# Phascolarctidae

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Family of marsupials

Phascolarctidae[1] Temporal range: Late Oligocene–Recent PreꞒ Ꞓ O S D C P T J K Pg N Phascolarctos cinereus Scientific classification Kingdom: Animalia Phylum: Chordata Class: Mammalia Infraclass: Marsupialia Order: Diprotodontia Suborder: Vombatiformes Infraorder: Phascolarctomorphia Aplin & Archer, 1987 Family: Phascolarctidae Owen, 1839 Genera †Invictokoala †Litokoala †Lumakoala? †Koobor †Madakoala †Nimiokoala †Perikoala †Priscakoala †Stelakoala Phascolarctos

The **Phascolarctidae** (*φάσκωλος (phaskolos)* - pouch or bag, *ἄρκτος (arktos)* - bear, from the Greek *phascolos* + *arctos* meaning pouched bear) is a [family](/source/Family_(taxonomy)) of [marsupials](/source/Marsupialia) of the order [Diprotodontia](/source/Diprotodontia), consisting of only one extant species, the [koala](/source/Koala),[1] and six well-known fossil species, with another six less well known fossil species, and two fossil species of the genus *Koobor*, whose taxonomy is debatable but are placed in this group. The closest relatives of the Phascolarctidae are the [wombats](/source/Wombat), which comprise the family [Vombatidae](/source/Vombatidae).[2]

The fossil record of the family dates back to the [Middle Miocene](/source/Middle_Miocene)[3] or [Late Oligocene](/source/Late_Oligocene).[4]

## Classification

**Family Phascolarctidae**[1]

- Genus †*[Nimiokoala](/source/Nimiokoala)* - †*[Nimiokoala greystanesi](https://en.wikipedia.org/w/index.php?title=Nimiokoala_greystanesi&action=edit&redlink=1)*

- Genus †*[Invictokoala](/source/Invictokoala)*[5] - †*Invictokoala monticola*

- Genus †*[Madakoala](/source/Madakoala)* - †*Madakoala robustus* - †*Madakoala wellsi* - †*Madakoala devisi*

- Genus †*[Litokoala](/source/Litokoala)* - †*Litokoala garyjohnstoni* - †*Litokoala kutjamarpensis* - †*Litokoala kanunkaensis*

- Genus †*[Lumakoala](/source/Lumakoala)* - †*Lumakoala blackae*[6]

- Genus †*[Koobor](/source/Koobor)* - †*Koobor jimbarrati* - †*Koobor notabilis*

- Genus †*[Perikoala](/source/Perikoala)* - †*Perikoala palankarinnica* - †*Perikoala robustus*

- Genus *[Phascolarctos](/source/Phascolarctos)* - †*Phascolarctos maris* - [Koala](/source/Koala) - *Phascolarctos cinereus* - †[Giant koala](/source/Giant_koala) - *Phascolarctos stirtoni* - †*Phascolarctos yorkensis* (formerly *Cundokoala yorkensis*)[7]

- Genus †*[Priscakoala](/source/Priscakoala)* - †*Priscakoala lucyturnbullae*[4]

- Genus †*[Stelakoala](https://en.wikipedia.org/w/index.php?title=Stelakoala&action=edit&redlink=1)*[*[citation needed](https://en.wikipedia.org/wiki/Wikipedia:Citation_needed)*] - †*[Stelakoala riversleighensis](https://en.wikipedia.org/w/index.php?title=Stelakoala_riversleighensis&action=edit&redlink=1)*

## References

1. ^ [***a***](#cite_ref-msw3_1-0) [***b***](#cite_ref-msw3_1-1) [***c***](#cite_ref-msw3_1-2) [Groves, C. P.](/source/Colin_Groves) (2005). ["Family Phascolarctidae"](http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=11000003). In [Wilson, D. E.](/source/Don_E._Wilson); Reeder, D. M (eds.). [*Mammal Species of the World: A Taxonomic and Geographic Reference*](http://www.google.com/books?id=JgAMbNSt8ikC&pg=PA43) (3rd ed.). Johns Hopkins University Press. p. 43. [ISBN](/source/ISBN_(identifier)) [978-0-8018-8221-0](https://en.wikipedia.org/wiki/Special:BookSources/978-0-8018-8221-0). [OCLC](/source/OCLC_(identifier)) [62265494](https://search.worldcat.org/oclc/62265494).

1. **[^](#cite_ref-msw3_b_2-0)** [Groves, C. P.](/source/Colin_Groves) (2005). ["Suborder Vombatiformes"](http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=11000002). In [Wilson, D. E.](/source/Don_E._Wilson); Reeder, D. M (eds.). [*Mammal Species of the World: A Taxonomic and Geographic Reference*](http://www.google.com/books?id=JgAMbNSt8ikC&pg=PA43-44) (3rd ed.). Johns Hopkins University Press. pp. 43–44. [ISBN](/source/ISBN_(identifier)) [978-0-8018-8221-0](https://en.wikipedia.org/wiki/Special:BookSources/978-0-8018-8221-0). [OCLC](/source/OCLC_(identifier)) [62265494](https://search.worldcat.org/oclc/62265494).

1. **[^](#cite_ref-3)** Benton, M.J. (2005). *Vertebrate Palaeontology*. Oxford. p. 314.

1. ^ [***a***](#cite_ref-Black2012_4-0) [***b***](#cite_ref-Black2012_4-1) [Karen Black](/source/Karen_H._Black); et al. (2012). "New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity". *Journal of Vertebrate Paleontology*. **32** (1): 125–138. [Bibcode](/source/Bibcode_(identifier)):[2012JVPal..32..125B](https://ui.adsabs.harvard.edu/abs/2012JVPal..32..125B). [doi](/source/Doi_(identifier)):[10.1080/02724634.2012.626825](https://doi.org/10.1080%2F02724634.2012.626825). [S2CID](/source/S2CID_(identifier)) [86152273](https://api.semanticscholar.org/CorpusID:86152273).

1. **[^](#cite_ref-Invictokoala_5-0)** Gilbert J. Price and Scott A. Hocknull (2011). "*Invictokoala monticola* gen. et sp. nov. (Phascolarctidae, Marsupialia), a Pleistocene plesiomorphic koala holdover from Oligocene ancestors". *Journal of Systematic Palaeontology*. **9** (2): 327–335. [Bibcode](/source/Bibcode_(identifier)):[2011JSPal...9..327P](https://ui.adsabs.harvard.edu/abs/2011JSPal...9..327P). [doi](/source/Doi_(identifier)):[10.1080/14772019.2010.504079](https://doi.org/10.1080%2F14772019.2010.504079). [S2CID](/source/S2CID_(identifier)) [84316375](https://api.semanticscholar.org/CorpusID:84316375).

1. **[^](#cite_ref-6)** Arthur I. Crichton, Robin M. D. Beck, Aidan M. C. Couzens, Trevor H. Worthy, Aaron B. Camens and Gavin J. Prideaux; et al. (2023). ["A probable koala from the Oligocene of central Australia provides insights into early diprotodontian evolution"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10477348). *Scientific Reports*. **13** (1). 14521. [Bibcode](/source/Bibcode_(identifier)):[2023NatSR..1314521C](https://ui.adsabs.harvard.edu/abs/2023NatSR..1314521C). [doi](/source/Doi_(identifier)):[10.1038/s41598-023-41471-0](https://doi.org/10.1038%2Fs41598-023-41471-0). [PMC](/source/PMC_(identifier)) [10477348](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10477348). [PMID](/source/PMID_(identifier)) [37666885](https://pubmed.ncbi.nlm.nih.gov/37666885).{{[cite journal](https://en.wikipedia.org/wiki/Template:Cite_journal)}}: CS1 maint: multiple names: authors list ([link](https://en.wikipedia.org/wiki/Category:CS1_maint:_multiple_names:_authors_list))

1. **[^](#cite_ref-Talent_7-0)** Talent, John A. (2012). *Earth and Life*. [Springer](/source/Springer_Science%2BBusiness_Media). p. 1047. [ISBN](/source/ISBN_(identifier)) [9789048134281](https://en.wikipedia.org/wiki/Special:BookSources/9789048134281).

## External links

Wikimedia Commons has media related to [Phascolarctidae](https://commons.wikimedia.org/wiki/Category:Phascolarctidae).

[Wikispecies](/source/Wikispecies) has information related to ***[Phascolarctidae](https://species.wikimedia.org/wiki/Phascolarctidae)***.

- [Koalas with evolutionary information](https://koalas.org/koala-info.html).

v t e Extant Diprotodontia species Kingdom Animalia Phylum Chordata Class Mammalia Infraclass Marsupialia Suborder Vombatiformes Phascolarctidae Phascolarctos Koala (P. cinereus) Vombatidae (wombats) Vombatus Common wombat (V. ursinus) Lasiorhinus Southern hairy-nosed wombat (L. latifrons) Northern hairy-nosed wombat (L. krefftii) Suborder Phalangeriformes (possums) Phalangeridae (including cuscuses) Ailurops (bear cuscuses) Talaud bear cuscus (A. melanotis) Sulawesi bear cuscus (A. ursinus) Phalanger Gebe cuscus (P. alexandrae) Mountain cuscus (P. carmelitae) Ground cuscus (P. gymnotis) Eastern common cuscus (P. intercastellanus) Woodlark cuscus (P. lullulae) Blue-eyed cuscus (P. matabiru) Telefomin cuscus (P. matanim) Southern common cuscus (P. mimicus) Northern common cuscus (P. orientalis) Ornate cuscus (P. ornatus) Rothschild's cuscus (P. rothschildi) Silky cuscus (P. sericeus) Stein's cuscus (P. vestitus) Spilocuscus Admiralty Island cuscus (S. kraemeri) Common spotted cuscus (S. maculatus) Waigeou cuscus (S. papuensis) Black-spotted cuscus (S. rufoniger) Blue-eyed spotted cuscus (S. wilsoni) Strigocuscus Sulawesi dwarf cuscus (S. celebensis) Banggai cuscus (S. pelegensis) Trichosurus (brushtail possums) Northern brushtail possum (T. arnhemensis) Short-eared possum (T. caninus) Mountain brushtail possum (T. cunninghami) Coppery brushtail possum (T. johnstonii) Common brushtail possum (T. vulpecula) Wyulda Scaly-tailed possum (W. squamicaudata) Burramyidae (pygmy possums) Burramys Mountain pygmy possum (B. parvus) Cercartetus Long-tailed pygmy possum (C. caudatus) Western pygmy possum (C. concinnus) Tasmanian pygmy possum (C. lepidus) Eastern pygmy possum (C. nanus) Tarsipedidae Tarsipes Honey possum (T. rostratus) Petauridae Dactylopsila Great-tailed triok (D. megalura) Long-fingered triok (D. palpator) Tate's triok (D. tatei) Striped possum (D. trivirgata) Gymnobelideus Leadbeater's possum (G. leadbeateri) Petaurus Northern glider (P. abidi) Yellow-bellied glider (P. australis) Biak glider (P. biacensis) Sugar glider (P. breviceps) Mahogany glider (P. gracilis) Squirrel glider (P. norfolcensis) Pseudocheiridae Hemibelideus Lemur-like ringtail possum (H. lemuroides) Petauroides (greater gliders) Central greater glider (P. armillatus) Northern greater glider (P. minor) Southern greater glider (P. volans) Petropseudes Rock-haunting ringtail possum (P. dahli) Pseudocheirus Common ringtail possum (P. peregrinus) Pseudochirulus Lowland ringtail possum (P. canescens) Weyland ringtail possum (P. caroli) Cinereus ringtail possum (P. cinereus) Painted ringtail possum (P. forbesi) Herbert River ringtail possum (P. herbertensis) Masked ringtail possum (P. larvatus) Pygmy ringtail possum (P. mayeri) Vogelkop ringtail possum (P. schlegeli) Pseudochirops D'Albertis's ringtail possum (P. albertisii) Green ringtail possum (P. archeri) Plush-coated ringtail possum (P. corinnae) Reclusive ringtail possum (P. coronatus) Coppery ringtail possum (P. cupreus) Acrobatidae Acrobates Feathertail glider (A. pygmaeus) Distoechurus Feather-tailed possum (D. pennatus) Suborder Macropodiformes Macropodidae (includes wallabies) Lagostrophus Banded hare-wallaby (L. fasciatus) Dendrolagus (tree-kangaroos) Bennett's tree-kangaroo (D. bennettianus) Doria's tree-kangaroo (D. dorianus) Goodfellow's tree-kangaroo (D. goodfellowi) Grizzled tree-kangaroo (D. inustus) Lumholtz's tree-kangaroo (D. lumholtzi) Matschie's tree-kangaroo (D. matschiei) Dingiso (D. mbaiso) Ifola (D. notatus) Golden-mantled tree-kangaroo (D. pulcherrimus) Lowlands tree-kangaroo (D. spadix) Tenkile (D. scottae) Seri's tree-kangaroo (D. stellarum) Ursine tree-kangaroo (D. ursinus) Dorcopsis Black dorcopsis (D. atrata) White-striped dorcopsis (D. hageni) Gray dorcopsis (D. luctuosa) Brown dorcopsis (D. muelleri) Dorcopsulus Macleay's dorcopsis (D. macleayi) Small dorcopsis (D. vanheurni) Lagorchestes (hare-wallabies) Spectacled hare-wallaby (L. conspicillatus) Rufous hare-wallaby (L. hirsutus) Macropus Western grey kangaroo (M. fuliginosus) Eastern grey kangaroo (M. giganteus) Notamacropus Agile wallaby (N. agilis) Black-striped wallaby (N. dorsalis) Tammar wallaby (N. eugenii) Western brush wallaby (N. irma) Parma wallaby (N. parma) Whiptail wallaby (N. parryi) Red-necked wallaby (N. rufogriseus) Onychogalea (nail-tail wallabies) Bridled nail-tail wallaby (O. fraenata) Northern nail-tail wallaby (O. unguifera) Osphranter Antilopine kangaroo (O. antilopinus) Black wallaroo (O. bernardus) Common wallaroo (O. robustus) Red kangaroo (O. rufus) Petrogale (rock-wallabies) P. brachyotis species group: Short-eared rock-wallaby (P. brachyotis) Monjon (P. burbidgei) Nabarlek (P. concinna) Eastern short-eared rock-wallaby (P. wilkinsi) P. xanthopus species group: Proserpine rock-wallaby (P. persephone) Rothschild's rock-wallaby (P. rothschildi) Yellow-footed rock-wallaby (P. xanthopus) P. lateralis/penicillata species group: Allied rock-wallaby (P. assimilis) Cape York rock-wallaby (P. coenensis) Godman's rock-wallaby (P. godmani) Herbert's rock-wallaby (P. herberti) Unadorned rock-wallaby (P. inornata) Black-flanked rock-wallaby (P. lateralis) Mareeba rock-wallaby (P. mareeba) Brush-tailed rock-wallaby (P. penicillata) Purple-necked rock-wallaby (P. purpureicollis) Mount Claro rock-wallaby (P. sharmani) Setonix Quokka (S. brachyurus) Thylogale (pademelons) Tasmanian pademelon (T. billardierii) Brown's pademelon (T. browni) Dusky pademelon (T. brunii) Calaby's pademelon (T. calabyi) Mountain pademelon (T. lanatus) Red-legged pademelon (T. stigmatica) Red-necked pademelon (T. thetis) Wallabia Swamp wallaby (W. bicolor) Potoroidae Aepyprymnus Rufous rat-kangaroo (A. rufescens) Bettongia (bettongs) Eastern bettong (B. gaimardi) Boodie (B. lesueur) Woylie (B. ogilbyi) Northern bettong (B. tropica) Potorous (potoroos) Long-footed potoroo (P. longipes) Long-nosed potoroo (P. tridactylus) Gilbert's potoroo (P. gilbertii) Hypsiprymnodontidae Hypsiprymnodon Musky rat-kangaroo (H. moschatus)

v t e Vombatiformes Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Diprotodontia †Thylacoleonidae Enigmaleo Lekaneleo Microleo Thylacoleo Wakaleo Phascolarctidae †Invictokoala †Koobor †Litokoala †Madakoala †Nimiokoala †Perikoala Phascolarctos †Priscakoala Vombatomorphia †Thylacoleonidae? †Ilariidae Ilaria Kuterintja †Wynyardiidae Muramura Namilamadeta Wynyardia Vombatoidea †Marada? †Mukupirna Vombatidae Lasiorhinus †Nimbavombatus †Phascolonus †Ramsayia †Rhizophascolonus †Sedophascolomys Vombatus †Warendja †Diprotodontoidea Palorchestidae Palorchestes Propalorchestes Diprotodontidae Alkwertatherium Ambulator Diprotodon Euowenia Euryzygoma Hulitherium Kolopsis Kolopsoides Maokopia Meniscolophus Neohelos Ngapakaldia Nimbadon Nototherium Pitikantia Plaisiodon Pyramios Raemeotherium Silvabestius Sthenomerus Zygomaturus

Taxon identifiers Phascolarctidae Wikidata: Q431978 Wikispecies: Phascolarctidae ADW: Phascolarctidae AFD: Phascolarctidae BioLib: 33119 BOLD: 81250 CoL: 6268B EoL: 1700 EPPO: 1PHOSF GBIF: 9373 iNaturalist: 42981 IRMNG: 103175 ITIS: 552319 MSW: 11000003 NCBI: 38624 Open Tree of Life: 327628 Paleobiology Database: 40173

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