# Mouse-eared bat

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{{Short description|Genus of bats}}
{{For|the species sometimes known as the '''mouse eared bat'''|Greater mouse-eared bat}}
{{redirect-distinguish|Myotis|Mitosis|Myositis}}
{{Automatic taxobox
| name = Mouse-eared bats
| fossil_range = [Early Oligocene](/source/Early_Oligocene) to present, {{fossil range|33|0|ref=<ref name="Gunnel2017"/>}}
| image = Myotis mystacinus.jpg
| image_caption = [Whiskered bat](/source/Whiskered_bat) (''Myotis mystacinus'')
| image_alt = A whiskered bat (''Myotis mystacinus''), who is quite displeased at being handled.
| taxon = Myotis
| authority = [Kaup](/source/Johann_Jakob_Kaup), 1829
| type_species = ''[Vespertilio myotis](/source/Vespertilio_myotis)''
| type_species_authority = [Borkhausen](/source/Moritz_Balthasar_Borkhausen), 1797
| subdivision_ranks = Species
| subdivision = See text
}}
The '''mouse-eared bats''' or '''myotises''' are a diverse and widespread [genus](/source/genus) (''Myotis'') of [bat](/source/bat)s within the family [Vespertilionidae](/source/Vespertilionidae). The noun "''myotis''" itself is a [Neo-Latin](/source/Neo-Latin) construction, from the [Greek](/source/Greek_language) "''muós'' (meaning "mouse") and "''oûs''" (meaning ear), literally translating to "mouse-eared".<ref>{{cite book |url= https://books.google.com/books?id=uEWl0ZM6DfUC |page= 69 |title= The Wild Mammals of Missouri |first1= Charles Walsh |last1= Schwartz |first2= Elizabeth Reeder |last2= Schwartz |edition= illustrated |publisher= University of Missouri Press |year= 2001 |isbn= 978-0-8262-1359-4}}</ref>

==Relationships==
''Myotis'' has historically been included in the subfamily [Vespertilioninae](/source/Vespertilioninae), but was classified in its own subfamily, [Myotinae](/source/Myotinae), by [Nancy Simmons](/source/Nancy_Simmons) in 1998. In her 2005 classification in ''[Mammal Species of the World](/source/Mammal_Species_of_the_World)'', Simmons listed the genera ''[Cistugo](/source/Cistugo)'' and ''[Lasionycteris](/source/Lasionycteris)'' in the Myotinae in addition to ''Myotis'' itself.<ref name=S499>Simmons, 2005, p. 499</ref> However, molecular data indicate that ''Cistugo'' is distantly related to all other Vespertilionidae, so  it was reclassified into its own family, the Cistugidae,<ref>Lack et al., 2010</ref> and that ''Lasionycteris'' belongs in the Vespertilioninae.<ref name=Rea2010>Roehrs et al., 2010</ref> The genus ''[Submyotodon](/source/Submyotodon)'' has since been added to the subfamily, making it and ''Myotis'' its only members.<ref name="Ruedi 2015">{{cite journal|doi=10.11646/zootaxa.3920.2.6|pmid=25781252|url=https://www.researchgate.net/publication/272830083|title=Molecular phylogeny and morphological revision of Myotis bats (Chiroptera: Vespertilionidae) from Taiwan and adjacent China|journal=Zootaxa|volume=3920|issue=2|pages=301–342|date=2015-02-20|last1=Ruedi|first1=Manuel|last2=Csorba|first2=Gábor|last3=Lin|first3=Liang-Kong|last4=Chou|first4=C-H}}</ref>

==Appearance and behavior==
Their ears are normally longer than they are wide, with a long and lance-shaped [tragus](/source/Tragus_(ear)), hence their English and zoological names. The species within this genus vary in size from very large to very small for [vesper bat](/source/vesper_bat)s, with a single pair of mammary glands.

Mouse-eared bats are generally [insectivore](/source/insectivore)s. ''[M. vivesi](/source/Myotis_vivesi)'', and several members of the trawling bat ecomorph ''Leuconoe'', have relatively large feet with long toes, and take small fish from the water surface (they also take insects).<ref>Levin, E.; A. Barnea; Y. Yovel; and Y. Yom-Tov (2006). Have introduced fish initiated piscivory among the long-fingered bat? Mammalian Biology 71(3): 139–143.</ref>

==Longevity==
''Myotis'' species are remarkably long-lived for their size; in 2018, researchers <!-- at [University College Dublin](/source/University_College_Dublin) --> revealed that a longitudinal study appears to indicate that ''Myotis'' [telomere](/source/telomere)s do not shrink with age, and that [telomerase](/source/telomerase) does not appear to be present in the ''Myotis'' metabolism. 13 species of ''Myotis'' bats live longer than 20 years and 4 species live longer than 30 years.<ref name=Telomerase>[https://www.science.org/doi/10.1126/sciadv.aao0926 Growing old, yet staying young: The role of telomeres in bats' exceptional longevity], {{doi|10.1126/sciadv.aao0926}}</ref><ref name=Motherboard>[https://www.vice.com/en/article/bats-dont-die-old-age-telomeres-longevity/ These Bats Don't Seem to Die of Old Age—Can They Help Extend the Human Lifespan?], by Kate Lunau, at ''[Vice](/source/Vice_(magazine))''; published February 7, 2018; retrieved June 12, 2018</ref> The longest-living species of ''Myotis'', and longest-living bat in general, is thought to be the [Siberian bat](/source/Siberian_bat) (''M. sibiricus''); one individual discovered in 2005 was found to be over 41 years old at the time.<ref>{{Cite journal|last1=Podlutsky|first1=A. J.|last2=Khritankov|first2=A. M.|last3=Ovodov|first3=N. D.|last4=Austad|first4=S. N.|date=2005-11-01|title=A New Field Record for Bat Longevity|journal=The Journals of Gerontology Series A: Biological Sciences and Medical Sciences|volume=60|issue=11|pages=1366–1368|doi=10.1093/gerona/60.11.1366|pmid=16339320|issn=1079-5006|doi-access=free}}</ref>

==Species==
{{cladogram|align=right
|caption=Relationships among ''Myotis'' species according to molecular data<ref name=phylogenetics>Stadelmann et al., 2007, fig. 2; Lack et al., 2010, figs. 1, 2</ref>
|clades=
{{clade
|label1='''Myotinae'''
|1={{clade
   |1=''[Submyotodon](/source/Submyotodon)''
   |label2=''[Myotis](/source/Myotis)''
   |2={{clade
      |1=Most Old World species
      |2={{clade
         |1=Most [Nearctic](/source/Nearctic) species
         |2={{clade
            |1=''[Myotis brandtii](/source/Myotis_brandtii)'' & ''[Myotis sibiricus](/source/Myotis_sibiricus)''
            |2=[Neotropical](/source/Neotropical) and some Nearctic species
            }}
         }}
      }}
   }}
}}
}}
Traditionally, ''Myotis'' was divided into three large subgenera—''Leuconoe'', ''Myotis'', and ''Selysius''. However, molecular data indicate that these subgenera are not natural groups, but instead unnatural assemblages of [convergently](/source/convergent_evolution) similar species.<ref name=S500>Simmons, 2005, p. 500</ref> Instead, ''Myotis'' species largely fall in two main [clade](/source/clade)s, one containing Old World and the other New World species.<ref name=phylogenetics/> The [ITIS](/source/Integrated_Taxonomic_Information_System) presently divides it into three subgenera: '''''Chrysopteron''''' (containing most reddish-colored Old World species), '''''Myotis''''' (containing almost all other Old World species), and '''''Pizonyx''''' (containing all New World species and the Eurasian ''[Myotis brandtii](/source/Myotis_brandtii)'' and ''[Myotis sibiricus](/source/Siberian_bat)'', which are more closely related to New World species than to other Old World species).<ref>{{Cite web|title=ITIS - Report: Myotis|url=https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=179987#null|access-date=2022-02-01|website=www.itis.gov}}</ref><ref>Stadelmann et al., 2007, fig. 2</ref> The Asian species ''[Myotis latirostris](/source/Myotis_latirostris)'' falls outside the clade formed by these main groups, and has since been reclassified into a separate genus, ''[Submyotodon](/source/Submyotodon)'', alongside several others.<ref>Lack et al., 2010, p. 984</ref>

The earliest known fossil ''Myotis'' is the extinct, large-sized species †''[Myotis belgicus](/source/Myotis_belgicus)'' <small>Gunnel, Smith & Smith, 2017</small> from the [Early Oligocene](/source/Early_Oligocene) of Belgium.<ref name="Gunnel2017">{{Cite journal |last1=Gunnell |first1=Gregg F. |last2=Smith |first2=Richard |last3=Smith |first3=Thierry |date=2017-03-08 |title=33 million year old Myotis (Chiroptera, Vespertilionidae) and the rapid global radiation of modern bats |journal=PLOS ONE |language=en |volume=12 |issue=3 |article-number=e0172621 |doi=10.1371/journal.pone.0172621 |doi-access=free |issn=1932-6203 |pmc=5342209 |pmid=28273112 |bibcode=2017PLoSO..1272621G }}</ref>
[[File:Myotis emarginatus - Ph. Karol Tabarelli de Fatis.jpg|thumb|[Geoffroy's bat](/source/Geoffroy's_bat)]]
[[File:Black-winged Myotis imported from iNaturalist photo 6268974 on 26 January 2022 (cropped).jpg|thumb|[Black-winged myotis](/source/Reddish-black_myotis)]]
[[File:Vespertilion bechstein.jpg|thumb|[Bechstein's bat](/source/Bechstein's_bat)]]
[[File:Myotis crypticus - Manuel Ruedi.jpg|thumb|[Cryptic myotis](/source/Cryptic_myotis)]]
[[File:Myotis myotis (cropped).jpg|thumb|[Greater mouse-eared bat](/source/Greater_mouse-eared_bat)]]
[[File:Myotis yumanensis (Yuma myotis) (11362476624).jpg|thumb|[Yuma myotis](/source/Yuma_myotis)]]
[[File:Myotis daubentoni.jpg|thumb|[Daubenton's bat](/source/Daubenton's_bat)]]
[[File:Southeastern Myotis profile.JPG|thumb|[Southeastern myotis](/source/Southeastern_myotis)]]
[[File:Myotis brandtii - Manuel Ruedi.JPG|thumb|[Brandt's bat](/source/Brandt's_bat)]]
[[File:Myotis vivesi.jpg|thumb|[Fish-eating bat](/source/Myotis_vivesi)]]
''Myotis'' is a highly species-rich genus, and the classification of many species remains unsettled. The taxonomy below is based on that of the [ITIS](/source/Integrated_Taxonomic_Information_System) in 2021.<ref>{{Citation|last=Mammal Diversity Database|title=Mammal Diversity Database|work=Zenodo |date=2021-08-10|doi=10.5281/zenodo.5175993 |url=https://zenodo.org/record/5175993|access-date=2021-09-11}}</ref> Some differences in taxonomy from the 2005 third edition of ''[Mammal Species of the World](/source/Mammal_Species_of_the_World)''<ref>Simmons, 2005, pp. 500–518</ref> are indicated in footnotes.

{{div col|colwidth=30em}}
* Subgenus ''[Chrysopteron](/source/Chrysopteron)'':
** ''[Myotis anjouanensis](/source/Myotis_anjouanensis)'' <small>(Dorst, 1960)</small> - Anjouan myotis
** ''[Myotis bartelsii](/source/Myotis_bartelsii)'' <small>(Jentink, 1910)</small> - Bartels's myotis
** ''[Myotis bocagii](/source/Myotis_bocagii)'' <small>(Peters, 1870)</small> - rufous mouse-eared bat
** ''[Myotis dieteri](/source/Myotis_dieteri)'' <small>(Happold, 2005)</small> - Kock's mouse-eared bat
** ''[Myotis emarginatus](/source/Myotis_emarginatus)'' <small>(E. Geoffroy, 1806)</small> - Geoffroy's bat
** ''[Myotis formosus](/source/Myotis_formosus)'' <small>(Hodgson, 1835)</small> - Hodgson's bat, copper-winged bat
** ''[Myotis goudotii](/source/Myotis_goudotii)'' <small>(A. Smith, 1834)</small> - Malagasy mouse-eared bat
** ''[Myotis hermani](/source/Myotis_hermani)'' <small>Thomas, 1923</small> - Herman's myotis
** ''[Myotis morrisi](/source/Myotis_morrisi)'' <small>Hill, 1971</small> - Morris's bat
** ''[Myotis nimbaensis](/source/Myotis_nimbaensis)'' <small>(Simmons et al., 2021)</small> - Nimba mountain bat
** ''[Myotis rufoniger](/source/Myotis_rufoniger)'' <small>(Tomes, 1858)</small>{{#tag:ref|Split from ''M.&nbsp;formosus'' (Csorba et al., 2014).|group=footnote}} - reddish-black myotis
** ''[Myotis rufopictus](/source/Myotis_rufopictus)'' <small>(Waterhouse, 1845)</small>{{#tag:ref|Split from ''M.&nbsp;formosus'' (Csorba et al., 2014).|group=footnote}} - orange-fingered myotis
** ''[Myotis scotti](/source/Myotis_scotti)'' <small>Thomas, 1927</small> - Scott's mouse-eared bat
** ''[Myotis tricolor](/source/Myotis_tricolor)'' <small>(Temminck, 1832)</small> - Cape hairy bat, little brown bat, Temminck's mouse-eared bat, Cape myotis, tricoloured mouse-eared bat, Cape hairy myotis, Temminck's hairy bat, three-coloured bat
** ''[Myotis weberi](/source/Myotis_weberi)'' <small>(Jentink, 1890)</small>{{#tag:ref|Split from ''M.&nbsp;formosus'' (Csorba et al., 2014).|group=footnote}} - Weber's myotis
** ''[Myotis welwitschii](/source/Myotis_welwitschii)'' <small>(Gray, 1866)</small> - Welwitsch's bat, Welwitsch's mouse-eared bat, Welwitsch's myotis
* Subgenus ''Myotis'':
** ''[Myotis adversus](/source/Myotis_adversus)'' <small>(Horsfield, 1824)</small> - large-footed bat, large-footed mouse-eared bat, large-footed myotis
** ''[Myotis aelleni](/source/Myotis_aelleni)'' <small>(Baud, 1979)</small> - southern myotis (disputed species)
** ''[Myotis alcathoe](/source/Myotis_alcathoe)'' <small>(von Helversen and  Heller, 2001)</small> - Alcathoe bat
** ''[Myotis altarium](/source/Myotis_altarium)'' <small>(Thomas, 1911)</small> - Szechwan myotis
** ''[Myotis alticraniatus](/source/Myotis_alticraniatus)'' <small>Osgood, 1932</small> - Indochinese whiskered myotis
** ''[Myotis ancricola](/source/Myotis_ancricola)'' <small>Kruskop, Borisenko, Dudorova, & Artyushin, 2018</small> - valley myotis
** ''[Myotis annamiticus](/source/Myotis_annamiticus)'' <small>(Kruskop and Tsytsulina, 2001)</small> - Annamit myotis
** ''[Myotis annatessae](/source/Myotis_annatessae)'' <small>Kruskop & Borisenko, 2013</small> - Anna Tess's myotis
** ''[Myotis annectans](/source/Myotis_annectans)'' <small>(Dobson, 1871)</small> - hairy-faced bat
** ''[Myotis ater](/source/Myotis_ater)'' <small>(Peters, 1866)</small> - Peters's myotis, small black myotis
** ''[Myotis badius](/source/Myotis_badius)'' <small>Tiunov, Kruskop, & Feng Jiang, 2011</small> - chestnut myotis
** ''[Myotis bechsteinii](/source/Myotis_bechsteinii)'' <small>(Kuhl, 1817)</small> - Bechstein's bat
** ''[Myotis blythii](/source/Myotis_blythii)'' <small>(Tomes, 1857)</small> - lesser mouse-eared bat
** ''[Myotis bombinus](/source/Myotis_bombinus)'' <small>(Thomas, 1906)</small> - Far Eastern myotis, bombinus bat
** ''[Myotis borneoensis](/source/Myotis_borneoensis)'' <small>Hill & Francis, 1984</small>{{#tag:ref|Split from ''M.&nbsp;montivagus'' (Görföl et al., 2013).|group=footnote}} - Bornean whiskered myotis
** ''[Myotis browni](/source/Myotis_browni)'' <small>E. H. Taylor, 1934</small> - Brown's whiskered myotis
** ''[Myotis bucharensis](/source/Myotis_bucharensis)'' <small>(Kuzyakin, 1950)</small> - Bocharic myotis, Bokhara whiskered bat
** ''[Myotis capaccinii](/source/Myotis_capaccinii)'' <small>(Bonaparte, 1837)</small> - long-fingered bat
** ''[Myotis chinensis](/source/Myotis_chinensis)'' <small>(Tomes, 1857)</small> - large myotis
** ''[Myotis crypticus](/source/Myotis_crypticus)'' <small>Ruedi, Ibáñez, Salicini, Juste & Puechmaille, 2019</small> - cryptic myotis
** ''[Myotis csorbai](/source/Myotis_csorbai)'' <small>(Topál, 1997)</small> - Csorba's mouse-eared bat
** ''[Myotis dasycneme](/source/Myotis_dasycneme)'' <small>(Boie, 1825)</small> - pond bat
** ''[Myotis daubentonii](/source/Myotis_daubentonii)'' <small>(Kuhl, 1817)</small> - Daubenton's bat
** ''[Myotis davidii](/source/Myotis_davidii)'' <small>(Peters, 1869)</small> - David's myotis
** ''[Myotis escalerai](/source/Myotis_escalerai)'' <small>Cabrera, 1904</small>{{#tag:ref|Split from ''M.&nbsp;nattereri'' (Ibáñez et al., 2006).|group=footnote}} - Escalera's bat
** ''[Myotis federatus](/source/Myotis_federatus)'' <small>Thomas, 1916</small>{{#tag:ref|Split from ''M.&nbsp;montivagus'' (Görföl et al., 2013).|group=footnote}} - Malaysian whiskered myotis
** ''[Myotis fimbriatus](/source/Myotis_fimbriatus)'' <small>(Peters, 1871)</small> - fringed long-footed myotis
** ''[Myotis frater](/source/Myotis_frater)'' <small>G.M.&nbsp;Allen, 1923</small> - fraternal myotis
** ''[Myotis gomantongensis](/source/Myotis_gomantongensis)'' <small>Francis and  Hill, 1998</small> - Gomantong myotis
** ''[Myotis hajastanicus](/source/Myotis_hajastanicus)'' <small>Argyropulo, 1939</small> - Armenian whiskered bat, Hajastan myotis, Armenian myotis (disputed species)
** ''[Myotis hasseltii](/source/Myotis_hasseltii)'' <small>(Temminck, 1840)</small> - lesser large-footed bat
** ''[Myotis horsfieldii](/source/Myotis_horsfieldii)'' <small>(Temminck, 1840)</small> - Horsfield's bat
** ''[Myotis hoveli](/source/Myotis_hoveli)'' <small>Harrison, 1964</small> - Hovel's myotis
** ''[Myotis hyrcanicus](/source/Myotis_hyrcanicus)'' <small>Benda ''et al.'', 2012</small> - Hyrcanian myotis
** ''[Myotis ikonnikovi](/source/Myotis_ikonnikovi)'' <small>Ognev, 1912</small> - Ikonnikov's bat
** ''[Myotis indochinensis](/source/Myotis_indochinensis)'' <small>Son ''et al.'', 2013</small> - Indochinese myotis
** ''[Myotis insularum](/source/Myotis_insularum)'' <small>(Dobson, 1878)</small> - insular myotis
** ''[Myotis laniger](/source/Myotis_laniger)'' <small>Peters, 1871</small> - Chinese water myotis
** ''[Myotis longicaudatus](/source/Myotis_longicaudatus)'' <small>Ognev, 1927</small>{{#tag:ref|Split from ''M.&nbsp;frater'' (Ruedi et al., 2015).|group=footnote}} - long-tailed myotis
** ''[Myotis longipes](/source/Myotis_longipes)'' <small>(Dobson, 1873)</small> - Kashmir cave bat
** ''[Myotis macrodactylus](/source/Myotis_macrodactylus)'' <small>(Temminck, 1840)</small> - eastern long-fingered bat, big-footed myotis
** ''[Myotis macropus](/source/Myotis_macropus)'' <small>(Gould, 1854)</small> - southern myotis, large-footed myotis
** ''[Myotis macrotarsus](/source/Myotis_macrotarsus)'' <small>(Waterhouse, 1845)</small> - pallid large-footed myotis, Philippine large-footed myotis
** ''[Myotis melanorhinus](/source/Myotis_melanorhinus)'' <small>Merriam, 1890</small> - dark-nosed small-footed myotis (disputed species)
** ''[Myotis moluccarum](/source/Myotis_moluccarum)'' <small>(Thomas, 1915)</small> - Maluku myotis, Arafura large-footed bat
** ''[Myotis montivagus](/source/Myotis_montivagus)'' <small>(Dobson, 1874)</small> - Burmese whiskered bat
** ''[Myotis muricola](/source/Myotis_muricola)'' <small>(Gray, 1846)</small> - wall-roosting mouse-eared bat, Nepalese whiskered myotis
** ''[Myotis myotis](/source/Myotis_myotis)'' <small>(Borkhausen, 1797)</small> - greater mouse-eared bat
** ''[Myotis mystacinus](/source/Myotis_mystacinus)'' <small>(Kuhl, 1817)</small> - whiskered bat
** ''[Myotis nattereri](/source/Myotis_nattereri)'' <small>(Kuhl, 1817)</small> - Natterer's bat
** ''[Myotis nipalensis](/source/Myotis_nipalensis)'' <small>Dobson, 1871</small> - Nepal myotis
** ''[Myotis pequinius](/source/Myotis_pequinius)'' <small>Thomas, 1908</small> - Beijing mouse-eared bat, Peking myotis
** ''[Myotis petax](/source/Myotis_petax)'' <small>Hollister, 1912</small>{{#tag:ref|Split from ''M.&nbsp;daubentonii'' (Matveev et al., 2005). Includes ''M.&nbsp;abei'' <small>(Tsytsulina, 2004, as ''daubentonii'')</small>.|group=footnote}} - eastern water bat, Sakhalin bat
** ''[Myotis peytoni](/source/Myotis_peytoni)'' <small>Wroughton & Ryley, 1913</small>{{#tag:ref|Split from ''M.&nbsp;montivagus'' (Görföl et al., 2013).|group=footnote}} - Peyton's myotis
** ''[Myotis phanluongi](/source/Myotis_phanluongi)'' <small>Borisenko, Kruskop and  Ivanova, 2008</small> - Phan Luong's myotis{{#tag:ref|A new species (Borisenko et al., 2008).|group=footnote}}
** ''[Myotis pilosus](/source/Myotis_pilosus)'' <small>Peters, 1869</small> - Rickett's big-footed bat
** ''[Myotis pruinosus](/source/Myotis_pruinosus)'' <small>Yoshiyuki, 1971</small> - frosted myotis
** ''[Myotis punicus](/source/Myotis_punicus)'' <small>Felten, Spitzenberger and Storch, 1977</small> - Felten's myotis
** ''[Myotis ridleyi](/source/Myotis_ridleyi)'' <small>Thomas, 1898</small> - Ridley's bat
** ''[Myotis rosseti](/source/Myotis_rosseti)'' <small>(Oey, 1951)</small> - thick-thumbed myotis
** ''[Myotis schaubi](/source/Myotis_schaubi)'' <small>Kormos, 1934</small> - Schaub's myotis
** ''[Myotis secundus](/source/Myotis_secundus)'' <small>Ruedi, Csorba, Lin, & Chou, 2015</small> - long-toed myotis
** ''[Myotis sicarius](/source/Myotis_sicarius)'' <small>Thomas, 1915</small> - Mandelli's mouse-eared bat
** ''[Myotis siligorensis](/source/Myotis_siligorensis)'' <small>(Horsfield, 1855)</small> - Himalayan whiskered bat
** ''[Myotis soror](/source/Myotis_soror)'' <small>Ruedi, Csorba, Lin, & Chou, 2015</small> - reddish myotis
** ''[Myotis sowerbyi](/source/Myotis_sowerbyi)'' <small>Howell, 1926</small> - Sowerby's whiskered myotis
** ''[Myotis stalkeri](/source/Myotis_stalkeri)'' <small>Thomas, 1910</small> - Kei myotis
** ''[Myotis tschuliensis](/source/Myotis_tschuliensis)'' <small>Kuzyakin, 1935</small> - Tschuli myotis
** ''[Myotis yanbarensis](/source/Myotis_yanbarensis)'' <small>Maeda and  Matsumara, 1998</small> - Yanbaru whiskered bat
** ''[Myotis zenatius](/source/Myotis_zenatius)''  <small>Ibáñez, Juste, Salicini, Puechmaille & Ruedi, 2019</small> - Zenati myotis
* Subgenus ''[Pizonyx](/source/Pizonyx)'':
** ''[Myotis albescens](/source/Myotis_albescens)'' <small>(E. Geoffroy, 1806)</small> - silver-tipped myotis
** ''[Myotis armiensis](/source/Myotis_armiensis)'' <small>Carrión-Bonilla & Cook, 2020</small> - Armién's myotis
** ''[Myotis atacamensis](/source/Myotis_atacamensis)'' <small>(Lataste, 1892)</small> - Atacama myotis
** ''[Myotis attenboroughi](/source/Myotis_attenboroughi)'' <small>Moratelli et al., 2017</small>{{#tag:ref|Split from ''M.&nbsp;nigricans'' (Moratelli et al., 2017).|group=footnote}} - Sir David Attenborough's myotis
** ''[Myotis auriculus](/source/Myotis_auriculus)'' <small>(Baker and  Stains, 1955)</small> - southwestern myotis
** ''[Myotis austroriparius](/source/Myotis_austroriparius)'' <small>(Rhoads, 1897)</small> - southeastern myotis
** ''[Myotis bakeri](/source/Myotis_bakeri)'' <small>Moratelli, Novaes, Bonilla, & D. E. Wilson, 2019</small> - Baker's myotis
** ''[Myotis brandtii](/source/Myotis_brandtii)'' <small>(Eversmann, 1845)</small> - Brandt's bat
** ''[Myotis californicus](/source/Myotis_californicus)'' <small>(Audubon and  Bachman, 1842)</small> - California myotis
** ''[Myotis caucensis](/source/Myotis_caucensis)'' <small>Allen, 1914</small> - Colombian black myotis
** ''[Myotis chiloensis](/source/Myotis_chiloensis)'' <small>(Waterhouse, 1840)</small> - Chilean myotis
** ''[Myotis ciliolabrum](/source/Myotis_ciliolabrum)'' <small>(Merriam, 1886)</small> - western small-footed bat, western small-footed myotis
** ''[Myotis clydejonesi](/source/Myotis_clydejonesi)'' <small>Moratelli, D. E. Wilson, A. L. Gardner, Fisher, & Gutierrez, 2016</small> - Clyde Jones's myotis
** ''[Myotis cobanensis](/source/Myotis_cobanensis)'' <small>(Goodwin, 1955)</small> - Guatemalan myotis
** ''[Myotis diminutus](/source/Myotis_diminutus)'' <small>Moratelli & Wilson, 2011</small> - diminutive myotis
** ''[Myotis dinellii](/source/Myotis_dinellii)'' <small>Thomas, 1902</small>{{#tag:ref|Split from ''M.&nbsp;levis'' (Barquez et al., 2006).|group=footnote}} - Dinelli's myotis
** ''[Myotis dominicensis](/source/Myotis_dominicensis)'' <small>Miller, 1902</small> - Dominican myotis
** ''[Myotis elegans](/source/Myotis_elegans)'' <small>Hall, 1962</small> - elegant myotis
** ''[Myotis evotis](/source/Myotis_evotis)'' <small>(H. Allen, 1864)</small> - long-eared myotis
** ''[Myotis findleyi](/source/Myotis_findleyi)'' <small>Bogan, 1978</small> - Findley's myotis
** ''[Myotis fortidens](/source/Myotis_fortidens)'' <small>Miller and  Allen, 1928</small> - cinnamon myotis
** ''[Myotis grisescens](/source/Myotis_grisescens)'' <small>A.H. Howell, 1909</small> - gray bat
** ''[Myotis handleyi](/source/Myotis_handleyi)'' <small>Moratelli, A. L. Gardner, J. A. Oliveira, & D. E. Wilson, 2013</small> - Handley's myotis
** ''[Myotis izecksohni](/source/Myotis_izecksohni)'' <small>Moratelli, Peracchi, Dias & de Oliveira, 2011</small> - Izecksohn's myotis<ref name="Moratelli">{{cite journal|doi=10.1016/j.mambio.2011.01.003|title=Geographic variation in South American populations of Myotis nigricans ( ) (Chiroptera, Vespertilionidae), with the description of two new species|journal=Mammalian Biology|volume=76|issue=5|pages=592–607|year=2011|last1=Moratelli|first1=Ricardo|last2=Peracchi|first2=Adriano L.|last3=Dias|first3=Daniela|last4=De Oliveira|first4=João A.}}</ref>
** ''[Myotis keaysi](/source/Myotis_keaysi)'' <small>J.A. Allen, 1914</small> - hairy-legged myotis
** ''[Myotis keenii](/source/Myotis_keenii)'' <small>(Merriam, 1895)</small> -  Keen's myotis
** ''[Myotis larensis](/source/Myotis_larensis)'' <small>LaVal, 1973</small> - Lara myotis
** ''[Myotis lavali](/source/Myotis_lavali)'' <small>Moratelli, Peracchi, Dias, & Oliveira, 2011</small> - LaVal's Myotis
** ''[Myotis leibii](/source/Myotis_leibii)'' <small>(Audubon and  Bachman, 1842)</small> - eastern small-footed bat
** ''[Myotis levis](/source/Myotis_levis)'' <small>(I. Geoffroy, 1824)</small> - yellowish myotis
** ''[Myotis lucifugus](/source/Myotis_lucifugus)'' <small>(Le Conte, 1831)</small> - little brown bat, little brown myotis
** ''[Myotis martiniquensis](/source/Myotis_martiniquensis)'' <small>LaVal, 1973</small> - Schwartz's myotis
** ''[Myotis midastactus](/source/Myotis_midastactus)'' <small>Moratelli & Wilson, 2014</small> - golden myotis{{#tag:ref|Split from ''M.&nbsp;simus'' (Moratelli & Wilson, 2014).|group=footnote}}
** ''[Myotis nesopolus](/source/Myotis_nesopolus)'' <small>Miller, 1900</small> - Curacao myotis
** ''[Myotis nigricans](/source/Myotis_nigricans)'' <small>(Schinz, 1821)</small> - black myotis
** ''[Myotis nyctor](/source/Myotis_nyctor)'' <small>LaVal & Schwartz, 1974</small> - Barbados myotis{{#tag:ref|Split from ''M.&nbsp;martiniquensis'' (Larsen et al., 2012).|group=footnote}}
** ''[Myotis occultus](/source/Myotis_occultus)'' <small>Hollister, 1909</small> - Arizona myotis
** ''[Myotis oxyotus](/source/Myotis_oxyotus)'' <small>(Peters, 1867)</small> - montane myotis
** ''[Myotis peninsularis](/source/Myotis_peninsularis)'' <small>Miller, 1898</small> - peninsular myotis
** ''[Myotis pilosatibialis](/source/Myotis_pilosatibialis)'' <small>LaVal, 1973</small> - northern hairy-legged myotis
** ''[Myotis planiceps](/source/Myotis_planiceps)'' <small>Baker, 1955</small> - flat-headed myotis
** ''[Myotis riparius](/source/Myotis_riparius)'' <small>Handley, 1960</small> - riparian myotis
** ''[Myotis ruber](/source/Myotis_ruber)'' <small>(E. Geoffroy, 1806)</small> - red myotis
** ''[Myotis septentrionalis](/source/Myotis_septentrionalis)'' <small>(Trouessart, 1897)</small> - northern long-eared bat, northern myotis
** ''[Myotis sibiricus](/source/Myotis_sibiricus)'' <small>(Kastschenko, 1905)</small> - Siberian bat or Siberian whiskered myotis{{#tag:ref|Split from ''M.&nbsp;brandtii'' (Kruskop, Borisenko, Ivanova, Lim & Eger, 2012).|group=footnote}}
** ''[Myotis simus](/source/Myotis_simus)'' <small>Thomas, 1901</small> - velvety myotis
** ''[Myotis sodalis](/source/Myotis_sodalis)'' <small>Miller and  Allen, 1928</small> - Indiana bat
** ''[Myotis thysanodes](/source/Myotis_thysanodes)'' <small>Miller, 1897</small> - fringed myotis
** ''[Myotis velifer](/source/Myotis_velifer)'' <small>(J.A. Allen, 1890)</small> - cave myotis
** ''[Myotis vivesi](/source/Myotis_vivesi)'' <small>Menegaux, 1901</small> - fish-eating bat, fish-eating myotis
** ''[Myotis volans](/source/Myotis_volans)'' <small>(H. Allen, 1866)</small> - long-legged myotis
** ''[Myotis yumanensis](/source/Myotis_yumanensis)'' <small>(H. Allen, 1864)</small> - Yuma myotis
* Unclassified & dubious species:
** ''[Myotis australis](/source/Myotis_australis)'' <small>(Dobson, 1878)</small> - Australian myotis (disputed species)
** ''[Myotis oreias](/source/Myotis_oreias)'' <small>(Temminck, 1840)</small> - Singapore whiskered bat
{{div col end}}

==See also==
* [Bat adenovirus TJM](/source/Bat_adenovirus_TJM)

== Notes ==
{{Reflist|group=footnote}}

==References==
{{Reflist|30em}}

==Literature cited==
* Borisenko, A.V., Kruskop, S.V. and Ivanova, N.V. 2008. A new mouse-eared bat (Mammalia: Chiroptera: Vespertilionidae) from Vietnam. Russian Journal of Theriology 7(2):57–69.
* Han, N., Zhang, J., Reardon, T., Lin, L., Zhang, J. and Zhang, S. 2010. [http://www.bioone.org/doi/full/10.3161/150811010X538016 Revalidation of ''Myotis taiwanensis'' Ärnbäck-Christie-Linde 1908 and its molecular relationship with ''M. adversus'' (Horsfield 1824) (Vespertilionidae, Chiroptera)] (subscription required). Acta Chiropterologica 12(2):449–456.*Happold, M. 2005. A new species of ''Myotis'' (Chiroptera: Vespertilionidae) from central Africa. Acta Chiropterologica 7(1):9–21.
* Ibáñez, C., García-Mudarra, J.L., Ruedi, M., Stadelmann, B. and Juste, J. 2006. The Iberian contribution to cryptic diversity in European bats. Acta Chiropterologica 8(2):277–297.
* Jiang, T., Sun, K., Chou, C., Zhang, Z. and Feng, J. 2010. First record of ''Myotis flavus'' (Chiroptera: Vespertilionidae) from mainland China and a reassessment of its taxonomic status. Zootaxa 2414:41–51.
* Lack, J.B., Roehrs, Z.P., Stanley, C.E., Ruedi, M. and Van Den Bussche, R.A. 2010. [https://dx.doi.org/10.1644/09-MAMM-A-192.1 Molecular phylogenetics of ''Myotis'' indicate familial-level divergence for the genus ''Cistugo'' (Chiroptera)] (subscription required). Journal of Mammalogy 91(4):976–992.
* Matveev, V.A., Kruskop, S.V. and Kramerov, D.A. 2005. Revalidation of ''Myotis petax'' Hollister, 1912 and its new status in connection with ''M. daubentonii'' (Kuhl, 1817) (Vespertilionidae, Chiroptera). Acta Chiropterologica 7(1):23–37.
* Mayer, F., Dietz, C. and Kiefer, A. 2007. [https://dx.doi.org/10.1186/1742-9994-4-4 Molecular species identification boosts bat diversity]. Frontiers in Zoology 4(1):239–255.
* {{cite journal|last1= Moratelli|first1=R.|last2=Wilson|first2=D. E.|last3=Novaes|first3=R. L. M.|last4=Helgen|first4=K. M.|last5=Gutiérrez|first5=E. E.|title= Caribbean ''Myotis'' (Chiroptera, Vespertilionidae), with description of a new species from Trinidad and Tobago|journal= Journal of Mammalogy|volume=98|issue=4|pages=994–1008|date= 2017-06-07|doi= 10.1093/jmammal/gyx062|doi-access=free}}
* Roehrs, Z.P., Lack, J.B. and Van Den Bussche, R.A. 2010. [https://dx.doi.org/10.1644/09-MAMM-A-325.1 Tribal phylogenetic relationships within Vespertilioninae (Chiroptera: Vespertilionidae) based on mitochondrial and nuclear sequence data] (subscription required). Journal of Mammalogy 91(5):1073–1092.
* Simmons, N.B. 2005. Order Chiroptera. Pp.&nbsp;312–529 in Wilson, D.E. and Reeder, D.M. (eds.). [http://www.bucknell.edu/msw3 Mammal Species of the World: A Taxonomic and Geographic Reference]. 3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142&nbsp;pp.&nbsp;{{ISBN|978-0-8018-8221-0}}
* Stadelmann, B., Lin, L.-K., Kunz, T.H. and Ruedi, M. 2007. [https://dx.doi.org/10.1016/j.ympev.2006.06.019 Molecular phylogeny of New World ''Myotis'' (Chiroptera, Vespertilionidae) inferred from mitochondrial and nuclear DNA genes] (subscription required). Molecular Phylogenetics and Evolution 43(1):32–48.
* Tsytsulina, K. 2004. On the taxonomical status of ''Myotis abei'' Yoshikura, 1944 (Chiroptera, Vespertilionidae). Zoological Science 21:963–966.
* Simmons, Nancy B.; Flanders, J.; Bakwo Fils, E. M.; Parker, Guy; Suter, Jamison D.; Bamba, Seinan; Keita, Mamady Kobele; Morales, Ariadna E.; Frick, Winifred F. 2021. [http://digitallibrary.amnh.org/handle/2246/7249 A new dichromatic species of Myotis (Chiroptera: Vespertilionidae) from the Nimba Mountains, Guinea (American Museum novitates, no. 3963)] American Museum Novitates. ISSN 0003-0082.

==External links==
{{Wikispecies|''Myotis''}}
{{Commons category|Myotis}}
* [http://animaldiversity.ummz.umich.edu/accounts/Myotis/classification/ ''Myotis''] at [Animal Diversity Web](/source/Animal_Diversity_Web)

{{Vespertilionidae}}
{{Myotinae nav}}

{{Taxonbar|from=Q223384}}

{{DEFAULTSORT:Mouse-Eared Bats}}
Category:Mouse-eared bats
Category:Extant Rupelian first appearances

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Adapted from the Wikipedia article [Mouse-eared bat](https://en.wikipedia.org/wiki/Mouse-eared_bat) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Mouse-eared_bat?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
