{{Short description|Subfamily of moths}} {{Automatic taxobox | image = Psaphida P1530619a.jpg | image_caption = ''Psaphida'' | taxon = Amphipyrinae | authority = [[Achille Guenée|Guenée]], 1837 | subdivision_ranks = Tribes | subdivision = * [[Amphipyra|Amphipyrini]] <small>Guenée, 1837</small> * [[Psaphidini]] <small>Grote, 1896</small> }} [[File:Amphipyra pyramidoides PG958579a.jpg| thumb| ''Amphipyra pyramidoides'', Copper underwing, Oklahoma]] [[File:Grote's Sallow (Copivaleria grotei) - Guelph, Ontario.jpg|thumb|''Copivaleria grotei'', Grote's Sallow, Ontario, Canada]] [[File:Emarginea percara PG957404a.jpg|thumb|''Emarginea percara'', Beloved emarginea, Oklahoma]] '''Amphipyrinae''' is a subfamily of [[owlet moth]]s in the family [[Noctuidae]]. There are more than 50 genera and 210 described species in Amphipyrinae, although the classifications are likely to change over time.
This subfamily has been used as a catchall for members of Noctuidae that don't fit well into other subfamilies. As such, many of its members lack morphological traits that would allow assignment into one of the other subfamilies. Genetic analysis conducted on Amphipyrinae is improving the classification of these genera.<ref name=Keegan2019/>
In 2021, phylogenetic research resulted in 11 genera being transferred from Amphipyrinae to six different subfamilies, and it is expected that more members of Amphipyrinae will be reassigned as further research is done.<ref name=Keegan2021/>
Psaphidini is included here as a tribe, but is sometimes treated as the subfamily Psaphidinae. The Australian genera in Acronictinae are sometimes considered part of Amphipyrinae.<ref name=Keegan2019/><ref name=Keegan2021/>
==Genera== These 52 genera belong to the subfamily Amphipyrinae:<ref name=itis/><ref name=gbif/><ref name=bugguide/><ref name=Keegan2021/> ; Tribe [[Amphipyrini]] <small>Guenée, 1837</small> :: ''[[Amphipyra]]'' <small>Ochsenheimer, 1816</small> ; Tribe [[Psaphidini]] <small>Grote, 1896</small> :: Subtribe [[Feraliina]] <small>Poole, 1995</small> ::: ''[[Apsaphida]]'' <small>Franclemont, 1973</small> ::: ''[[Feralia (moth)|Feralia]]'' <small>Grote, 1874</small> ::: ''[[Miracavira]]'' <small>Franclemont, 1937</small> ::: ''[[Paratrachea]]'' <small>Hampson, 1908</small> ::: ''[[Viridemas]]'' <small>Smith, 1908</small> :: Subtribe [[Nocloina]] <small>Poole, 1995</small> ::: ''[[Emarginea]]'' <small>Guenée, 1852</small> ::: ''[[Euamiana]]'' <small>Barnes & Benjamin, 1927</small> ::: ''[[Lythrodes]]'' <small>Smith, 1903</small> ::: ''[[Nocloa]]'' <small>Smith, 1906</small> ::: ''[[Oslaria]]'' <small>Dyar, 1904</small> ::: ''[[Paramiana]]'' <small>Barnes & Benjamin, 1924</small> ::: ''[[Petalumaria]]'' <small>Buckett & Bauer, 1968</small> ::: ''[[Prothrinax]]'' <small>Hampson, 1908</small> ::: ''[[Redingtonia]]'' <small>Barnes & McDunnough, 1912</small> ::: ''[[Ruacodes]]'' <small>Hampson, 1908</small> ::: ''[[Walterella]]'' <small>Dyar, 1921</small> :: Subtribe [[Psaphidina]] <small>Grote, 1896</small> ::: ''[[Brachionycha]]'' <small>Hübner, 1819</small> ::: ''[[Copivaleria]]'' <small>Grote, 1883</small> ::: ''[[Pleromella]]'' <small>Dyar, 1921</small> ::: ''[[Provia (moth)|Provia]]'' <small>Barnes & McDunnough, 1910</small> ::: ''[[Psaphida]]'' <small>Walker, 1865</small> ::: ''[[Pseudocopivaleria]]'' <small>Buckett & Bauer, 1966</small> :: Subtribe [[Triocnemidina]] <small>Poole, 1995</small> ::: ''[[Crimona]]'' <small>Smith, 1902</small> ::: ''[[Oxycnemis]]'' <small>Grote, 1882</small> ::: ''[[Policocnemis]]'' <small>Benjamin, 1932</small> ::: ''[[Triocnemis]]'' <small>Grote, 1881</small> ::: ''[[Unciella]]'' <small>Troubridge, 2008</small> :: Not assigned to a subtribe :: ''[[Allomeganephria]]'' <small>Ronkay, Ronkay, Gyulai & Hacker, 2010</small> :: ''[[Asteroscopus]]'' <small>Boisduval, 1828</small> :: ''[[Benedekia]]'' <small>Ronkay, Ronkay, Gyulai & Hacker, 2010</small> :: ''[[Beshkovietta]]'' <small>Ronkay, Ronkay, Gyulai & Hacker, 2010</small> :: ''[[Copitype]]'' <small>Hampson, 1906</small> :: ''[[Diphtherocome]]'' <small>Warren, 1907</small> :: ''[[Fansipania]]'' <small>Ronkay, Ronkay, Gyulai & Hacker, 2010</small> :: ''[[Flexivaleria]]'' :: ''[[Gracilisinensis]]'' <small>Gyulai, Saldaitis & Vaitonis, 2017</small> :: ''[[Himalaea]]'' <small>Hreblay & Ronkay, 1998</small> :: ''[[Sirioba]]'' <small>Nye, 1975</small> :: ''[[Speidelia]]'' <small>Ronkay, 2000</small> :: ''[[Valeria (moth)|Valeria]]'' <small>Stephens, 1829</small> :: ''[[Vigentinocome]]'' ; Not assigned to a tribe : ''[[Bityla]]'' <small>Walker, 1865</small> : ''[[Centrogone]]'' <small>Hampson, 1908</small> : ''[[Eutamsia]]'' <small>Fletcher D.S., 1961</small> : ''[[Exathetis]]'' <small>Janse, 1938</small> : ''[[Heptapotamia]]'' <small>Alphéraky, 1882</small> : ''[[Leucosigma]]'' <small>Druce, 1908</small> : ''[[Mazuca]]'' <small>Walker, 1866</small> : ''[[Nacna]]'' <small>Fletcher, 1961</small> : ''[[Phidrimana]]'' <small>Kononenko, 1989</small> : ''[[Thoracolopha]]'' <small>Turner, 1939</small>
==References== {{Reflist|refs= <ref name=itis> {{Cite web| title=Amphipyrinae Report | url=https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=693991 | website=Integrated Taxonomic Information System | access-date=2021-11-19 }}</ref>
<ref name=gbif> {{Cite web| title=Noctuidae | url=https://www.gbif.org/species/7015 | website=GBIF, Global Biodiversity Information Facility | access-date=2021-11-19 }}</ref>
<ref name=bugguide> {{Cite web| title=Amphipyrinae subfamily Information | url=https://bugguide.net/node/view/12343 | website=BugGuide.net | access-date=2021-11-19 }}</ref>
<ref name=Keegan2019> {{Cite journal | title = Towards resolving and redefining Amphipyrinae (Lepidoptera, Noctuoidea, Noctuidae): a massively polyphyletic taxon | date = 2019 | last1 = Keegan | first1 = Kevin L. | last2 = Lafontaine | first2 = James Donald | last3 = Wahlberg | first3 = Niklas | last4 = Wagner | first4 = David L. | journal = Systematic Entomology | volume = 44| issue = 2 | pages = 451–464 | url = https://www.biorxiv.org/content/biorxiv/early/2018/10/12/271478.full.pdf | doi = 10.1111/syen.12336 | bibcode = 2019SysEn..44..451K | s2cid = 92559412 }}</ref>
<ref name=Keegan2021> {{Cite journal | title = Toward a Stable Global Noctuidae (Lepidoptera) Taxonomy | date = 2021 | last1 = Keegan | first1 = Kevin | last2 = Rota | first2 = Jadranka | last3 = Zahiri | first3 = Reza | last4 = Zilli | first4 = Alberto | last5 = Wahlberg | first5 = Niklas | last6 = Schmidt | first6 = B. | last7 = Lafontaine | first7 = J. | last8 = Goldstein | first8 = Paul | last9 = Wagner | first9 = David | display-authors = 4 | journal = Insect Systematics and Diversity | volume = 5 | issue = 3 | url = https://www.researchgate.net/publication/351287266 | doi = 10.1093/isd/ixab005 | doi-access = free }}</ref> }}
==Further reading== {{refbegin}} * {{Cite book | title = List of the Lepidoptera of North America | date = 1868 | last1 = Grote | first1 = Aug. R. | last2 = Robinson | first2 = C. T. | publisher = American Entomological Society | isbn = 978-0-665-04389-5 | url = https://archive.org/details/cihm_04389 }} * {{Cite journal | title = Annotated check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico | date = 2010 | last1 = Lafontaine | first1 = J. Donald | last2 = Schmidt | first2 = B. Christian | journal = ZooKeys | issue = 40| pages = 127–47 | doi = 10.3897/zookeys.40.414 | doi-access = free | bibcode = 2010ZooK...40....1L }} * {{Cite journal | title = Additions and corrections to the checklist of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico, III | date = 2015 | last1 = Lafontaine | first1 = J. Donald | last2 = Schmidt | first2 = B. Christian | journal = ZooKeys | issue = 527| pages = 227–236 | doi = 10.3897/zookeys.527.6151 | pmc=4668890 | pmid=26692790 | doi-access = free | bibcode = 2015ZooK..527..127L }} * {{Cite report | title = Annotated taxonomic checklist of the Lepidoptera of North America, North of Mexico | date = 2016 | last1 = Pohl | first1 = Greg | last2 = Patterson | first2 = Bob | last3 = Pelham | first3 = Jonathan | url = https://www.researchgate.net/publication/302570819 | doi = 10.13140/RG.2.1.2186.3287| doi-access = free }} {{refend}}
==External links== {{refbegin}} * {{Commons-inline}} {{refend}}
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[[Category:Noctuidae]] [[Category:Taxa named by Achille Guenée]]
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