# Phaseoleae

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Tribe of legumes

Phaseoleae Phaseolus coccineus Scientific classification Kingdom: Plantae Clade: Embryophytes Clade: Tracheophytes Clade: Spermatophytes Clade: Angiosperms Clade: Eudicots Clade: Rosids Order: Fabales Family: Fabaceae Subfamily: Faboideae Clade: Meso-Papilionoideae Clade: Non-protein amino acid-accumulating clade Clade: Millettioids Tribe: Phaseoleae Bronn ex DC. (1825) Subtribes See text Synonyms Phaseolinae Bronn (1822) Erythrineae (Benth.) Hassk. (1844) Glycineae Burnett (1835)

The [plant](/source/Plant) [tribe](/source/Tribe_(biology)) **Phaseoleae** is one of the subdivisions of the [legume](/source/Legume) [subfamily](/source/Subfamily) [Faboideae](/source/Faboideae), in the unranked [NPAAA clade](/source/Non-protein_amino_acid-accumulating_clade). This group includes many of the [beans](/source/Bean) cultivated for human and animal food, most importantly from the [genera](/source/Genus) *[Glycine](/source/Glycine_(plant))*, *[Phaseolus](/source/Phaseolus)*, and *[Vigna](/source/Vigna)*.

## Taxonomy

Although the tribe as defined in the late 20th century does not appear to be [monophyletic](/source/Monophyletic), there does seem to be a monophyletic group which roughly corresponds to the tribe Phaseoleae (with some changes). The earlier concept of Phaseoleae is [paraphyletic](/source/Paraphyly) relative to the tribes [Abreae](/source/Abreae) and [Psoraleeae](/source/Psoraleeae), plus most of [Millettieae](/source/Millettieae) and parts of [Desmodieae](/source/Desmodieae).[1][2]

The following subtribes and [genera](/source/Genera) are recognized by the [USDA](/source/USDA):[3]

**Cajaninae[4]**

- *[Adenodolichos](/source/Adenodolichos)* Harms

- *[Bolusafra](/source/Bolusafra)* Kuntze

- *[Cajanus](/source/Cajanus)* Adans.

- *[Carrissoa](/source/Carrissoa)* Baker f.

- *[Chrysoscias](/source/Chrysoscias)* E. Mey.[a]

- *[Dunbaria](/source/Dunbaria)* Wight & Arn.

- *[Eriosema](/source/Eriosema)* (DC.) Desv.

- *[Paracalyx](/source/Paracalyx)* Ali

- *[Rhynchosia](/source/Rhynchosia)* Lour.

**Clitoriinae[5]**

- *[Barbieria](/source/Barbieria)* DC.

- *[Centrosema](/source/Centrosema)* (DC.) Benth.

- *[Clitoria](/source/Clitoria)* L.

- *[Clitoriopsis](/source/Clitoriopsis)* R. Wilczek

- *[Periandra](/source/Periandra)* Mart. *ex* Benth.

**Diocleinae[6]**

- *[Bionia](/source/Bionia)* Mart. *ex* Benth.[b]

- *[Camptosema](/source/Camptosema)* Hook. & Arn.

- *[Canavalia](/source/Canavalia)* Adans.

- *[Cleobulia](/source/Cleobulia)* Mart. *ex* Benth.

- *[Collaea](/source/Collaea)* DC.

- *[Cratylia](/source/Cratylia)* Mart. *ex* Benth.

- *[Cymbosema](/source/Cymbosema)* Benth.[c]

- *[Dioclea](/source/Dioclea_(plant))* Kunth

- *[Galactia](/source/Galactia)* P. Browne

- *[Lackeya](/source/Lackeya)* Fortunato *et al.*[d]

- *[Luzonia](/source/Luzonia)* Elmer

- *[Macropsychanthus](/source/Macropsychanthus)* Harms *ex* K. Schum. & Lauterb.

- *[Neorudolphia](/source/Neorudolphia)* Britton

- *[Rhodopis](/source/Rhodopis_(plant))* Urb.

**Glycininae[7]**

- *[Afroamphica](/source/Afroamphica)* H.Ohashi & K.Ohashi[8]

- *[Amphicarpaea](/source/Amphicarpaea)* Elliott *ex* Nutt.

- *[Calopogonium](/source/Calopogonium)* Desv.

- *[Cologania](/source/Cologania)* Kunth

- *[Dumasia](/source/Dumasia)* DC.

- *[Eminia](/source/Eminia_(plant))* Taub.

- *[Glycine](/source/Glycine_(plant))* Willd.

- *[Herpyza](/source/Herpyza)* C. Wright

- *[Neocollettia](/source/Neocollettia)* Hemsl.

- *[Neonotonia](/source/Neonotonia)* J. A. Lackey

- *[Neorautanenia](/source/Neorautanenia)* Schinz

- *[Neustanthus](/source/Neustanthus)* Benth.

- *[Nogra](/source/Nogra)* Merr.

- *[Pachyrhizus](/source/Pachyrhizus)* Rich. *ex* DC.

- *[Phylacium](/source/Phylacium)* Benn.

- *[Pseudeminia](/source/Pseudeminia)* Verdc.

- *[Pseudovigna](/source/Pseudovigna)* (Harms) Verdc.

- *[Pueraria](/source/Pueraria)* DC.

- *[Sinodolichos](/source/Sinodolichos)* Verdc.

- *[Teramnus](/source/Teramnus)* P. Browne

- *[Teyleria](/source/Teyleria)* Backer

- *[Toxicopueraria](/source/Toxicopueraria)* A.N. Egan & B. Pan bis[9][10]

- *[Weizhia](/source/Weizhia)* G.Y.Li, Z.H.Chen, K.W.Jiang & B.Pan bis[11]

**Kennediinae[12]**

- *[Hardenbergia](/source/Hardenbergia)* Benth.

- *[Kennedia](/source/Kennedia)* Vent.

- *[Vandasina](/source/Vandasina)* Rauschert

**Ophrestiinae[13]**

- *[Cruddasia](/source/Cruddasia)* Prain

- *[Ophrestia](/source/Ophrestia)* H. M. L. Forbes

- *[Pseudoeriosema](/source/Pseudoeriosema)* Hauman

**Phaseolinae[14][15]**

- *[Alistilus](/source/Alistilus)* N. E. Br.

- *[Ancistrotropis](/source/Ancistrotropis)* A. Delgado[e]

- *[Austrodolichos](/source/Austrodolichos)* Verdc.

- *[Cochliasanthus](/source/Cochliasanthus)* Trew[e]

- *[Condylostylis](/source/Condylostylis)* Piper[e]

- *[Dipogon](/source/Dipogon_(plant))* Liebm.

- *[Dolichopsis](/source/Dolichopsis)* Hassl.

- *[Dolichos](/source/Dolichos_(plant))* L.

- *[Helicotropis](/source/Helicotropis)* A. Delgado[e]

- *[Lablab](/source/Lablab)* Adans.

- *[Leptospron](/source/Leptospron)* (Benth.) A. Delgado[e]

- *[Macroptilium](/source/Macroptilium)* (Benth.) Urb.

- *[Macrotyloma](/source/Macrotyloma)* (Wight & Arn.) Verdc.

- *[Mysanthus](/source/Mysanthus)* G. P. Lewis & A. Delgado

- *[Nesphostylis](/source/Nesphostylis)* Verdc.

- *[Oryxis](/source/Oryxis)* A. Delgado & G. P. Lewis

- *[Oxyrhynchus](/source/Oxyrhynchus_(plant))* Brandegee

- *[Phaseolus](/source/Phaseolus)* L.

- *[Physostigma](/source/Physostigma)* Balf.

- *[Ramirezella](/source/Ramirezella_(plant))* Rose

- *[Sigmoidotropis](/source/Sigmoidotropis)* (Piper) A. Delgado[e]

- *[Spathionema](/source/Spathionema)* Taub.

- *[Sphenostylis](/source/Sphenostylis)* E. Mey.

- *[Strophostyles](/source/Strophostyles)* Elliott

- *[Vatovaea](/source/Vatovaea)* Chiov.

- *[Vigna](/source/Vigna)* Savi

- *[Wajira](/source/Wajira)* Thulin

***incertae sedis***

- *[Apios](/source/Apios)* Fabr.

- *[Butea](/source/Butea)* Roxb. *ex* Willd.

- *[Cochlianthus](/source/Cochlianthus)* Benth.

- *[Decorsea](/source/Decorsea)* R. Vig.

- *[Diphyllarium](/source/Diphyllarium)* Gagnep.

- *[Dysolobium](/source/Dysolobium)* (Benth.) Prain

- *[Erythrina](/source/Erythrina)* L.

- *[Haymondia](/source/Haymondia)* A.N. Egan & B. Pan bis[9][16][f]

- *[Mastersia](/source/Mastersia)* Benth.

- *[Meizotropis](/source/Meizotropis)* Voigt[g]

- *[Mucuna](/source/Mucuna)* Adans.

- *[Otoptera](/source/Otoptera)* DC.

- *[Psophocarpus](/source/Psophocarpus)* Neck. *ex* DC.

- *[Shuteria](/source/Shuteria)* Wight & Arn.

- *[Spatholobus](/source/Spatholobus)* Hassk.

- *[Strongylodon](/source/Strongylodon)* Vogel

## Notes

1. **[^](#cite_ref-5)** Possibly synonymous with *[Rhynchosia](/source/Rhynchosia)*

1. **[^](#cite_ref-8)** Possibly synonymous with *[Camptosema](/source/Camptosema)*

1. **[^](#cite_ref-9)** Possibly synonymous with *[Dioclea](/source/Dioclea_(plant))*

1. **[^](#cite_ref-10)** Possibly synonymous with *[Galactia](/source/Galactia)*

1. ^ [***a***](#cite_ref-exVigna_20-0) [***b***](#cite_ref-exVigna_20-1) [***c***](#cite_ref-exVigna_20-2) [***d***](#cite_ref-exVigna_20-3) [***e***](#cite_ref-exVigna_20-4) [***f***](#cite_ref-exVigna_20-5) American genera formerly part of *[Vigna](/source/Vigna)*[15]

1. **[^](#cite_ref-22)** USDA/GRIN places it under Glycininae following *Pueraria* DC., in conflict with A.N. Egan & B. Pan.[10]

1. **[^](#cite_ref-23)** Possibly synonymous with *[Butea](/source/Butea)*

## See also

- [Helena M. L. Forbes](/source/Helena_M._L._Forbes)

## References

1. **[^](#cite_ref-1)** Wojciechowski MF, Lavin M, Sanderson MJ (2004). ["A phylogeny of legumes (Leguminosae) based on analysis of the plastid *matK* gene resolves many well-supported subclades within the family"](https://doi.org/10.3732%2Fajb.91.11.1846). *[Am J Bot](/source/American_Journal_of_Botany)*. **91** (11): 1846–1862. [Bibcode](/source/Bibcode_(identifier)):[2004AmJB...91.1846W](https://ui.adsabs.harvard.edu/abs/2004AmJB...91.1846W). [doi](/source/Doi_(identifier)):[10.3732/ajb.91.11.1846](https://doi.org/10.3732%2Fajb.91.11.1846). [PMID](/source/PMID_(identifier)) [21652332](https://pubmed.ncbi.nlm.nih.gov/21652332).

1. **[^](#cite_ref-2)** Li H, Wang W, Lin L, Zhu X, Li J, Zhu X, Chen Z (2013). ["Diversification of the phaseoloid legumes: Effects of climate change, range expansion and habit shift"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3793175). *Frontiers in Plant Science*. **4** (386): 1–8. [Bibcode](/source/Bibcode_(identifier)):[2013FrPS....4..386L](https://ui.adsabs.harvard.edu/abs/2013FrPS....4..386L). [doi](/source/Doi_(identifier)):[10.3389/fpls.2013.00386](https://doi.org/10.3389%2Ffpls.2013.00386). [PMC](/source/PMC_(identifier)) [3793175](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3793175). [PMID](/source/PMID_(identifier)) [24130564](https://pubmed.ncbi.nlm.nih.gov/24130564).

1. **[^](#cite_ref-GRIN-Phaseoleae_3-0)** ["Tribe Phaseoleae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomyfamily?type=tribe&id=1571). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021. [follow tribe links and genera lists for the accepted genera in each tribe]

1. **[^](#cite_ref-GRIN-Cajaninae_4-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Cajaninae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1507&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-GRIN-Clitoriinae_6-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Clitoriinae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1511&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-GRIN-Diocleinae_7-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Diocleinae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1515&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-GRIN-Glycininae_11-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Glycininae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1520&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-12)** [A Phylogenetic Study of Amphicarpaea with a New Genus Afroamphica](https://www.researchgate.net/publication/331283552_A_Phylogenetic_Study_of_Amphicarpaea_with_a_New_Genus_Afroamphica_Leguminosae_Tribe_Phaseoleae)

1. ^ [***a***](#cite_ref-Egan1_13-0) [***b***](#cite_ref-Egan1_13-1) Egan AN, Pan B (2015). ["Resolution of polyphyly in *Pueraria* (Leguminosae, Papilionoideae): The creation of two new genera, *Haymondia* and *Toxicopueraria*, the resurrection of *Neustanthus*, and a new combination in *Teyleria*"](https://doi.org/10.11646%2Fphytotaxa.218.3.1). *[Phytotaxa](/source/Phytotaxa)*. **218** (3): 201–226. [Bibcode](/source/Bibcode_(identifier)):[2015Phytx.218..201E](https://ui.adsabs.harvard.edu/abs/2015Phytx.218..201E). [doi](/source/Doi_(identifier)):[10.11646/phytotaxa.218.3.1](https://doi.org/10.11646%2Fphytotaxa.218.3.1).

1. ^ [***a***](#cite_ref-Egan2_14-0) [***b***](#cite_ref-Egan2_14-1) Egan AN, Vatanparast M, Cagle W (2016). ["Parsing polyphyletic *Pueraria*: Delimiting distinct evolutionary lineages through phylogeny"](http://thescholarship.ecu.edu/bitstream/10342/4191/1/Cagle_ecu_0600M_11006.pdf) (PDF). *Molecular Phylogenetics and Evolution*. **104**: 44–59. [Bibcode](/source/Bibcode_(identifier)):[2016MolPE.104...44E](https://ui.adsabs.harvard.edu/abs/2016MolPE.104...44E). [doi](/source/Doi_(identifier)):[10.1016/j.ympev.2016.08.001](https://doi.org/10.1016%2Fj.ympev.2016.08.001). [hdl](/source/Hdl_(identifier)):[10342/4191](https://hdl.handle.net/10342%2F4191). [PMID](/source/PMID_(identifier)) [27495827](https://pubmed.ncbi.nlm.nih.gov/27495827).

1. **[^](#cite_ref-15)** Jiang, K.-W., Chen, Y., Li, H.-D., Ma, D.-D., Pan, B., Song, Z.-Q., Wan, K.-Y., Chen, Z.-H. & Li, G.-Y. (2023) *Weizhia* (Fabaceae: Phaseoleae), a new endemic genus from East China supported by morphological and molecular evidence. *Phytotaxa* 603 (3): 235–248. [https://doi.org/10.11646/phytotaxa.603.3.3](https://doi.org/10.11646/phytotaxa.603.3.3)

1. **[^](#cite_ref-GRIN-Kennediinae_16-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Kennediinae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1522&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-GRIN-Ophrestiinae_17-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Ophrestiinae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1525&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. **[^](#cite_ref-GRIN-Phaseolinae_18-0)** ["Genera and generic subdivisons of Fabaceae Lindl., nom. cons.: Phaseolinae"](https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomygenuslist?id=1527&type=subtribe). *[Germplasm Resources Information Network](/source/Germplasm_Resources_Information_Network) (GRIN)*. [Agricultural Research Service](/source/Agricultural_Research_Service) (ARS), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture) (USDA). Retrieved 27 October 2021.

1. ^ [***a***](#cite_ref-Delgado-Salinas-2011_19-0) [***b***](#cite_ref-Delgado-Salinas-2011_19-1) Delgado-Salinas A, Thulin M, Pasquet R, Weeden N, Lavin M (2011). "*Vigna* (Leguminosae) *sensu lato*: the names and identities of the American segregate genera". *[Am J Bot](/source/American_Journal_of_Botany)*. **98** (10): 1694–715. [Bibcode](/source/Bibcode_(identifier)):[2011AmJB...98.1694D](https://ui.adsabs.harvard.edu/abs/2011AmJB...98.1694D). [doi](/source/Doi_(identifier)):[10.3732/ajb.1100069](https://doi.org/10.3732%2Fajb.1100069). [PMID](/source/PMID_(identifier)) [21980163](https://pubmed.ncbi.nlm.nih.gov/21980163).

1. **[^](#cite_ref-Egan3_21-0)** Egan AN, Puttock CF (2016). ["The genus *Haymondia* A.N.Egan & B.Pan bis (Fabaceae) in Thailand"](https://doi.org/10.20531%2Ftfb.2016.44.1.06). *Thai Forest Bulletin (Botany)*. **44** (1): 26–31. [doi](/source/Doi_(identifier)):[10.20531/tfb.2016.44.1.06](https://doi.org/10.20531%2Ftfb.2016.44.1.06).

## External links

- Data related to [Phaseoleae](https://species.wikimedia.org/wiki/Special:Search/Phaseoleae) at Wikispecies

- Media related to [Phaseoleae](https://commons.wikimedia.org/wiki/Category:Phaseoleae) at Wikimedia Commons

Taxon identifiers Phaseoleae Wikidata: Q141198 Wikispecies: Phaseoleae APNI: 4230028 FNA: 20648 GRIN: 1571 iNaturalist: 507503 NCBI: 163735 Open Tree of Life: 563857 Tropicos: 50133648 VASCAN: 419

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Adapted from the Wikipedia article [Phaseoleae](https://en.wikipedia.org/wiki/Phaseoleae) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Phaseoleae?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
