# Protobothrops > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Protobothrops > Markdown URL: https://mediated.wiki/source/Protobothrops.md > Source: https://en.wikipedia.org/wiki/Protobothrops > Source revision: 1351248598 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) Genus of snakes Protobothrops Protobothrops cornutus Scientific classification Kingdom: Animalia Phylum: Chordata Class: Reptilia Order: Squamata Suborder: Serpentes Family: Viperidae Subfamily: Crotalinae Genus: Protobothrops Hoge & Romano-Hoge, 1983 Type species Bothrops flavoviridis Hallowell, 1861 ***Protobothrops*** is a [genus](/source/Genus) of [pit vipers](/source/Pit_viper) found in Asia.[1] It is also known as the **lance headed pit vipers**.[2] The [venom](/source/Venom) of *Protobothrops* can be diverse and unique across individual organisms of the same and differing species and has demonstrated the ability to evolve in an accelerated manner.[3][4] One possible explanation for these traits is that the [habitat](/source/Habitat) in which *Protobothrops* reside contains a large variety of prey which may have prompted the diversification of the [venom](/source/Venom) related [genes](/source/Gene).[3][4] ## Species Image Scientific name Distribution Protobothrops cornutus (M.A. Smith, 1930) Vietnam and Southern China. Protobothrops dabieshanensis Huang, T. Pan, Han, L. Zhang, Hou, Yu, Zheng & B. Zhang, 2012 China Protobothrops elegans (Gray, 1849) Ryukyu Islands, specifically the Yaeyama Islands. Protobothrops flavirostris Grassby-Lewis, Brakels, Maury, Sitthivong, Frohlich, Pawangkhanant, Idiiatullina, Nguyen & Poyarkov, 2025[5] Vang Vieng District, Vientiane Province, Laos Protobothrops flavoviridis (Hallowell, 1861) Ryukyu Islands, including Okinawa and the Amami Islands. Protobothrops himalayanus H. Pan, Chettri, D. Yang, K. Jiang, K. Wang, L. Zhang & Vogel, 2013 China (southern Tibet), Nepal, Bhutan, India (Sikkim) Protobothrops jerdonii (Günther, 1875) India, Nepal, Myanmar, China, and Vietnam. Protobothrops kaulbacki (M.A. Smith, 1940) Myanmar, China Protobothrops kelomohy Sumontha, Vasaruchapong, Chomngam, Suntrarachun, Pawangkhanant, Sompan, Smit, Kunya & Chanhome, 2020 Northern Thailand Protobothrops mangshanensis (Zhao, 1990) China (Guangdong, Hunan) Protobothrops maolanensis J. Yang, Orlov & Y. Wang, 2011 China (Guizhou) Protobothrops mucrosquamatus (Cantor, 1839) India (Assam and Mizoram) and Bangladesh, to Myanmar, China (including Hainan, and as far north as Gansu and as far east as Zhejiang), Laos, northern and central Vietnam, also found in northern Thailand as well as in Taiwan Protobothrops sieversorum (Ziegler, Herrmann, David, Orlov & Pauwels, 2000) Vietnam Protobothrops tokarensis (Nagai, 1928) Kagoshima, Kyushu and the Ryukyu Islands Protobothrops trungkhanhensis Orlov, Ryabov & Nguyen, 2009 Vietnam, China Protobothrops xiangchengensis (Zhao, Y. Jiang & Huang, 1979) China (Yunnan, Sichuan) *[Nota bene](/source/Nota_bene)*: A [binomial authority](/source/Binomial_nomenclature) in parentheses indicates that the species was originally described in a genus other than *Protobothrops*. ## References 1. **[^](#cite_ref-RDB_1-0)** *[Protobothrops](http://reptile-database.reptarium.cz/search.php?submit=Search&genus=Protobothrops)* at the [Reptarium.cz Reptile Database](http://reptile-database.reptarium.cz/). Accessed 13 August 2022.. 1. **[^](#cite_ref-2)** Grassby-Lewis, Rupert J.; Brakels, Peter; Maury, Nathanaël; Sitthivong, Saly; Frohlich, David; Pawangkhanant, Parinya; Idiiatullina, Sabira S.; Nguyen, Tan Van; Poyarkov, Nikolay A. (2025-02-25). ["A new endemic karst-associated species of lance-headed pit viper (Squamata, Viperidae, Protobothrops) from Laos"](https://herpetozoa.pensoft.net/article/146004/). *Herpetozoa*. **38**: 43–60. [doi](/source/Doi_(identifier)):[10.3897/herpetozoa.38.e146004](https://doi.org/10.3897%2Fherpetozoa.38.e146004). [ISSN](/source/ISSN_(identifier)) [2682-955X](https://search.worldcat.org/issn/2682-955X). 1. ^ [***a***](#cite_ref-:0_3-0) [***b***](#cite_ref-:0_3-1) Shibata, Hiroki; Chijiwa, Takahito; Oda-Ueda, Naoko; Nakamura, Hitomi; Yamaguchi, Kazuaki; Hattori, Shousaku; Matsubara, Kazumi; Matsuda, Yoichi; Yamashita, Akifumi; Isomoto, Akiko; Mori, Kazuki (2018-07-26). ["The habu genome reveals accelerated evolution of venom protein genes"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6062510). *Scientific Reports*. **8** (1): 11300. [Bibcode](/source/Bibcode_(identifier)):[2018NatSR...811300S](https://ui.adsabs.harvard.edu/abs/2018NatSR...811300S). [doi](/source/Doi_(identifier)):[10.1038/s41598-018-28749-4](https://doi.org/10.1038%2Fs41598-018-28749-4). [PMC](/source/PMC_(identifier)) [6062510](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6062510). [PMID](/source/PMID_(identifier)) [30050104](https://pubmed.ncbi.nlm.nih.gov/30050104). 1. ^ [***a***](#cite_ref-:1_4-0) [***b***](#cite_ref-:1_4-1) Inamaru, Kento; Takeuchi, Ami; Maeda, Marie; Shibata, Hiroki; Fukumaki, Yasuyuki; Oda-Ueda, Naoko; Hattori, Shosaku; Ohno, Motonori; Chijiwa, Takahito (2020-03-12). ["Discovery of the Gene Encoding a Novel Small Serum Protein (SSP) of *Protobothrops flavoviridis* and the Evolution of SSPs"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7150969). *Toxins*. **12** (3): 177. [doi](/source/Doi_(identifier)):[10.3390/toxins12030177](https://doi.org/10.3390%2Ftoxins12030177). [PMC](/source/PMC_(identifier)) [7150969](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7150969). [PMID](/source/PMID_(identifier)) [32178380](https://pubmed.ncbi.nlm.nih.gov/32178380). 1. **[^](#cite_ref-5)** Grassby-Lewis, Rupert J.; Brakels, Peter; Maury, Nathanaël; Sitthivong, Saly; Frohlich, David; Pawangkhanant, Parinya; Idiiatullina, Sabira S.; Nguyen, Tan Van; Poyarkov, Nikolay A. (2025-02-25). ["A new endemic karst-associated species of lance-headed pit viper (Squamata, Viperidae, Protobothrops) from Laos"](https://herpetozoa.pensoft.net/article/146004/element/2/15//). *Herpetozoa*. **38**: 43–60. [doi](/source/Doi_(identifier)):[10.3897/herpetozoa.38.e146004](https://doi.org/10.3897%2Fherpetozoa.38.e146004). [ISSN](/source/ISSN_(identifier)) [2682-955X](https://search.worldcat.org/issn/2682-955X). Taxon identifiers Protobothrops Wikidata: Q2697677 Wikispecies: Protobothrops ADW: Protobothrops CoL: 6XLB EoL: 41189 GBIF: 2444145 iNaturalist: 30994 IRMNG: 1020246 ITIS: 1058406 NCBI: 103943 Open Tree of Life: 267855 This snake article is a stub. 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