon, 1866. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411706 on 2021-05-13</ref>

This family is in the superfamily Truncatelloidea and in the order Littorinimorpha (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

== Description == Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water.<ref name=":0" />

== 2005 taxonomy == The family Cochliopidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):<ref name="Bouchet 2005">{{Bouchet 2005}}</ref> *Cochliopinae Tryon, 1866 - synonyms: Mexithaumatinae D. W. Taylor, 1966, Paludiscalinae D. W. Taylor, 1966 *Littoridininae Thiele, 1928 *Semisalsinae Guiusti & Pezzoli, 1980 - synonym: Heleobiini Bernasconi, 1991

== Genera == Liu et al. (2001)<ref name="Liu 2001"/> have recognized 34 genera with more than 260 species within the subfamily Cochliopinae.<ref name="Liu 2001"/>

Strong et al. (2008)<ref name="Strong 2008">Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". ''Hydrobiologia'' 595: 149-166. {{hdl|10088/7390}} {{doi|10.1007/s10750-007-9012-6}}.</ref> have recognized 246 freshwater species within Cochliopidae.<ref name="Strong 2008"/>

Genera within the family Cochliopidae include:

subfamily Cochliopinae *''Aphaostracon'' F. G. Thompson, 1968 *''Coahuilix'' Taylor, 1966 *''Cochliopa'' Stimpson, 1865 - type genus of the family Cochliopidae<ref name="Bouchet 2005"/> *''Cochliopina'' Morrison, 1946 *''Emmericiella'' Pilsbry, 1909 *''Lobogenes'' Pilsbry & Bequaert, 1927 *''Mexithauma'' D. W. Taylor, 1966 *''Nanivitrea'' Thiele, 1927 *''Subcochliopa'' J. P. E. Morrison, 1946 *''Tepalcatia'' F. G. Thompson & Hershler, 2002

subfamily Littoridininae *''Antrobia'' Hubricht, 1971 - with the only species ''Antrobia culveri'' Hubricht, 1971<ref name="Kabat 1993">Kabat A. R. & Hershler R. (1993). "The prosobranch snail family Hydrobiidae (Gastropoda: Rissooidea): review of classification and supraspecific taxa". ''Smithsonian Contributions to Zoology'' '''547''': 1-94. [http://www.sil.si.edu/smithsoniancontributions/Zoology/pdf_hi/SCTZ-0547.pdf PDF].</ref> - Tumbling Creek cavesnail *''Littoridina'' Souleyet, 1852 * ''Pseudocaspia'' Starobogatov, 1972 *''Pseudotryonia'' Hershler, 2001<ref name="Hershler 2011"/> **''Pseudotryonia adamantina''<ref name="Hershler 2011"/> **''Pseudotryonia alamosae''<ref name="Hershler 2011"/> **''Pseudotryonia brevissima''<ref name="Hershler 2011"/> **''Pseudotryonia grahamae''<ref name="Hershler 2011"/> **''Pseudotryonia mica'' Hershler, Liu & Landye, 2011<ref name="Hershler 2011">Hershler R., Liu H.-P. & Landye J. J. (2011). "Two new genera and four new species of freshwater cochliopid gastropods (Rissooidea) from northeastern Mexico". ''Journal of Molluscan Studies'' '''77'''(1): 8-23. {{doi|10.1093/mollus/eyq033}}.</ref> **''Pseudotryonia pasajae'' Hershler, Liu & Landye, 2011<ref name="Hershler 2011"/>

subfamily Semisalsinae *''Heleobia'' Stimpson, 1865<ref name="Bouchet 2005"/><ref name="Kroll 2012">Kroll O., Hershler R., Albrecht C., Terrazas E. M., Apaza R., Fuentealba C., Wolff C. & Wilke T. (2012). "The endemic gastropod fauna of Lake Titicaca: correlation between molecular evolution and hydrographic history". ''Ecology and Evolution'' '''2'''(7): 1517-1530. {{doi|10.1002/ece3.280}}.</ref> *''Heleobops'' Thompson, 1968<ref name="Kroll 2012"/> *''Paludiscala'' D. W. Taylor, 1966 *''Semisalsa'' Radoman, 1974<ref name="Bouchet 2005"/><ref name="Kroll 2012"/>

subfamily ? (either Cochliopinae or Littoridininae) *''Aroapyrgus'' H. B. Baker, 1931 *''Balconorbis'' Hershler & Longley, 1986 *† ''Carinulorbis'' Yen, 1949 *''Chorrobius'' Hershler, Liu & Landye, 2011<ref name="Hershler 2011"/> *''Dyris'' Conrad, 1871 - it has extant species and also 26 species in Miocene Pebas Formation<ref name="Wesselingh 2006"/> *''Eremopyprgus'' Hershler, 1999<ref>Hershler R., Liu H.-P. & Landye J. J. (2002). "A new species of ''Eremopyprgus'' (Hydrobiidae: Cochliopinae) from the Chihuahuan desert, Mexico: Phylogentic relationships and biogeography". ''Journal of Molluscan Studies'' '''68''': 7-13. [http://si-pddr.si.edu/dspace/handle/10088/7300 PDF].</ref><ref name="Hershler 1999">Hershler R. (1999). "A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part II. Genera ''Colligyrus'', ''Fluminicola'', ''Pristinicola'', and ''Tryonia''". ''The Veliger'' '''42'''(4): 306-337. [http://si-pddr.si.edu/dspace/handle/10088/7293 PDF] {{Webarchive|url=https://web.archive.org/web/20120327053907/http://si-pddr.si.edu/dspace/handle/10088/7293 |date=2012-03-27 }}.</ref> *† ''Feliconcha'' Wesselingh, Anderson & Kadolsky, 2006 - two species from Miocene of the Pebas Formation<ref name="Wesselingh 2006">Wesselingh F. P., Anderson L. C. & Kadolsky D. (2006). "Molluscs from the Miocene Pebas Formation of Peruvian and Colombian Amazonia". ''Scripta Geologica'' '''1333''': 19-290. [https://www.repository.naturalis.nl/document/41346 PDF].</ref> **''Feliconcha feliconcha'' Wesselingh, Anderson & Kadolsky, 2006<ref name="Wesselingh 2006"/> **''Feliconcha reticulata'' Wesselingh, Anderson & Kadolsky, 2006<ref name="Wesselingh 2006"/> *(probably extant)<ref name="Wesselingh 2006"/> ''Glabertryonia'' Wesselingh, Anderson & Kadolsky, 2006 - three species<ref name="Wesselingh 2006"/> **† ''Glabertryonia glabra'' Wesselingh, Anderson & Kadolsky, 2006 - from Miocene of the Pebas Formation<ref name="Wesselingh 2006"/> **† ''Glabertryonia'' sp. 1 - from Pliocene of the Las Piedras Formation<ref name="Wesselingh 2006"/> **(probably extant) ''Glabertryonia'' sp. 2 - from Holocene of Surinam, probably extant<ref name="Wesselingh 2006"/> *''Juturnia'' Hershler, Liu & Stockwell, 2002 *''Lithococcus'' Pilsbry, 1911 *''Mesobia'' F. G. Thompson & Hershler, 1991 *''Mexipyrgus'' Taylor, 1966 *''Minckleyella'' Hershler, Liu & Landye, 2011<ref name="Hershler 2011"/> - with the only species ''Minckleyella balnearis'' Hershler, Liu & Landye, 2011<ref name="Hershler 2011"/> * ''Onobops'' Thompson, 1968<ref name="Wesselingh 2006"/> * ''Pyrgophorus'' Ancey, 1888<ref name="Wesselingh 2006"/> * ''Sioliella'' Haas, 1949 * ''Spurwinkia'' Davis, Mazurkiewicz & Mandracchia, 1982 * ''Texadina'' Abbott & Ladd, 1951 * ''Thalassobia'' Bourguignat in Mabille, 1877

== Cladogram == A cladogram based on sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of species within Cochliopidae:<ref name="Liu 2001">Liu H.-P., Hershler R. & Thompson F. G. (2001). "Phylogenetic Relationships of the Cochliopinae (Rissooidea: Hydrobiidae): An Enigmatic Group of Aquatic Gastropods". ''Molecular Phylogenetics and Evolution'' '''21'''(1): 17-25. {{doi|10.1006/mpev.2001.0988}}.</ref>

{{clade |label1='''Cochliopidae''' |1={{clade |1={{clade |1={{clade |1=''Onobops jacksoni'' |2=''Heleobops docimus'', ''Heleobops dalmatica'' }} |2={{clade |1={{clade |1={{clade |1=''Cochliopina riograndensis'' |2={{clade |1=''Cochliopa'' sp. |2=''Lithococcus multicarinatus'' }} }} |2={{clade |1=''Mexithauma quadripaludium'' |2=''Aroapyrgus'' sp. }} }} |2={{clade |1={{clade |1={{clade |1=''Eremopyrgus eganensis'' |2=''Zetekina'' sp. 1, ''Zetekina'' sp. 2 }} |2={{clade |1={{clade |1="''Tryonia''" ''brevissima'', "''Tryonia''" ''alamosae'' |2=''Aphaostracon'' sp. }} |2={{clade |1={{clade |1={{clade |1=''Littoridinops monroensis'', ''Littoridinops palustris'' |2=''Pyrgophorus platyrachis'' }} |2={{clade |1="''Tryonia''" ''kosteri'' |2=''Durangonella coahuilae'' }} }} |2=''Spurwikinia salsa'' }} }} }} |2={{clade |1=''Tryonia clathrata'', ''Tryonia rowlandsi'', ''Tryonia aequicostata'' |2=''Mexipyrgus carranzae'' }} }} }} }} |2="''Tryonia''" ''robusta'' }} }}

==References== {{Reflist}}

== Further reading == * Hershler R., Davis C. L., Kitting C. L & Liu H.-P. (2007). "Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California". ''Journal of Molluscan Studies'' '''73'''(4): 323-332. [http://si-pddr.si.edu/dspace/handle/10088/6210 PDF]. * Hershler R. & Thompson F. G. (1992). "A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae)". ''Malacological Review'', Supplement '''5''': 1-140. pages 16–18. * [https://dx.doi.org/10.1016/j.ympev.2012.10.025 Wilke T., Haase M., Hershler R., Liu H.-P., Misof B. & Ponder W. (2013) ''Pushing short DNA fragments to the limit: Phylogenetic relationships of ‘hydrobioid’ gastropods (Caenogastropoda: Rissooidea)''. Molecular Phylogenetics and Evolution, 66: 715-736]

== External links == {{Commons category|Cochliopidae}} {{Wikispecies|Cochliopidae}} * [http://www.marinespecies.org/aphia.php?p=taxdetails&id=411706 Cochliopidae on WoRMS]

Category:Cochliopidae Category:Taxa described in 1866 Category:Taxa named by George Washington Tryon