# Tentaculites

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{{Short description|Extinct genus of invertebrates}}
{{Automatic taxobox
| fossil_range = {{fossil range | Early Ordovician | Late Devonian | ref=<ref name="Traverse2007">{{Cite book | last1 = Traverse | first1 = A. | chapter = What Paleopalynology Is and Is Not | title = Paleopalynology | series = Topics in Geobiology | volume = 28 | pages = 1–43 | year = 2007 | isbn = 978-1-4020-6684-9 | doi = 10.1007/978-1-4020-5610-9_1}}</ref> }}
| image = Tentaculites bellulus.jpg
| image_caption = ''[Tentaculites bellulus](/source/Tentaculites_bellulus)'' from the Middle Devonian of Wisconsin
| taxon = Tentaculites
| authority = von Schlotheim, 1820
| subdivision_ranks = Species
| subdivision =
* ''T. bellulus'' <small>Hall, 1876</small>
* ''T. exaltatus'' <small>Talent, 1963</small>
* ''T. grandis'' <small>Roemer, 1870</small>
* ''[T. oswegoensis](/source/Tentaculites_oswegoensis)'' <small>F.B. Meek & A.H. Worthen, 1865</small>
}}

'''''Tentaculites''''' is an extinct genus of conical fossils of uncertain affinity, class [Tentaculita](/source/Tentaculita), although it is not the only member of the class. It is known from  Lower Ordovician to Upper Devonian deposits<ref name="Traverse2007" /> both as calcitic shells with a brachiopod-like microstructure<ref name="Towe1978">{{Cite journal | last1 = Towe | first1 = K. M. | title = Tentaculites: Evidence for a Brachiopod Affinity? | journal = Science | volume = 201 | issue = 4356 | pages = 626–628 | year = 1978 | pmid = 17794124 | doi = 10.1126/science.201.4356.626| bibcode = 1978Sci...201..626T | s2cid = 22806221 }}</ref> and carbonaceous 'linings'.<ref>Wood, G.D., Miller, M.A., and Bergstrom, S.M. 2004. Late Devonian (Frasnian) tentaculite organic remains in palynological preparations, Radom−Lublin region, Poland. Memoirs of the Association of Australian Palaeontologists 29: 253–258.</ref><ref name="Filipiak2009">{{Cite journal | last1 = Filipiak | first1 = P. | last2 = Jarzynka | first2 = A. | title = Organic Remains of Tentaculitids: New Evidence from Upper Devonian of Poland | journal = Acta Palaeontologica Polonica | volume = 54 | pages = 111–116 | year = 2009 | doi = 10.4202/app.2009.0111| doi-access = free }}</ref>

== Affinity==
The taxonomic classification of tentaculitids is uncertain.  Some grouped them with [pteropod](/source/pteropod)s, but there is no modern support and only superficial similarity.{{who|<!--mentioned in following reference but basis for relationship not specified-->|date=December 2015}}<ref>Ager, 1963, Principles of Palaeontology</ref> They may also be related to other conical shells of uncertain affinity including [cornulitids](/source/Cornulitida), ''[Anticalyptraea](/source/Anticalyptraea)'', [microconchids](/source/Microconchida) and [trypanoporids](/source/Trypanoporida).<ref name='Vinn2010'>{{cite journal
 |author=Vinn, O.
 |title=Adaptive strategies in the evolution of encrusting tentaculitoid tubeworms
 |journal= 	Palaeogeography, Palaeoclimatology, Palaeoecology 
 |year=2010
 |volume=292
 |issue=1–2
 |pages=211–221
 |url=https://www.researchgate.net/publication/222285765
 |access-date = 2014-01-11
 |doi=10.1016/j.palaeo.2010.03.046
|bibcode=2010PPP...292..211V
 }}</ref>
Their shell microstructure has warranted their comparison with the [brachiopod](/source/brachiopod)s and [phoronid](/source/phoronid)s,<ref name="Towe1978" /><ref name='VinnZaton2008'>{{cite journal
 | doi =  10.1080/11035897.2012.669788
 | title = Phenetic phylogenetics of tentaculitoids — extinct problematic calcareous tube-forming organisms
 | year = 2012
 | author = Vinn, O.
 | author2 = Zatoń, M. 
 | journal = GFF
 | volume = 134
 | issue = 2
 | pages = 145–156  
 | s2cid = 83591938
 | url = https://www.researchgate.net/publication/234023270
 | access-date = 2014-06-11}}</ref>
and the possible Ediacaran lophophorate ''[Namacalathus](/source/Namacalathus)''.<ref>1. Zhuravlev, A.Y., Wood, R.A., and Penny, A.M. (2015). Ediacaran skeletal metazoan interpreted as a lophophorate. Proc. R. Soc. B 282, 20151860. Available at: http://rspb.royalsocietypublishing.org/lookup/doi/10.1098/rspb.2015.1860.</ref>

== Morphology ==
thumb|right|Sketch of a ''Tentaculites ornatus'', Silurian.
[[File:Tentaculitid041011.jpg|thumb|right|Tentaculitid from the New Creek Limestone ([Lochkovian](/source/Lochkovian), Early Devonian) of New Creek, [West Virginia](/source/West_Virginia).]]
Tentaculitids have ribbed, cone-shaped shells which range in length from 5 to 20&nbsp;mm.  Some species septate; their embryonic shell, which is retained, forms a small, sometimes spherical, chamber.<ref name="Filipiak2009" />

==Ecology==
Some species are inferred to have been planktonic.<ref>{{cite journal|author=LARSSON K. |year=1979 |title=Silurian tentaculitids from Gotland and Scania|journal=Fossils and Strata |volume=11 |pages=180}}</ref>

== See also ==
*[Mari Mari Group](/source/Mari_Mari_Group), fossil formation in the [state of Amazonas](/source/Amazonas_(Brazilian_state)) of northwestern Brazil

== References ==
{{Reflist}}

==Further reading==
{{Commons category|Tentaculites}}
* Treatise on Invertebrate Paleontology, Part W - Miscellanea. ''Geological Society of America and University of Kansas Press'', 1962. [LCCN](/source/Library_of_Congress_Control_Number) 53012913

{{Taxonbar|from=Q4130259}}
{{Authority control}}

Category:Devonian animals
Category:Late Devonian animals
Category:Tentaculita
Category:Early Ordovician first appearances
Category:Late Devonian genus extinctions
Category:Silurian animals of North America
Category:Paleozoic life of Ontario
Category:Paleozoic life of Alberta
Category:Paleozoic life of the Northwest Territories
Category:Paleozoic life of Nova Scotia
Category:Paleozoic life of Quebec
Category:Paleozoic life of Yukon
Category:Prehistoric animal genera

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Adapted from the Wikipedia article [Tentaculites](https://en.wikipedia.org/wiki/Tentaculites) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Tentaculites?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
