# Tryella

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Genus of true bugs

Tryella Tryella graminea Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hemiptera Suborder: Auchenorrhyncha Family: Cicadidae Subfamily: Cicadettinae Tribe: Lamotialnini Genus: Tryella Moulds, 2003 Type species Tryella ochra Synonyms[1] Tyrella Moulds, 2003

*Tryella willsi*, Australia

***Tryella*** is a [genus](/source/Genus) of [cicadas](/source/Cicada) found in [Australia](/source/Australia) and [New Guinea](/source/New_Guinea). The group are commonly called **bullets**, the name derived from markings behind their heads, which together with their compact bodies make them resemble bullets while in flight.[2]

## Taxonomy

For many years, the members were classified in the genus *[Abricta](/source/Abricta)*. However, recent review of the genus has shown it to be a disparate group of species, and the [Australian](/source/Australia) members moved to other genera. Australian entomologist [Maxwell Sydney Moulds](/source/Maxwell_Sydney_Moulds) conducted a morphological analysis of the genus and found the cicadas split naturally into clades according to biogeographical region.[3]

Of the 15 Australian species, the floury baker was the earliest offshoot. Unpublished data confirmed it was quite genetically distant from the other 14 species and so it was classified in a new monotypic genus *[Aleeta](/source/Aleeta)*, while the others were placed in the genus *Tryella*. The genus name is derived from the [Ancient Greek](/source/Ancient_Greek) *tryelis* "[ladle](/source/Ladle_(spoon))" and relates to the ladle-shaped uncal lobes of the species in the genus.[3]

Phylogenetic evidence supports *Tryella* and *Aleeta* as being the closest relatives to the famous [periodical cicadas](/source/Periodical_cicadas) (genus *[Magicicada](/source/Periodical_cicadas)*) of [North America](/source/North_America) despite being widely geographically separated from them.[4]

## Species

The [type species](/source/Type_species) is *[Tryella ochra](/source/Tryella_ochra)*. Within the group, cladistic morphological analysis showed *T. rubra* to be the earliest offshoot within the genus. The relationships between other species was not able to be clarified.[3]

As of 2025 there were 16 described species in the genus.[5][6]

- *[Tryella adela](/source/Tryella_adela)* Moulds, 2003 (NT) - small maroon bullet

- *[Tryella burnsi](/source/Tryella_burnsi)* Moulds, 2003 (QLD) - brown bullet

- *[Tryella castanea](/source/Tryella_castanea)* (Distant, 1905) (NT, QLD, WA) - small rusty bullet

- *[Tryella crassa](/source/Tryella_crassa)* Moulds, 2003 (NT, QLD) - dusky bullet

- *[Tryella fumipennis](/source/Tryella_fumipennis)* Emery, Emery, Hutchinson & Ong, 2022 (WA) - smoky-winged bullet

- *[Tryella graminea](/source/Tryella_graminea)* Moulds, 2003 (NT, QLD) - grass bullet

- *[Tryella infuscata](/source/Tryella_infuscata)* Moulds, 2003 (NT, QLD) - large maroon bullet

- *[Tryella kauma](/source/Tryella_kauma)* Moulds, 2003 (QLD) - whispering bullet

- *[Tryella lachlani](/source/Tryella_lachlani)* Moulds, 2003 (QLD, PNG) - golden black bullet

- *[Tryella noctua](/source/Tryella_noctua)* (Distant, 1913) (WA) - chocolate bullet

- *[Tryella occidens](/source/Tryella_occidens)* Moulds, 2003 (WA) - dusty brown bullet

- *[Tryella ochra](/source/Tryella_ochra)* Moulds, 2003 (NT, QLD) - golden-brown bullet

- *[Tryella rubra](/source/Tryella_rubra)* (Goding & Froggatt, 1904) (WA, NT, QLD) - large rusty bullet

- *[Tryella stalkeri](/source/Tryella_stalkeri)* (Distant, 1907) (WA) - honey bullet

- *[Tryella willsi](/source/Tryella_willsi)* (Distant, 1882) (QLD) - black bullet

- *[Tryella wuggubun](/source/Tryella_wuggubun)* Emery, Emery, Hutchinson & Ong, 2022 (NT, WA) - Kimberley bullet

## Distribution and habitat

The species are found across eastern, central and northern Australia, and south-western New Guinea, predominantly on trees, especially [eucalypts](/source/Eucalypt).[3]

## Behaviour

Male *Tryella* cicadas call during the day and dusk. Adults emerge from pupation after monsoonal rain.[3] They characteristically sit facing downwards on branches,[2] of generally less than 5 cm (2.0 in) in diameter.[3] One species, *T. graminea*, is found on grass.[3]

## References

1. **[^](#cite_ref-itis_1-0)** ["ITIS report, *Tryella*"](https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=1217779). Retrieved 2024-12-03.

1. ^ [***a***](#cite_ref-popple_2-0) [***b***](#cite_ref-popple_2-1) Popple, Lindsay (2006). ["Genus *Tryella* Moulds, 2003 (Bullets)"](http://sci-s03.bacs.uq.edu.au/ins-info/Abr2.htm). *The cicadas of central eastern Australia*. University of Queensland. Retrieved 5 June 2013.

1. ^ [***a***](#cite_ref-moulds_2003_3-0) [***b***](#cite_ref-moulds_2003_3-1) [***c***](#cite_ref-moulds_2003_3-2) [***d***](#cite_ref-moulds_2003_3-3) [***e***](#cite_ref-moulds_2003_3-4) [***f***](#cite_ref-moulds_2003_3-5) [***g***](#cite_ref-moulds_2003_3-6) Moulds, Maxwell Sydney (2003). ["An appraisal of the cicadas of the genus Abricta Stål and allied genera (Hemiptera: Auchenorrhyncha: Cicadidae)"](https://web.archive.org/web/20111204205204/http://australianmuseum.net.au/Uploads/Journals/17962/1386_complete.pdf) (PDF). *Records of the Australian Museum*. **55** (3): 245–304. [doi](/source/Doi_(identifier)):[10.3853/j.0067-1975.55.2003.1386](https://doi.org/10.3853%2Fj.0067-1975.55.2003.1386). Archived from [the original](http://australianmuseum.net.au/Uploads/Journals/17962/1386_complete.pdf) (PDF) on 2011-12-04. Retrieved 2013-06-05.

1. **[^](#cite_ref-4)** Marshall, David C.; Moulds, Max; Hill, Kathy B. R.; Price, Benjamin W.; Wade, Elizabeth J.; Owen, Christopher L.; Goemans, Geert; Marathe, Kiran; Sarkar, Vivek; Cooley, John R.; Sanborn, Allen F. (2018-05-28). ["A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification"](https://www.mapress.com/j/zt/article/view/zootaxa.4424.1.1). *Zootaxa*. **4424** (1): 1–64. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.4424.1.1](https://doi.org/10.11646%2Fzootaxa.4424.1.1). [ISSN](/source/ISSN_(identifier)) [1175-5334](https://search.worldcat.org/issn/1175-5334). [PMID](/source/PMID_(identifier)) [30313477](https://pubmed.ncbi.nlm.nih.gov/30313477).

1. **[^](#cite_ref-Dmitriev2024_5-0)** Dmitriev, D.A. (2024). ["World Auchenorrhyncha Database"](https://hoppers.speciesfile.org). Retrieved 2024-08-20.

1. **[^](#cite_ref-afd_6-0)** ["Australian Faunal Directory, Genus Tryella Moulds, 2003"](https://biodiversity.org.au/afd/taxa/Tryella). Retrieved 2024-12-03.

Taxon identifiers Tryella Wikidata: Q16746868 Wikispecies: Tryella AFD: Tryella BOLD: 342173 CoL: 8HSHR EoL: 3741668 GBIF: 4778707 iNaturalist: 368560 IRMNG: 1378407 NCBI: 1218665 Open Tree of Life: 865657

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