# Stipeae
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> Source: https://en.wikipedia.org/wiki/Stipeae
> Source revision: 1290709213
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{{Short description|Tribe of grasses}}
{{Automatic taxobox
|image = Stipa gigantea 3.JPG
|image_caption = ''[Celtica gigantea](/source/Celtica_gigantea)''
|display_parents = 3
|taxon = Stipeae
|authority = [Dumort.](/source/Dumort.)
|synonyms =
* supertribe Stipodae L. Liu
* subtribe Stipinae Griseb.
* Aciachninae Caro
* Ortachninae Caro
|synonyms_ref = [Barkworth ME, Arriaga MO, Smith JF, Jacobs SWL, Valdes-Reyna J, Bushman BS (2008) Molecules and Morphology in South American Stipeae (Poaceae). Systematic Botany 33(4), 719-731.]
==Description==
The defining morphological features of the Stipeae include single-flowered [spikelets](/source/spikelets) lacking a [rachilla](/source/Spikelet) extension, and the [lemma](/source/lemma_(botany))s (the external bract) have either a sharp point or a terminal [awn](/source/Awn_(botany)) (long bristle).[Cialdella AM, Giussani LM, Aagesen L, Zuloaga FO, Morrone O (2007) A phylogeny of Piptochaetium (Poaceae : Pooideae : Stipeae) and related genera based on a combined analysis including trnL-F, rp116, and morphology. Systematic Botany 32(3), 545-559.]
==Genera==
The tribe includes 32 genera:[{{cite journal|last1=Soreng|first1=Robert J.|last2=Peterson|first2=Paul M.|last3=Romaschenko|first3=Konstantin|last4=Davidse|first4=Gerrit|last5=Teisher|first5=Jordan K.|last6=Clark|first6=Lynn G.|last7=Barberá|first7=Patricia|last8=Gillespie|first8=Lynn J.|last9=Zuloaga|first9=Fernando O.|title=A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications|journal=Journal of Systematics and Evolution|volume=55|issue=4|year=2017|pages=259–290|issn=1674-4918|doi=10.1111/jse.12262|doi-access=free|hdl=10261/240149|hdl-access=free}} {{open access}}][Peterson PM, Romaschenko K, Soreng RJ, Valdés Reyna J (2019) A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of ''Eriocoma, Neotrinia, Oloptum'', and five new genera: ''Barkworthia, × Eriosella, Pseudoeriocoma, Ptilagrostiella'', and ''Thorneochloa''. ''PhytoKeys'' 126: 89-125. https://doi.org/10.3897/phytokeys.126.34096][{{cite web |title=Poaceae Barnhart |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30000032-2 |website=[Plants of the World Online](/source/Plants_of_the_World_Online) |publisher=[Royal Botanic Gardens, Kew](/source/Royal_Botanic_Gardens%2C_Kew) |access-date=27 October 2024}}]
{{columns-list|colwidth=22em|
*''[Achnatherum](/source/Achnatherum)'' {{small|P.Beauv.}} (syn. ''Aristella'')
*''[Aciachne](/source/Aciachne)'' {{small|Benth.}}
*''[Amelichloa](/source/Amelichloa)'' {{small|Arriaga & Barkworth}}
*''[Anemanthele](/source/Anemanthele)'' {{small|Veldkamp}}
*''[Austrostipa](/source/Austrostipa)'' {{small|S.W.L.Jacobs & J.Everett}}
*''[Barkworthia](/source/Barkworthia)'' {{small|Romasch., P.M.Peterson & Soreng}}
*''[Celtica](/source/Celtica_(plant))'' {{small|F.M.Vázquez & Barkworth}}
*''[Eriocoma](/source/Eriocoma)'' {{small|Nutt.}}
*''[× Eriosella](/source/%C3%97_Eriosella)'' {{small|Romasch.}}
*''[Hesperostipa](/source/Hesperostipa)'' {{small|(Elias) Barkworth}}
*''[Jarava](/source/Jarava)'' {{small|Ruiz & Pav.}}
*''[Lorenzochloa](/source/Lorenzochloa)'' {{small|Reeder & C.Reeder}}
*''[Macrochloa](/source/Macrochloa)'' {{small|Kunth}}
*''[Nassella](/source/Nassella)'' {{small|(Trin.) É.Desv.}}
*''[Neotrinia](/source/Neotrinia)'' {{small|(Tzvelev) M.Nobis, P.D.Gudkova & A.Nowak}}
*''[Oloptum](/source/Oloptum)'' {{small|Röser & Hamasha}}
*''[Ortachne](/source/Ortachne)'' {{small|Nees ex Steud.}}
*''[Orthoraphium](/source/Orthoraphium)'' {{small|Nees}}
*''[Oryzopsis](/source/Oryzopsis)'' {{small|Michx.}}
*''[Pappostipa](/source/Pappostipa)'' {{small|(Speg.) Romasch., P.M.Peterson & Soreng}}
*''[Patis](/source/Patis_(plant))'' {{small|Ohwi}}
*''[Piptatheropsis](/source/Piptatheropsis)'' {{small|Romasch., P.M.Peterson & Soreng}}
*''[Piptatherum](/source/Piptatherum)'' {{small|P.Beauv.}}
*''[Piptochaetium](/source/Piptochaetium)'' {{small|J.Presl.}}
*''[Psammochloa](/source/Psammochloa)'' {{small|Htchc.}}
*''[Pseudoeriocoma](/source/Pseudoeriocoma)'' {{small|Romasch., P.M.Peterson & Soreng}}
*''[Ptilagrostiella](/source/Ptilagrostiella)'' {{small|Romasch., P.M.Peterson & Soreng}}
*''[Ptilagrostis](/source/Ptilagrostis)'' {{small|Griseb.}}
*''[Stipa](/source/Stipa)'' {{small|L.}}
*''[Stipellula](/source/Stipellula)'' {{small|Röser & Hamasha}} (syn. ''Stipella'')
*''[Thorneochloa](/source/Thorneochloa)'' {{small|Romasch., P.M.Peterson & Soreng}}
*''[Timouria](/source/Timouria)'' {{small|Roshev.}}
*''[Trikeraia](/source/Trikeraia)'' {{small|Bor}}
}}
Many species initially placed into ''Stipa'' have now been split off into new genera. Some recent papers have analysed relationships within and between the genera,[{{cite journal|last1=Barber|first1=Janet C.|last2=Hames|first2=Kasey A.|last3=Cialdella|first3=Ana Maria|last4=Giussani|first4=Liliana M.|last5=Morrone|first5=Osvaldo|title=Phylogenetic relationships of Piptochaetium Presl (Poaceae: Stipeae) and related genera reconstructed from nuclear and chloroplast sequence datasets |journal=Taxon|volume=58|issue=2|year=2009|pages=375–380|issn=0040-0262|doi=10.1002/tax.582005}} [https://www.researchgate.net/profile/Liliana_Giussani/publication/233665373_Phylogenetic_relationships_of_Piptochaetium_Presl_Poaceae_Stipeae_and_related_genera_reconstructed_from_nuclear_and_chloroplast_sequence_datasets/links/552e76700cf22d43716e78ae/Phylogenetic-relationships-of-Piptochaetium-Presl-Poaceae-Stipeae-and-related-genera-reconstructed-from-nuclear-and-chloroplast-sequence-datasets.pdf pdf]][Jacobs SWL, Bayer R, Everett J, Arriaga MO, Barkworth ME, Sabin-Badereau A, Torres MA, Vazquez FM, Bagnall N (2007) Systematics of the tribe Stipeae (Gramineae) using molecular data. Aliso 23, 349-361.][Jacobs SWL, Everett J, Barkworth ME, Hsiao C (2000) Relationships within the Stipoid grasses (Gramineae). In 'Grass Systematics and Evolution.' (Eds SWL Jacobs and J Everett) pp. 75-82. (CSIRO: Melbourne)] but a complete analysis has not yet been performed. Stipoid grasses use the [C3 photosynthetic pathway](/source/C3_photosynthesis) and live in temperate areas worldwide.[{{cite journal|author=Romaschenko K, Peterson PM, Soreng RJ, Garcia-Jacas N, Futorna O, Susanna A |date=2008| title=Molecular phylogenetic analysis of the American Stipeae (Poaceae) resolves ''Jarava'' sensu lato polyphyletic: evidence for a new genus, ''Pappostipa''|journal= Journal of the Botanical Research Institute of Texas|volume= 2|issue=1|pages= 165–192|jstor=41971613}} [https://repository.si.edu/bitstream/handle/10088/6448/Romaschenko_etal_Pappostipa_highres.pdf?sequence=1&isAllowed=y pdf]]
Known [fossil](/source/fossil)s date from the late [Miocene](/source/Miocene).[{{cite journal|last1=Thomasson|first1=Joseph R.|title=Berriochloa Gabeli and Berrichloa Huletti (Graminae: Stipeae), two new grass species from the late miocene Ash Hollow Formationof Nebraska and Kansas|journal=Journal of Paleontology|volume=79|issue=1|year=2005|pages=185–199|issn=0022-3360|doi=10.1666/0022-3360(2005)079<0185:BGABHG>2.0.CO;2|bibcode=2005JPal...79..185T }} [https://www.researchgate.net/profile/Joseph_Thomasson/publication/250070847_Berriochloa_gabeli_and_Berriochloa_huletti_Gramineae_Stipeae_two_new_grass_species_from_the_late_miocene_ash_hollow_formation_of_Nebraska_and_Kansas/links/56324ae608ae584878091c65/Berriochloa-gabeli-and-Berriochloa-huletti-Gramineae-Stipeae-two-new-grass-species-from-the-late-miocene-ash-hollow-formation-of-Nebraska-and-Kansas.pdf pdf]]
== References ==
{{Reflist}}
== External links ==
* [https://www.delta-intkey.com/grass/www/stipa-sl.htm Stipa L. sensu lato] (''sensu lato'' refers to the tribe) ''The grass genera of the world'' website by L. Watson, [T.D. Macfarlane](/source/Terry_Desmond_Macfarlane), and M.J. Dallwitz
*{{Commons category-inline|Stipeae}}
*{{Wikispecies-inline|Stipeae}}
{{Taxonbar|from=Q7617501}}
Category:Stipeae
Category:Poaceae tribes
Category:Taxa described in 1824
Category:Taxa named by Barthélemy Charles Joseph Dumortier
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Adapted from the Wikipedia article [Stipeae](https://en.wikipedia.org/wiki/Stipeae) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Stipeae?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.