Cassidae

{{Short description|Family of gastropods}} {{Automatic taxobox | image = Semicassis pyrum (top).JPG | image_caption = Lateral view of a shell of ''Semicassis pyrum'' | taxon = Cassidae | authority = Latreille, 1825 | synonyms_ref = <ref name="WoRMS" /> | synonyms = * Cassididae Gyllenhal, 1813 * Oocorythidae P. Fischer, 1885 | type_genus = | subdivision_ranks = Genera | subdivision = }}

The '''Cassidae''' are a taxonomic family of medium-sized, large, and sometimes very large sea snails commonly called '''helmet snails''' or '''bonnet snails'''. These are marine gastropod mollusks in the superfamily Tonnoidea and the clade Littorinimorpha.<ref name="WoRMS">Gofas, S. (2010). Cassidae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=22999 on 2011-01-19</ref>

About 60 species comprise the family Cassidae; an example is ''Cypraecassis rufa''.

==Nomenclature== Despite its incorrect formation (the correct one would be Cassididae, based on the genitive form of Cassis), the ICZN has placed the name Cassidae Latreille, 1825 on the official list of family names, therefore avoiding homonymy with Cassididae Stephens, 1831 (based on Cassida Linnaeus, 1758, a chrysomelid beetle); Opinion 1023 (1974, Bulletin of Zoological Nomenclature 31: 127–129).<ref name="WoRMS" />

==Distribution== Species of this family occur in tropical and temperate seas from the intertidal zone to depths of 100 m (330 ft), buried in the sand during the day and becoming active at night.

==Shell description== Members of this family are shaped rather like bonnets or helmets, as their common name suggests. The shells are large, thick, subglobular with dextrally coiled, sometimes varicose, whorls, and a short spire. The coiling may be trochospiral or convoluted. The shells of many species have great variability, which has led to many misidentifications, resulting in many synonyms.

Many species have a large and solid shield over the parietal body or beside the thick, plicated columella. Many species show blunt knobs and thickened axial ridges, known as varices. The thin, horny operculum is oval in shape and covers a long aperture. The siphonal canal is straight or slightly curved. The outer lip is somewhat thicker at its margin and toothed on the inside.

== Anatomy == These snails have a large mantle and a large, muscular foot. Their large head has an extensible snout. The eyes are at the base of the single pair of tentacles.

==Feeding habits== Helmet bonnets prey on echinoderms (especially sea urchins), starting by gripping them using the foot. The snail then makes a hole in the urchin through the combined action of a secretion which is rich in sulfuric acid and by rasping with their radula. The acid secretion is provided by two large proboscis glands.<ref>[http://md1.csa.com/partners/viewrecord.php?requester=gs&collection=ENV&recid=263350&q=Cassidae&uid=790834666&setcookie=yes Hughes, RN | Hughes, HPI, ''Morphological and Behavioural Aspects of Feeding in the Cassidae (Tonnacea, Mesogastropoda)'' ; Malacologia. Vol. 20, no. 2, pp. 385-402. 1981. ] {{webarchive |url=https://web.archive.org/web/20120829162506/http://md1.csa.com/partners/viewrecord.php?requester=gs&collection=ENV&recid=263350&q=Cassidae&uid=790834666&setcookie=yes |date=August 29, 2012 }}</ref>

==Taxonomy== In their taxonomy of the Gastropoda of 2005 Bouchet & Rocroi still listed Cassidae as Cassinae Latreille, 1825, a subfamily of Tonnidae Suter, 1913 (1825), following in this Riedel (1995).<ref>Riedel F. 1995. An outline of Cassoidean phylogeny (Mollusca, Gastropoda). Contributions to Tertiary and Quaternary Geology 32(4): 97–132</ref> In his "Revision of the deep-water Cassidae" however, Beu raised in 2008 Cassinae to the rank of family.<ref>Beu A.G. (2008) Recent deep-water Cassidae of the world. A revision of Galeodea, Oocorys, Sconsia, Echinophoria and related taxa, with new genera and species (Mollusca, Gastropoda). In Héros V., Cowie R.H. & Bouchet P. (eds), Tropical Deep-Sea Benthos 25. Mémoires du Muséum National d'Histoire Naturelle 196: 269-387.</ref>

==Genera == Genera within the family include: *''Casmaria'' H. Adams and A. Adams, 1853 *''Cassis'' Scopoli, 1777 *''Cypraecassis'' Stutchbury, 1837 *''Dalium'' Dall, 1889 *''Echinophoria'' Sacco, 1890 *''Eucorys'' Beu, 2008 *''Galeodea'' Link, 1807 *''Microsconsia'' Beu, 2008 *''Oocorys'' P. Fischer, 1883 *''Phalium'' Link, 1807 - bonnet shells *''Sconsia'' Gray, 1847 *''Semicassis'' Mörch, 1852

;Genera brought into synonymy : * ''Bathygalea'' Woodring & Olsson, 1957: synonym of ''Echinophoria'' Sacco, 1890 * ''Benthodolium'' Verrill & Smith [in Verrill], 1884: synonym of ''Oocorys'' P. Fischer, 1884 * ''Bezoardica'' Schumacher, 1817: synonym of ''Phalium'' Link, 1807 * ''Bezoardicella'' Habe, 1961: synonym of ''Phalium'' Link, 1807 * ''Cassidaria'' Lamarck, 1816: synonym of ''Galeodea'' Link, 1807 * ''Cassidea'' Bruguière, 1789: synonym of ''Cassis'' Scopoli, 1777 * ''Echinora'' Schumacher, 1817: synonym of ''Galeodea'' Link, 1807 * ''Euspinacassis'' Finlay, 1926: synonym of ''Echinophoria'' Sacco, 1890 * ''Galeoocorys'' Kuroda & Habe, 1957: synonym of ''Galeodea'' Link, 1807 * ''Hadroocorys'' Quinn, 1980: synonym of ''Oocorys'' P. Fischer, 1884 * ''Morio'' Montfort, 1810: synonym of ''Galeodea'' Link, 1807 * ''Taieria'' Finlay & Marwick, 1937: synonym of ''Galeodea'' Link, 1807 * † ''Trachydolium'' Howe, 1926: synonym of ''Echinophoria'' Sacco, 1890 * ''Xenogalea'' Iredale, 1927: synonym of ''Semicassis'' Mörch, 1852 * ''Xenophalium'' Iredale, 1927: synonym of ''Semicassis'' Mörch, 1852

{{clear}} <gallery style="text-align:center;" mode="packed"> Image:Casmaria erinaceus forma vibex 01.jpg|''Casmaria erinaceus'' Image:Cassis cornuta 01.jpg|''Cassis cornuta'' Image:Cypraecassis rufa 01.JPG|''Cypraecassis rufa'' Image:Galeodea rugosa 002.jpg|''Galeodea rugosa'' Image:Phalium bandatum bandatum 001.jpg|''Phalium bandatum'' Image:Semicassis granulata granulata 01.JPG|''Semicassis granulata'' </gallery>

== Human uses == Some helmet shells are carved into cameos, starting in Italy in 1820. *The king helmet ''Cassis tuberosa'' was the first species to be made into cameos. * The black helmet ''Cassis madagascariensis'', also known as ''Cassis cameo'', has a dark brown or a claret-coloured shell layer under a yellowish outer layer. This gives it an onyx appearance and makes it one of the most useful shells for cameos. * The red helmet ''Cypraecassis rufa'' gives a sardonyx-like appearance because it has sard-coloured bands under its pale outer coating. * The horned helmet ''Cassis cornuta'' produces a white figure on an orange background. It is also used as a trumpet by native Filipinos.

==References== {{Reflist}} * Abbott, R.T. (1968). The Helmet Shells of the World (Cassidae) Part 1. ''Indo-Pacific Mollusca'' 2(9): 7-201 * Riedel F. 1995. An outline of Cassoidean phylogeny (Mollusca, Gastropoda). ''Contributions to Tertiary and Quaternary Geology'' 32(4): 97–132 * Kreipl K. 1997. ''Recent Cassidae''. Christa Hemmen, Wiesbaden, 151 p * Beu A.G. (2008) Recent Deep-water Cassidae of the World. A revision of Galeodea, Oocorys, Sconsia, Echinophoria and related taxa, with new genera and species (Mollusca, Gastropoda). In Héros V., Cowie R.H. & Bouchet P. (eds), ''Tropical Deep-Sea Benthos'' 25. ''Mémoires du Muséum National d'Histoire Naturelle'' 196: 269–387.

==External links== * https://www.biolib.cz/en/taxon/id24557/ accessed 5 December 2017 * [https://web.archive.org/web/20070221053405/http://www.mollusca.co.nz/checklist/checklist_frameset.htm Checklist of Mollusca] * [https://web.archive.org/web/20090312100559/http://www.itis.gov/servlet/SingleRpt/SingleRpt ITIS] * Steyn, D.G. & Lussi, M. 2005. ''Offshore Shells of Southern Africa'' {{ISBN|0-620-33607-2}} * http://www.gastropods.com/Taxon_pages/Family_CASSIDAE.shtml#CASSIDAECASSINAE

Category:Cassidae Category:Gastropod families Category:Taxa named by Pierre André Latreille