# Typophorus

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***Typophorus*** is a genus of [leaf beetles](/source/Leaf_beetle) in the subfamily [Eumolpinae](/source/Eumolpinae). There are at least 52 described species in *Typophorus*.[1][2][3][4][5][6] Similar genera include *[Paria](/source/Paria_(beetle))* and *[Tijucana](/source/Tijucana)*. *Typophorus* itself is probably [polyphyletic](/source/Polyphyletic).[7]

Two subspecies of *[Typophorus nigritus](/source/Typophorus_nigritus)*, *T. nigritus nitidulus* and *T. nigritus viridicyaneus*, are known as pests of [sweet potatoes](/source/Sweet_potato).[8]

## Species

- *[Typophorus annulatus](/source/Typophorus_annulatus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *[Typophorus apicicornis](/source/Typophorus_apicicornis)* [Jacoby](/source/Martin_Jacoby), 1891[9]
- *[Typophorus appendiculatus](/source/Typophorus_appendiculatus)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1948
- *[Typophorus atomarius](/source/Typophorus_atomarius)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[10]
- *[Typophorus basalis](/source/Typophorus_basalis)* [Baly](/source/Joseph_Sugar_Baly), 1859[11]
- *[Typophorus biplagiatus](/source/Typophorus_biplagiatus)* [Boheman](/source/Carl_Henrik_Boheman), 1858
- *[Typophorus bisignatus](/source/Typophorus_bisignatus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *[Typophorus bohumilae](/source/Typophorus_bohumilae)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[12]
- *[Typophorus centromaculatus](/source/Typophorus_centromaculatus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1888
- *[Typophorus cyanipennis](/source/Typophorus_cyanipennis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *[Typophorus diomedes](/source/Typophorus_diomedes)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1955[13]
- *[Typophorus erythocephalus](/source/Typophorus_erythocephalus)* [Jacoby](/source/Martin_Jacoby), 1882[14]
- *[Typophorus exaequatus](/source/Typophorus_exaequatus)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[12]
- *[Typophorus exilis](/source/Typophorus_exilis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1878[15]
- *[Typophorus fasciatus](/source/Typophorus_fasciatus)* ([Germar](/source/Ernst_Friedrich_Germar), 1824)
- *[Typophorus florigradus](/source/Typophorus_florigradus)* [Bechyné](/source/Jan_Bechyn%C3%A9) & Bechyné, 1961[16]
- *[Typophorus furvus](/source/Typophorus_furvus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1885[17]
- *[Typophorus inflatus](/source/Typophorus_inflatus)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1958[18]
- *[Typophorus intermedius](/source/Typophorus_intermedius)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *[Typophorus interstitalis](/source/Typophorus_interstitalis)* [Jacoby](/source/Martin_Jacoby), 1891[9]
- *[Typophorus jacobyi](/source/Typophorus_jacobyi)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[19][20] - *Typophorus jacobyi atramentaria* ([Weise](/source/Julius_Weise), 1921)[21][22] - *Typophorus jacobyi jacobyi* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884
- *[Typophorus kuscheli](/source/Typophorus_kuscheli)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[12]
- *[Typophorus lar](/source/Typophorus_lar)* ([Fabricius](/source/Johan_Christian_Fabricius), 1787)
- *[Typophorus limbatus](/source/Typophorus_limbatus)* [Jacoby](/source/Martin_Jacoby), 1891[9]
- *[Typophorus maculigera](/source/Typophorus_maculigera)* ([Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1888)[20]
- *[Typophorus mansuetus](/source/Typophorus_mansuetus)* [Bechyné](/source/Jan_Bechyn%C3%A9) & Bechyné, 1961[16]
- *[Typophorus melanocephalus](/source/Typophorus_melanocephalus)* [Jacoby](/source/Martin_Jacoby), 1876[23]
- *[Typophorus mexicanus](/source/Typophorus_mexicanus)* [Jacoby](/source/Martin_Jacoby), 1876[23]
- *[Typophorus minutus](/source/Typophorus_minutus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *[Typophorus nanus](/source/Typophorus_nanus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *[Typophorus nigra](/source/Typophorus_nigra)* ([Weise](/source/Julius_Weise), 1921)[21][20]
- *[Typophorus nigricollis](/source/Typophorus_nigricollis)* [Jacoby](/source/Martin_Jacoby), 1882[14]
- *[Typophorus nigritarsis](/source/Typophorus_nigritarsis)* ([Jacoby](/source/Martin_Jacoby), 1882)[20]
- *[Typophorus nigritus](/source/Typophorus_nigritus)* ([Fabricius](/source/Johan_Christian_Fabricius), 1801) - *Typophorus nigritus chalceus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5] - *Typophorus nigritus coronadoi* [Bechyné](/source/Jan_Bechyn%C3%A9), 1948 - *Typophorus nigritus cretifer* [Bechyné](/source/Jan_Bechyn%C3%A9), 1953[6] - *Typophorus nigritus erbeni* [Bechyné](/source/Jan_Bechyn%C3%A9), 1948 - *Typophorus nigritus interpositus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951 - *Typophorus nigritus lucens* [Bechyné](/source/Jan_Bechyn%C3%A9), 1953[6] - *Typophorus nigritus molnari* [Bechyné](/source/Jan_Bechyn%C3%A9), 1953[6] - *Typophorus nigritus nigritus* ([Fabricius](/source/Johan_Christian_Fabricius), 1801) - *Typophorus nigritus nitidulus* ([Fabricius](/source/Johan_Christian_Fabricius), 1801) - *Typophorus nigritus obliquus* [Baly](/source/Joseph_Sugar_Baly), 1859[11] - *Typophorus nigritus paradoxus* [Jacoby](/source/Martin_Jacoby), 1882[14] - *Typophorus nigritus punctatissimus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1948 - *Typophorus nigritus viridicyaneus* ([Crotch](/source/George_Robert_Crotch), 1873)[24] (sweetpotato leaf beetle)
- *[Typophorus picimanus](/source/Typophorus_picimanus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *[Typophorus pumilus](/source/Typophorus_pumilus)* ([LeConte](/source/John_Lawrence_LeConte), 1859)[25][26]
- *[Typophorus purulenis](/source/Typophorus_purulenis)* [Jacoby](/source/Martin_Jacoby), 1882[14]
- *[Typophorus pygmaeus](/source/Typophorus_pygmaeus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[10]
- *[Typophorus quinquemaculatus](/source/Typophorus_quinquemaculatus)* [Erichson](/source/Wilhelm_Ferdinand_Erichson), 1847
- *[Typophorus rufipes](/source/Typophorus_rufipes)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *[Typophorus santaremus](/source/Typophorus_santaremus)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1955[13]
- *[Typophorus scheerpeltzi](/source/Typophorus_scheerpeltzi)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[12]
- *[Typophorus sericeus](/source/Typophorus_sericeus)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1948
- *[Typophorus signatus](/source/Typophorus_signatus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1891[20]
- *[Typophorus simplex](/source/Typophorus_simplex)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[27]
- *[Typophorus spadiceus](/source/Typophorus_spadiceus)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *[Typophorus steinheili](/source/Typophorus_steinheili)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1878[15]
- *[Typophorus subbrunneus](/source/Typophorus_subbrunneus)* [Jacoby](/source/Martin_Jacoby), 1882[14]
- *[Typophorus sulcifer](/source/Typophorus_sulcifer)* [Bechyné](/source/Jan_Bechyn%C3%A9), 1954[28]
- *[Typophorus tarsalis](/source/Typophorus_tarsalis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1888
- *[Typophorus tibialis](/source/Typophorus_tibialis)* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *[Typophorus variabilis](/source/Typophorus_variabilis)* [Jacoby](/source/Martin_Jacoby), 1882[14]

*[incertae sedis](/source/Incertae_sedis)* species:

- *Typophorus variabilis* ([Fabricius](/source/Johan_Christian_Fabricius), 1801)

Species moved to *[Paria](/source/Paria_(beetle))*:[6][20]

- *Typophorus aeneipennis* [Baly](/source/Joseph_Sugar_Baly), 1878[29]
- *Typophorus atripennis* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1878[18]
- *Typophorus corumbanus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[12]
- *Typophorus epimeralis* [Bechyné](/source/Jan_Bechyn%C3%A9), 1955[13]
- *Typophorus erythopus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1888
- *Typophorus festinatus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1950[30]
- *Typophorus fulvipennis* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[10]
- *Typophorus geniculatus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1878
- *Typophorus horvathi* [Bechyné](/source/Jan_Bechyn%C3%A9), 1955[13]
- *Typophorus montanellus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1951[31]
- *Typophorus nigripennis* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1884[10]
- *Typophorus nigronotatus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]
- *Typophorus pusillus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1876
- *Typophorus quadriplagiatus* [Jacoby](/source/Martin_Jacoby), 1876[23]
- *Typophorus tinctus* [Bechyné](/source/Jan_Bechyn%C3%A9), 1957[22]

Species moved to *[Entreriosa](/source/Entreriosa)*:[6]

- *Typophorus umbratus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877[5]

Synonyms:[6]

- *Typophorus histrio* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877:[5] synonym of *[Paria opacicollis](/source/Paria_opacicollis)* [LeConte](/source/John_Lawrence_LeConte), 1859
- *Typophorus nobilis* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877:[5] synonym of *[Typophorus basalis](/source/Typophorus_basalis)* [Baly](/source/Joseph_Sugar_Baly), 1859
- *Typophorus sturmi* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877:[5] synonym of *Typophorus nigritus chalceus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877
- *Typophorus versutus* [Lefèvre](/source/%C3%89douard_Lef%C3%A8vre), 1877:[5] synonym of *Typophorus nigritus nitidulus* ([Fabricius](/source/Johan_Christian_Fabricius), 1801)

## References

1. ["*Typophorus* Report"](https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=719744). *Integrated Taxonomic Information System*. Retrieved 2018-05-04.

1. ["Browse *Typophorus*"](http://www.catalogueoflife.org/col/browse/tree/id/1f666207bd4c74c294e5024f3d4982a1). *Catalogue of Life*. Retrieved 2018-05-04.

1. ["*Typophorus*"](https://www.gbif.org/species/1049018). *GBIF*. Retrieved 2018-05-04.

1. ["*Typophorus* Genus Information"](https://bugguide.net/node/view/109035). *BugGuide.net*. Retrieved 2018-05-04.

1. Lefèvre, E. (1877). ["Descriptions de Coléoptères nouveaux ou peu connus de la Famille des Eumolpides (2e partie)"](https://biodiversitylibrary.org/page/8244637). *[Annales de la Société Entomologique de France](/source/Annales_de_la_Soci%C3%A9t%C3%A9_Entomologique_de_France)*. **7**: 309–326. 5.

1. Bechyné, J. (1953). ["Katalog der neotropischen Eumolpiden (Col. Phytoph. Chrysomeloidea)"](https://www.biodiversitylibrary.org/page/49645311) (in German). *Entomologische Arbeiten aus dem Museum G. Frey*. **4**: 26–303.

1. Flowers, R.W. (1996). ["The subfamily Eumolpinae (Coleoptera: Chrysomelidae) in Central America"](http://www.sidalc.net/cgi-bin/wxis.exe/?IsisScript=oet.xis&method=post&formato=2&cantidad=1&expresion=mfn=009919). *Revista de Biología Tropical Special Publication*. **2**: 1–59. Retrieved December 31, 2018.

1. Santoro, F. H.; Bezzi, A.; Vigevano, A.; Cantos, F. (1980). ["Biología del negrito de la batata, *Typophorus nigritus nitidulus* (F.) y ensayo preliminar sobre control químico de adultos (Coleoptera-Chrysomelidae-Eumolpinae)"](http://www.bfa.fcnym.unlp.edu.ar/catalogo/doc_num.php?explnum_id=271). *IDIA (Buenos Aires)*. **373–375**: 13–44.

1. Jacoby, M. (1890). ["Eumolpidae"](https://biodiversitylibrary.org/page/585383). *Insecta. Coleoptera, Supplement to Phytophaga*. *[Biologia Centrali-Americana](/source/Biologia_Centrali-Americana)*. Vol. 6, no. 1. pp. 183–241.

1. Lefèvre, E. (1884). ["Descriptions de quatre genres nouveaux et de plusieurs espèces nouvelles de coléoptères de la famille des eumolpides"](https://biodiversitylibrary.org/page/52116991). *Bulletin de la Société Entomologique de Belgique*. **1884**: cxciii–ccvi.

1. Baly, J. S. (1859). "Descriptions of new genera and species of Phytophagous insects". *[Annals and Magazine of Natural History](/source/Annals_and_Magazine_of_Natural_History)*. **4** (20): 124–128. 3. [doi:10.1080/00222935908697096](https://doi.org/10.1080/00222935908697096)

1. Bechyné, J. (1951). ["Liste provisoire des Eumolpides de Bolivie et observations diverses sur les espèces de l'Amérique du Sud (Col. Phytophaga)"](https://www.biodiversitylibrary.org/page/45935465). *Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München*. **2**: 227–352.

1. Bechyné, J. (1955). ["Reise des Herrn G. Frey in Südamerika: Eumolpidae (Col. Phytophaga)"](https://www.biodiversitylibrary.org/page/45918278). *Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München*. **6**: 569–657.

1. Jacoby, M. (1881). ["Eumolpidae"](https://biodiversitylibrary.org/page/584677). *Insecta. Coleoptera*. *[Biologia Centrali-Americana](/source/Biologia_Centrali-Americana)*. Vol. 6, no. 1. pp. 105–187.

1. Lefèvre, E. (1878). ["Voyage de M. E. Steinheil à la Nouvelle Grenade. Eumolpides"](https://biodiversitylibrary.org/page/12654884). *Mittheilungen des Münchener Entomologischen Vereins*. **2**: 112–133.

1. Bechyné, J. & Springlovà de Bechyné, B. (1961). ["Notas sobre Chrysomeloidea neotropicais"](http://chrysomelidae.miza-ucv.org.ve/sites/chrysomelidae.miza-ucv.org.ve/files/1961_Separata_151.pdf). *Boletim do Museu Paraense Emilio Goeldi*. **33**: 1–49. New Series.

1. Lefèvre, É. (1885). ["Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis"](https://biodiversitylibrary.org/page/27434526). *Mémoires de la Société Royale des Sciences de Liège*. **11** (16): 1–172. 2.

1. Bechyné, J. (1958). ["Notizen zu den neotropischen Chrysomeloidea (Col. Phytophaga)"](https://www.biodiversitylibrary.org/page/45990136). *Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München*. **9**: 478–706.

1. Lefèvre, E. (1884). ["\[Communications\]"](https://biodiversitylibrary.org/page/32550559). *[Bulletin de la Société entomologique de France](/source/Bulletin_de_la_Soci%C3%A9t%C3%A9_entomologique_de_France)*. **1884**: LXXV–LXXVI.

1. Bechyné, J. (1957). ["Eumolpides neo-tropicaux de la collection du Museo civico di Storia naturale "Giacomo Doria" di Genova (Col. Phytophaga)"](https://www.biodiversitylibrary.org/page/34909536). *Annali del Museo Civico di Storia Naturale Giacomo Doria*. **69**: 226–247.

1. Weise, J. (1921). ["Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn Dr. A. Roman in Amazonas 1914—1915. 6. Chrysomelidae"](https://biodiversitylibrary.org/page/6412960). *Arkiv för Zoologi*. **14** (1): 1–205.

1. Bechyné, J. (1957). ["Voyage de M. le Dr. A. Roman au Brésil (1914-1915) Eumolpides, Galerucides et Alticides (Col. Phytophaga)"](http://chrysomelidae.miza-ucv.org.ve/sites/chrysomelidae.miza-ucv.org.ve/files/1957_Separata_132.pdf). *Arkiv för Zoologi*. **11**: 133–152.

1. Jacoby, M. (1876). ["Description of new Genera and Species of Phytophagous Coleoptera"](https://biodiversitylibrary.org/page/29100094). *[Proceedings of the Zoological Society of London](/source/Proceedings_of_the_Zoological_Society_of_London)*. **1876**: 807–817.

1. Crotch, G. R. (1873). "Materials for the Study of the Phytophaga of the United States". *[Proceedings of the Academy of Natural Sciences of Philadelphia](/source/Proceedings_of_the_Academy_of_Natural_Sciences_of_Philadelphia)*. **25** (1): 19–83. [JSTOR 4624297](https://www.jstor.org/stable/4624297)

1. LeConte, J. L. (1859). ["The Coleoptera of Kansas and eastern New Mexico"](https://biodiversitylibrary.org/page/17592421). *[Smithsonian Contributions to Knowledge](/source/Smithsonian_Contributions_to_Knowledge)*. **11**: i–vi, 1–58. [doi:10.5962/bhl.title.18986](https://doi.org/10.5962/bhl.title.18986)

1. Wilcox, J. A. (1957). ["A Revision of the North American Species of *Paria* Lec. (Coleoptera: Chrysomelidae)"](https://biodiversitylibrary.org/page/57341153). *New York State Museum and Science Service Bulletin*. '***(365): 1–45.***

1. Lefèvre, E. (1884). ["Eumolpides du voyage du Dr E. Fromont au Brésil et à la Plata"](https://biodiversitylibrary.org/page/52116953). *Bulletin de la Société Entomologique de Belgique*. **1884**: clv–clvi.

1. Bechyné, J. (1954). ["Über die in Matto Grosso von F. Plaumann gesammelten Chrysomeloidea (Col. Phytophaga)"](http://chrysomelidae.miza-ucv.org.ve/sites/chrysomelidae.miza-ucv.org.ve/files/1954_Separata_91.pdf). *Entomologischen Arbeiten aus dem Museum G. Frey*. **5**: 116–133.

1. Bechyné, J. & Springlovà de Bechyné, B. (1961). ["Notas sobre Chrysomeloidea Neotropicais II"](http://chrysomelidae.miza-ucv.org.ve/sites/chrysomelidae.miza-ucv.org.ve/files/1961_Separata_152.pdf). *Boletim do Museu Paraense Emilio Goeldi*. **37**: 1–93. New Series.

1. Bechyné, J. (1950). ["Notes sur les Eumolpides néotropicaux des collections de l'Institut royal des Sciences naturelles de Belgique"](http://new-biblio.naturalsciences.be/rbins-publications/bulletin-of-the-royal-belgian-institute-of-natural-sciences-bulletin-de-linstitut-royal-des-sciences-naturelles-de-belgique-bulletin-van-het-koninklijk-belgisch-instituut-voor-natuurwetenschappen/26-1950/irscnb_p4087_01013ap_26_bulletin-54.pdf). *Bulletin of the Royal Belgian Institute of Natural Sciences*. **26** (54): 1–24.

1. Bechyné, J. (1951). ["Les Chrysomeloidea neotropicaux des collections du Muséum Zool. de l'Université a Helsingfors"](http://chrysomelidae.miza-ucv.org.ve/sites/chrysomelidae.miza-ucv.org.ve/files/1951_Separata_73.pdf). *Notulae Entomologicae*. **31**: 59–66.

## Further reading

- Riley, Edward G.; Clark, Shawn M.; Seeno, Terry N. (2003). *Catalog of the leaf beetles of America north of Mexico (Coleoptera: Megalopodidae, Orsodacnidae and Chrysomelidae, excluding Bruchinae)*. Special Publication No. 1. The Coleopterists' Society. ISBN 978-0-9726087-1-8.
- Lobl, I. & Smetana, A. (eds.) (2013). *Catalogue of Palaearctic Coleoptera, Volume 6: Chrysomeloidea*. Apollo Books. ISBN 978-90-04-26091-7.

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Adapted from the Wikipedia article [Typophorus](https://en.wikipedia.org/wiki/Typophorus) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Typophorus?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.