# Saccoloma

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{{Short description|Genus of ferns}}
{{Automatic taxobox
| image = Saccoloma inaequale herbarium.jpg
| image_caption = ''[Saccoloma inaequale](/source/Saccoloma_inaequale)''<br/> herbarium specimen
| taxon = Saccoloma
| authority = [Kaulf.](/source/Georg_Friedrich_Kaulfuss)
| type_species = ''Saccoloma elegans''
| type_species_authority = [Kaulf.](/source/Georg_Friedrich_Kaulfuss)
| subdivision_ranks = [Species](/source/Species)
| subdivision = See text
| synonyms_ref = <ref name=POWO_30010104-2/>
| synonyms =
* ''Neuropteris'' <small>Desv.</small>
* ''Orthiopteris'' <small>Copel.</small>
}}

'''''Saccoloma''''' is a [fern](/source/fern) genus in family [Saccolomataceae](/source/Saccolomataceae).<ref name=PPGI/><ref name=Phyto>[http://www.mapress.com/phytotaxa/content/2011/f/pt00019p054.pdf Christenhusz ''et al.'' "A linear sequence of extant families and genera of lycophytes and ferns"] ''Phytotaxa'' 19: 7-54. 18 Feb. 2011</ref> It is the only genus in the family in the [Pteridophyte Phylogeny Group](/source/Pteridophyte_Phylogeny_Group) classification of 2016 (PPG I), but further investigation is needed.<ref name=PPGI/> It is [pantropical](/source/pantropical) and its species are found in wet, shaded forest areas.<ref>[http://www.metodosenecologiaysistematica.org/vol5/vol5%281%29/saccoloma5%281%29.pdf Rojas-Alvarado "Novelties in the Saccoloma inaequale complex (Saccolomataceae) from the neotropics"] ''Métodos en Ecología y Sistemática'' 5(1): 1-16. Apr 2010</ref> ''Saccoloma'' species are characterized by an omega-shaped (Ω) vascular bundle in the cross-sections of their [petioles](/source/petiole_(botany)).<ref>[http://www.plantsystematics.org/imgs/robbin/r/Dennstaedtiaceae_Saccoloma_moranii_28675.html ''Saccoloma inaequale'' petiole] Image at PlantSystematics.org 01 Feb 2012</ref><ref>[http://www.plantsystematics.org/imgs/robbin/r/Dennstaedtiaceae_Saccoloma_dominguensis_39778.html ''Saccoloma dominguense'' petiole] Image at PlantSystematics.org 01 Feb 2012</ref> The common name '''soralpouch fern''' is used for ''Saccoloma''.<ref>{{PLANTS|id=SACCO|taxon=Saccoloma|accessdate=26 October 2015}}</ref>

==Phylogeny==
''Saccoloma'' includes ''Orthiopteris'' in the [Pteridophyte Phylogeny Group](/source/Pteridophyte_Phylogeny_Group) classification of 2016 (PPG I), with a total of 18 species.<ref name=PPGI/> {{As of|2019|August}}, [Plants of the World Online](/source/Plants_of_the_World_Online) accepts the following species (which may be based on a wider [circumscription](/source/Circumscription_(taxonomy)) of the genus). Synonyms are taken from the ''Checklist of Ferns and Lycophytes of the World''.<ref name=CFLW/>

Phylogeny of ''Saccoloma''<ref>{{cite journal |last1=Nitta |first1=Joel H. |last2=Schuettpelz |first2=Eric |last3=Ramírez-Barahona |first3=Santiago |last4=Iwasaki |first4=Wataru |display-authors=et al. |year=2022 |title=An Open and Continuously Updated Fern Tree of Life |journal=Frontiers in Plant Science |volume=13 |issue= |article-number= 909768| doi=10.3389/fpls.2022.909768 |pmid= 36092417|pmc= 9449725|bibcode= |doi-access=free}}</ref><ref>{{cite web |last1= |first1= |display-authors=et al. |year=2024 |title=Tree viewer: interactive visualization of FTOL |url=https://fernphy.github.io/viewer.html |version=FTOL v1.7.0 ([GenBank](/source/GenBank) release 261) |access-date=1 May 2025}}</ref>
{{Clade | style=font-size:90%;line-height:80%
|1={{clade
  |label1=''Orthiopteris''
  |1={{clade
    |1=''O. campylura'' <small>(Kunze) Copel.</small> (syn. ''Saccoloma campylurum'' <small>(Kunze) Mett.</small>; ''Saccoloma caudatum'' <small>Copel.</small>; ''Saccoloma kingii'' <small>(Bedd.) G.B.Nair</small>)
    |2=''O. ferulacea'' <small>(T. Moore) Copel.</small> (syn. ''Saccoloma ferulaceum'' <small>(T.Moore) R.M.Tryon & A.F.Tryon</small>)
     }}
  |label2=''Saccoloma''
  |2={{clade
    |1={{clade
      |1=''S. galeottii'' <small>(Fée) A.Rojas</small>
      |2=''S. sunduei'' <small>A.Rojas</small>
       }}
    |2={{clade
      |1={{clade
        |1=''S. domingense'' <small>(Spr.) Prantl</small>
        |2=''S. elegans'' <small>Kaulf.</small> (syn. ''Saccoloma chartaceum'' <small>G.B.Nair</small>)
         }}
      |2={{clade
        |1=''S. inaequale'' <small>(Kunze) Mett.</small>
        |2={{clade
          |1=''S. brasiliense'' <small>(C.Presl) Mett.</small>
          |2=''S. nigrescens'' <small>(Mett.) A.Rojas</small>
           }}
         }}
       }}
     }}
   }}
}}

Other species include:
*''Saccoloma acuminatum'' <small>(Rosenst.) Christenh.</small> (syn. ''Orthiopteris acuminata''<small>(Rosenst.) Copel.</small>)
*''Saccoloma cicutarioides'' <small>(Baker) Christenh.</small> (syn. ''Orthiopteris cicutarioides'' <small>(Baker) Copel.</small>)
*''Saccoloma firmum'' <small>(Kuhn) C.Chr.</small> (syn. ''Orthiopteris firma'' <small>(Kuhn) Brownlie</small>)
*''Saccoloma henriettae'' <small>(Baker) C.Chr.</small> (syn. ''Orthiopteris henriettae'' <small>(Baker) Copel.</small>)
*''[Saccoloma laxum](/source/Saccoloma_laxum)'' <small>R.C.Moran & B.Øllg.</small>
*''Saccoloma membranaceum'' <small>Mickel</small>
*''Saccoloma moranii'' <small>A.Rojas</small>
*''Saccoloma quadripinnatum'' <small>A.Rojas</small>
*''Saccoloma samoense'' <small>(Hovenkamp & T.T.Luong) Christenh.</small> (syn. ''Orthiopteris samoensis'' <small>Hovenk. & T. T. Luong</small>)
*''[Saccoloma squamosum](/source/Saccoloma_squamosum)'' <small>R.C.Moran</small>
*''Saccoloma tenue'' <small>(Brack.) Mett.</small> (syn. ''Orthiopteris tenuis'' <small>(Brack.) Brownlie</small>)
*''Saccoloma trichophyllum'' <small>(Copel.) G.B.Nair</small> (syn. ''Orthiopteris trichophylla'' <small>Copel.</small>)

==References==
{{Reflist|refs=
<ref name=CFLW>{{Citation |mode=cs1|last1=Hassler |first1=Michael |last2=Schmitt |first2=Bernd |date=June 2019 |title=Checklist of Ferns and Lycophytes of the World |volume=8 |issue=2 |url=https://worldplants.webarchiv.kit.edu/ferns/ |access-date=2019-08-14 |name-list-style=amp }}</ref>

<ref name=POWO_30010104-2>{{cite web |title=''Saccoloma'' Kaulf. |work=Plants of the World Online |publisher=Royal Botanic Gardens, Kew|url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30010104-2 |access-date=2019-08-14 }}</ref>

<ref name=PPGI>{{citation |mode=cs1|author=PPG I |year=2016 |title=A community-derived classification for extant lycophytes and ferns |journal=Journal of Systematics and Evolution |volume=54 |issue=6 |pages=563–603 |doi=10.1111/jse.12229|s2cid=39980610 |doi-access=free }}</ref>
}}

{{Commons}}
{{Wikispecies}}
{{Plant classification}}
{{Fern classification}}
{{Taxonbar|from=Q310724}}
Category:Polypodiales
Category:Fern genera
Category:Pantropical flora
Category:Taxonomy articles created by Polbot

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Adapted from the Wikipedia article [Saccoloma](https://en.wikipedia.org/wiki/Saccoloma) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Saccoloma?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.