# Pyrgini

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The **Pyrgini** are a [tribe](/source/Tribe_(biology)) in the [skipper butterfly](/source/Skipper_butterfly) [subfamily](/source/Subfamily) [Pyrginae](/source/Pyrginae). Formerly, when only four tribes of Pyrginae were recognized, the Pyrgini contained the largest number of [genera](/source/Genera) among these. But this overly wide delimitation has since turned out to be [paraphyletic](/source/Paraphyletic).[1]

One of the traditional Pyrginae tribes, the Eudamini, had to be raised to subfamily rank as [Eudaminae](/source/Eudaminae). Some genera now in the Eudaminae were placed in the Pyrgini in earlier times. In addition, a number of additional tribes – [Achlyodidini](/source/Achlyodidini), [Carcharodini](/source/Carcharodini) and [Erynnini](/source/Erynnini) – are now usually recognized again. These are close relatives of the Pyrgini *[sensu stricto](/source/Sensu_stricto)*, and may just as well be included in them as they used to be: together they do still form a [monophyletic](/source/Monophyletic) group. Most authors prefer to keep them separate however, as each is an [apomorphic](/source/Apomorph) and [biogeographically](/source/Biogeographic) distinct lineage of the Pyrgini *[sensu lato](/source/Sensu_lato)*. Of these newly recognized tribes, the Achlyodidini are closest to the Pyrgini, but conspicuously differ from the latter in their unusually-shaped wings.[1]

## Genera

The genera of the Pyrgini, after genomic research published in 2019, are:[2][1][3]

- *[Anisochoria](/source/Anisochoria)* Mabille, 1877
- *[Antigonus](/source/Antigonus_(butterfly))* Hübner, 1819
- *[Bralus](/source/Bralus)* Grishin, 2019[4]
- *[Burnsius](/source/Burnsius)* Grishin, 2019 (New World checkered-skippers)
- *[Canesia](/source/Canesia)* Grishin, 2019[4]
- *[Carrhenes](/source/Carrhenes)* Godman & Salvin, 1895
- *[Celotes](/source/Celotes)* Godman & Salvin, 1899 (streaky-skippers)
- *[Chirgus](/source/Chirgus)* Grishin, 2019
- *[Clito](/source/Clito_(butterfly))* Evans, 1953
- *[Diaeus](/source/Diaeus_(butterfly))* Godman & Salvin, 1895
- *[Eracon](/source/Eracon)* Godman & Salvin, 1894
- *[Heliopetes](/source/Heliopetes)* Billberg, 1820 (white-skippers)
- *[Onenses](/source/Onenses)* Godman & Salvin, 1895
- *[Paches](/source/Paches)* Godman & Salvin, 1895
- *[Plumbago](/source/Plumbago_(butterfly))* Evans, 1953
- *[Pseudodrephalys](/source/Pseudodrephalys)* Burns, 1998
- *[Pyrgus](/source/Pyrgus)* Hübner, 1819 (checkered-skippers)
- *[Santa](/source/Santa_(butterfly))* Grishin, 2019[4]
- *[Spioniades](/source/Spioniades)* Hübner, 1819
- *[Systasea](/source/Systasea)* W. H. Edwards, 1877 (powdered-skippers)
- *[Trina](/source/Trina_(butterfly))* Evans, 1953
- *[Xenophanes](/source/Xenophanes_(butterfly))* Godman & Salvin, 1895
- *[Zobera](/source/Zobera)* Freeman, 1970
- *[Zopyrion](/source/Zopyrion_(butterfly))* Godman & Salvin, 1896

## Footnotes

1. Brower & Warren (2007)

1. Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". *Proceedings of the National Academy of Sciences*. **116** (13): 6232–6237. [Bibcode:2019PNAS..116.6232L](https://ui.adsabs.harvard.edu/abs/2019PNAS..116.6232L). [doi:10.1073/pnas.1821304116](https://doi.org/10.1073/pnas.1821304116). [PMC 6442542](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6442542). [PMID 30877254](https://pubmed.ncbi.nlm.nih.gov/30877254)

1. Pelham, Jonathan P. (2021). [*A Catalogue of the Butterflies of the United States and Canada*](http://butterfliesofamerica.com/US-Can-Cat.htm)

1. Cong, Qian; Zhang, Jing; Shen, Jinhui; Grishin, Nick V. (2019-11-06). "Fifty new genera of Hesperiidae (Lepidoptera)". *Insecta Mundi*. **2019**: 0731. [doi:10.5281/zenodo.3677235](https://doi.org/10.5281/zenodo.3677235). [ISSN 0749-6737](https://www.worldcat.org/issn/0749-6737). [PMC 8791444](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8791444). [PMID 35087260](https://pubmed.ncbi.nlm.nih.gov/35087260)

## References

- (2007): [Tree of Life Web Project](/source/Tree_of_Life_Web_Project) – [Pyrgini](http://tolweb.org/Pyrgini/95263). Version of 2007-MAR-04. Retrieved 2009-DEC-26.

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Adapted from the Wikipedia article [Pyrgini](https://en.wikipedia.org/wiki/Pyrgini) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Pyrgini?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.