# Stuttgart Formation

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Geologic formation in Germany

Stuttgart Formation Stratigraphic range: Carnian ~235–222 Ma PreꞒ Ꞓ O S D C P T J K Pg N Type Formation Unit of Keuper Sub-units Ansbachersandstein & Schilfsandstein members Underlies Weser & Steigerwald Formations Overlies Grabfeld & Benk Formations Lithology Primary Sandstone Other Claystone Location Coordinates 49°12′N 9°18′E / 49.2°N 9.3°E / 49.2; 9.3 Approximate paleocoordinates 21°24′N 14°54′E / 21.4°N 14.9°E / 21.4; 14.9 Region Baden-Württemberg, Bavaria, North Rhine-Westphalia Country Germany, Switzerland Type section Named for Stuttgart Stuttgart Formation (Germany)

The **Stuttgart Formation** is a [geologic formation](/source/Formation_(geology)) in [Germany](/source/Germany). It preserves [fossils](/source/Fossils) dating back to the [Carnian](/source/Carnian) stage of the [Triassic](/source/Triassic) [period](/source/Period_(geology)).[1]

## Fossil content

See also: [Ansbachersandstein § Fossil content](/source/Ansbachersandstein#Fossil_content)

### Temnospondyls

- *[Cyclotosaurus buechneri](/source/Cyclotosaurus)*[2]

- *[C. robustus](/source/Cyclotosaurus)*[3]

- *[Hyperokynodon keuperinus](/source/Hyperokynodon)*[3]

- *[Mastodonsaurus andriani](/source/Mastodonsaurus)*[4]

- *[Metoposaurus diagnosticus](/source/Metoposaurus)*[5]

- *[Gerrothorax sp.](/source/Gerrothorax)*[6]

### Therapsids

- [Kannemeyeriiformes indet.](/source/Kannemeyeriiformes)(Possibly *[Woznikella triradiata](/source/Woznikella)*)[6]

### Reptiles

- *[Calamosuchus arenaceus](/source/Calamosuchus_arenaceus)*[7]

- *[Dyoplax arenaceus](/source/Dyoplax_arenaceus)*

- *[Thuringopelta werneburgi](/source/Thuringopelta)*[8]

- *[Zanclodon subcylindrodon](/source/Zanclodon)*[9]

- [Archosauria indet.](/source/Archosauria)[6]

### Fish

- *[Lissodus sp.](/source/Lissodus)*[6]

- *[Palaeobates sp.](/source/Palaeobates)*[10]

**Invertebrates[10]**

- *[Aviculomyalina angusta](https://en.wikipedia.org/w/index.php?title=Aviculomyalina&action=edit&redlink=1)*

- *[Cyzicus laxitextus](/source/Cyzicus)*

- *[C. cf. dorsorectus](/source/Cyzicus)*

- *[C. minutus](/source/Cyzicus)*

- *[Homomya bilonga](https://en.wikipedia.org/w/index.php?title=Homomya&action=edit&redlink=1)*

- *[H. simplex](https://en.wikipedia.org/w/index.php?title=Homomya&action=edit&redlink=1)*

- *[Myalina rotundata](https://en.wikipedia.org/w/index.php?title=Myalina&action=edit&redlink=1)*

- *[Mytilus acutefinitus](/source/Mytilus_(bivalve))*

- *[M. avirostrum](/source/Mytilus_(bivalve))*

- *[M. hasta](/source/Mytilus_(bivalve))*

- *[M. minutus](/source/Mytilus_(bivalve))*

- *[M. peregrinus](/source/Mytilus_(bivalve))*

- *[M. sulmensis](/source/Mytilus_(bivalve))*

- *[Lithophaga buchhornensis](/source/Lithophaga)*

- *[L. lennachensis](/source/Lithophaga)*

- *[L. producta](/source/Lithophaga)*

- *[L. vermiculata](/source/Lithophaga)*

- *[Mactromya altera](https://en.wikipedia.org/w/index.php?title=Mactromya&action=edit&redlink=1)*

- *[M. equisetitis](https://en.wikipedia.org/w/index.php?title=Mactromya&action=edit&redlink=1)*

- *[Modiolus eberstadtensis](/source/Modiolus_(bivalve))*

- *[M. formosissimus](/source/Modiolus_(bivalve))*

- *[M. mediocarinatus](/source/Modiolus_(bivalve))*

- *[M. minalatus](/source/Modiolus_(bivalve))*

- *[M. parallelus](/source/Modiolus_(bivalve))*

- *[M. parvoblongus](/source/Modiolus_(bivalve))*

- *[M. suprarectus](/source/Modiolus_(bivalve))*

- *[M. transiens](/source/Modiolus_(bivalve))*

- *[M. triangulus](/source/Modiolus_(bivalve))*

- *[Myoconcha aperina](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[M. longaperina](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[M. ovulum](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[M. scalprosa](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[M. cf. woehrmanni](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[M. aff. aquatensis](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[Parallelodon beyrichii](https://en.wikipedia.org/w/index.php?title=Parallelodon&action=edit&redlink=1)*

- *[Pinna mediokeuperina](/source/Pinna_(genus))*

- *[Pleuromya curta](https://en.wikipedia.org/w/index.php?title=Pleuromya&action=edit&redlink=1)*

- *[P. nitens](https://en.wikipedia.org/w/index.php?title=Pleuromya&action=edit&redlink=1)*

- *[Schafhaeutlia aff. liscaviensis](https://en.wikipedia.org/w/index.php?title=Schafhaeutlia&action=edit&redlink=1)*

- *[Thracia keuperina](https://en.wikipedia.org/w/index.php?title=Thracia_(genus)&action=edit&redlink=1)*

- *[Trigonodus cf. grandis](https://en.wikipedia.org/w/index.php?title=Trigonodus&action=edit&redlink=1)*

- *[T. pygmaeus](https://en.wikipedia.org/w/index.php?title=Trigonodus&action=edit&redlink=1)*

- *[T. singularis](https://en.wikipedia.org/w/index.php?title=Trigonodus&action=edit&redlink=1)*

- *[T. wuertembergicus](https://en.wikipedia.org/w/index.php?title=Trigonodus&action=edit&redlink=1)*

- *[Asmussia sp.](https://en.wikipedia.org/w/index.php?title=Asmussia&action=edit&redlink=1)*

- *[Gervillia sp.](/source/Gervillia)*

- *[?Gryphaea sp.](/source/Gryphaea)*

- *[Myoconcha sp.](https://en.wikipedia.org/w/index.php?title=Myoconcha&action=edit&redlink=1)*

- *[Mytilus sp.](/source/Mytilus_(bivalve))*

- *[Trigonodus sp.](https://en.wikipedia.org/w/index.php?title=Trigonodus&action=edit&redlink=1)*

### Flora

- *[Clathrophyllum meriani](https://en.wikipedia.org/w/index.php?title=Clathrophyllum&action=edit&redlink=1)*[10]

- *[Danaeopsis marantacea](https://en.wikipedia.org/w/index.php?title=Danaeopsis&action=edit&redlink=1)*[11]

- *[D. rumphi](https://en.wikipedia.org/w/index.php?title=Danaeopsis&action=edit&redlink=1)*[10]

- *[Desmiophyllum imhoffi](https://en.wikipedia.org/w/index.php?title=Desmiophyllum&action=edit&redlink=1)*[10]

- *[Dioonitocarpidium pennaeforme](https://en.wikipedia.org/w/index.php?title=Dioonitocarpidium&action=edit&redlink=1)*[10]

- *[Neocalamites meriani](/source/Neocalamites)*[10]

- *[Pagiophyllum foetterlei](/source/Pagiophyllum)*[10]

- *[Voltziopsis coburgensis](https://en.wikipedia.org/w/index.php?title=Voltziopsis&action=edit&redlink=1)*[10]

- *[Widdringtonites keuperianus](https://en.wikipedia.org/w/index.php?title=Widdringtonites&action=edit&redlink=1)*[10]

- *[Equisetites sp.](/source/Equisetites)*[10]

- *[Pterophyllum sp.](/source/Pterophyllum)*[10]

## See also

- [List of fossiliferous stratigraphic units in Germany](/source/List_of_fossiliferous_stratigraphic_units_in_Germany)

- [List of fossiliferous stratigraphic units in Switzerland](/source/List_of_fossiliferous_stratigraphic_units_in_Switzerland)

- [Benkersandstein](/source/Benkersandstein), contemporaneous ichnofossiliferous formation of Bavaria

- [Chañares Formation](/source/Cha%C3%B1ares_Formation), fossiliferous formation of the [Ischigualasto-Villa Unión Basin](/source/Ischigualasto-Villa_Uni%C3%B3n_Basin), Argentina

- [Candelária Formation](/source/Candel%C3%A1ria_Formation%2C_Paran%C3%A1_Basin), contemporaneous fossiliferous formation of the Paraná Basin, Brazil

- [Molteno Formation](/source/Molteno_Formation), contemporaneous fossiliferous formation of Lesotho and South Africa

- [Pebbly Arkose Formation](/source/Pebbly_Arkose_Formation), contemporaneous fossiliferous formation of Botswana, Zambia and Zimbabwe

- [Denmark Hill Insect Bed](/source/Denmark_Hill_Insect_Bed), contemporaneous fossiliferous unit of Queensland, Australia

- [Madygen Formation](/source/Madygen_Formation), contemporaneous Lagerstätte of Kyrgyzstan

## References

1. **[^](#cite_ref-FW_1-0)** [Stuttgart Formation](https://paleobiodb.org/classic/displayStrata?geological_group=&formation=Stuttgart&group_formation_member=Stuttgart) at [Fossilworks](/source/Fossilworks).org

1. **[^](#cite_ref-Witzmann2016_2-0)** Witzmann et al., 2016

1. ^ [***a***](#cite_ref-Schoch2000_3-0) [***b***](#cite_ref-Schoch2000_3-1) Schoch & Milner, 2000

1. **[^](#cite_ref-Munster1839_4-0)** Münster, 1839

1. **[^](#cite_ref-Sulej2002_5-0)** Sulej, 2002

1. ^ [***a***](#cite_ref-Schoch2012_6-0) [***b***](#cite_ref-Schoch2012_6-1) [***c***](#cite_ref-Schoch2012_6-2) [***d***](#cite_ref-Schoch2012_6-3) Schoch, 2012

1. **[^](#cite_ref-S&S25_7-0)** Sues, Hans-Dieter; Schoch, Rainer R. (2025-12-04). ["Synopsis of the Triassic reptiles from Germany"](https://doi.org/10.3897%2Ffr.28.164405). *Fossil Record*. **28** (2): 411–483. [doi](/source/Doi_(identifier)):[10.3897/fr.28.164405](https://doi.org/10.3897%2Ffr.28.164405). [ISSN](/source/ISSN_(identifier)) [2193-0074](https://search.worldcat.org/issn/2193-0074).

1. **[^](#cite_ref-8)** Sues, H.-D.; Schoch, R. R. (2025). "A doswelliid archosauriform from the Upper Triassic (Carnian) Stuttgart Formation of Thuringia (Germany)". *Journal of Vertebrate Paleontology*. e2455949. [doi](/source/Doi_(identifier)):[10.1080/02724634.2025.2455949](https://doi.org/10.1080%2F02724634.2025.2455949).

1. **[^](#cite_ref-Huene1905_9-0)** Huene, 1905

1. ^ [***a***](#cite_ref-Linck1968_10-0) [***b***](#cite_ref-Linck1968_10-1) [***c***](#cite_ref-Linck1968_10-2) [***d***](#cite_ref-Linck1968_10-3) [***e***](#cite_ref-Linck1968_10-4) [***f***](#cite_ref-Linck1968_10-5) [***g***](#cite_ref-Linck1968_10-6) [***h***](#cite_ref-Linck1968_10-7) [***i***](#cite_ref-Linck1968_10-8) [***j***](#cite_ref-Linck1968_10-9) [***k***](#cite_ref-Linck1968_10-10) [***l***](#cite_ref-Linck1968_10-11) Linck, 1968

1. **[^](#cite_ref-Kustatscher2012_11-0)** Kustatscher et al., 2012

### Bibliography

- Witzmann, F.; Sachs, S.; Nyhuis, C. J. (2016), ["A new species of *Cyclotosaurus* (Stereospondyli, Capitosauria) from the Late Triassic of Bielefeld, NW Germany, and the intrarelationships of the genus"](https://fr.copernicus.org/articles/19/83/2016/), *[Fossil Record](/source/Fossil_Record)*, **19** (2): 83–100, [Bibcode](/source/Bibcode_(identifier)):[2016FossR..19...83W](https://ui.adsabs.harvard.edu/abs/2016FossR..19...83W), [doi](/source/Doi_(identifier)):[10.5194/fr-19-83-2016](https://doi.org/10.5194%2Ffr-19-83-2016), retrieved 2020-10-19

- Kustatscher, E.; Kelber, K.-P.; Van Konijnenburg-van Cittert, J. H. A. (2012), "*Danaeopsis* Heer ex Schimper 1869 and its European Triassic species", *[Review of Palaeobotany and Palynology](/source/Review_of_Palaeobotany_and_Palynology)*, **183**: 32–49, [Bibcode](/source/Bibcode_(identifier)):[2012RPaPa.183...32K](https://ui.adsabs.harvard.edu/abs/2012RPaPa.183...32K), [doi](/source/Doi_(identifier)):[10.1016/j.revpalbo.2012.06.011](https://doi.org/10.1016%2Fj.revpalbo.2012.06.011)

- Schoch, R. R (2012), "A dicynodont mandible from the Triassic of Germany forms the first evidence of large herbivores in the Central European Carnian", *[Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen](/source/Neues_Jahrbuch_f%C3%BCr_Geologie_und_Pal%C3%A4ontologie_-_Abhandlungen)*, **263** (2): 119–123, [doi](/source/Doi_(identifier)):[10.1127/0077-7749/2012/0216](https://doi.org/10.1127%2F0077-7749%2F2012%2F0216)

- Sulej, T (2002), ["Species discrimination of the Late Triassic temnospondyl amphibian *Metoposaurus diagnosticus*"](https://www.app.pan.pl/archive/published/app47/app47-535.pdf) (PDF), *[Acta Palaeontologica Polonica](/source/Acta_Palaeontologica_Polonica)*, **47**: 535–546, retrieved 2020-10-19

- Schoch, R.; Milner, A. R. (2000), "Stereospondyli", *[Handbuch der Paläoherpetologie](https://en.wikipedia.org/w/index.php?title=Handbuch_der_Pal%C3%A4oherpetologie&action=edit&redlink=1)*, **3B**: 1–203

- Linck, O (1968), "Die marinen Muschelfauna des Schilfsandsteins von Eberstadt, Wuerttemberg, (Trias, Karn, mittl. Keuper 2) und deren Bedeutung", *[Jahreshefte des Vereins für Vaterlaendsiche Naturkunde in Wuerttemberg](https://en.wikipedia.org/w/index.php?title=Jahreshefte_des_Vereins_f%C3%BCr_Vaterlaendsiche_Naturkunde_in_Wuerttemberg&action=edit&redlink=1)*, **123**: 69–133

- [Huene](/source/Friedrich_von_Huene), F. von (1905), "Trias-Dinosaurier Europas - European Triassic dinosaurs", *[Zeitschrift der Deutschen Geologischen Gesellschaft](https://en.wikipedia.org/w/index.php?title=Zeitschrift_der_Deutschen_Geologischen_Gesellschaft&action=edit&redlink=1)*, **57**: 345–349

- Fraas, E (1896), *Die Schwäbischen Trias-Saurier nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt - Swabian Triassic dinosaurs based on the material in the Royal Natural History Collection compiled in Stuttgart*, Festgabe des Königlichen Naturalien-Cabinets In Stuttgart zur 42 Versammlung der Deutschen geologischen Gesellschaft in Stuttgart, August 1896. E. Schweizerbartische Verlag-handlung (E. Koch), Stuttgart, pp. 1–18

- Münster, G. G (1839), "*Mastodonsaurus andriani*", *[Beiträge zur Petrefactenkunde](https://en.wikipedia.org/w/index.php?title=Beitr%C3%A4ge_zur_Petrefactenkunde&action=edit&redlink=1)*: 102–103

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Adapted from the Wikipedia article [Stuttgart Formation](https://en.wikipedia.org/wiki/Stuttgart_Formation) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Stuttgart_Formation?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.