# Melolonthini

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{{short description|Tribe of beetles}}
{{Automatic taxobox
| image = Phyllophaga P1380631a.jpg
| image_caption = ''Phyllophaga'' sp.
| taxon = Melolonthini
| authority = Leach, 1819
| subdivision_ranks = Subtribes
| subdivision =
: Enariina <small>Dewailly, 1950</small>
: Heptophyllina <small>Medvedev, 1951</small>
: Leucopholina <small>Burmeister, 1855</small>
: Melolonthina <small>Leach, 1819</small>
: Pegylina <small>Lacroix, 1989</small>
: Rhizotrogina <small>Burmeister, 1855</small>
: Schizonychina <small>Burmeister, 1855</small>
}}

thumb|''Polyphylla occidentalis''
'''Melolonthini''' is a tribe of scarab beetles in the family [Scarabaeidae](/source/Scarabaeidae).<ref>{{Cite web|url=https://bugguide.net/node/view/2951|title=Subfamily Melolonthinae - May Beetles and Junebugs - BugGuide.Net|website=bugguide.net|access-date=2020-04-22}}</ref> There are over 250 genera in Melolonthini, occurring worldwide; there are over 300 species in North America alone, and more than 3000 worldwide.<ref name=bugref>{{cite web|url=https://bugguide.net/node/view/41314|title=Melolonthini Tribe Information|website=BugGuide.net|publisher=Iowa State University|accessdate=2018-01-25}}</ref><ref name=itis>{{cite web|url=https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=926272|title=Melolonthini Report|website=Integrated Taxonomic Information System|accessdate=2018-01-25}}</ref><ref name=Jia2021/>

==Subtribes and genera==
The tribe is divided in three subtribes:
*subtribe Enariina <small>Dewailly, 1950</small>
**''[Anenaria](/source/Anenaria)'' <small>Lacroix, 1993</small>
**''[Anthrencya](/source/Anthrencya)'' <small>Lacroix, 1993</small>
**''[Apicencya](/source/Apicencya)'' <small>Lacroix, 1993</small>
**''[Bisencya](/source/Bisencya)'' <small>Lacroix, 1993</small>
**''[Cherbezatina](/source/Cherbezatina)'' <small>Lacroix, 1993</small>
**''[Comencya](/source/Comencya)'' <small>Lacroix, 1993</small>
**''[Djadjoua](/source/Djadjoua)'' <small>Lacroix, 1993</small>
**''[Djafouna](/source/Djafouna)'' <small>Lacroix, 1993</small>
**''[Enaria](/source/Enaria)'' <small>Erichson, 1847</small>
**''[Encya](/source/Encya)'' <small>Dejean, 1833</small>
**''[Enthora](/source/Enthora)'' <small>Erichson, 1847</small>
**''[Eutrichesis](/source/Eutrichesis)'' <small>Waterhouse, 1882</small>
**''[Globencya](/source/Globencya)'' <small>Lacroix, 1993</small>
**''[Humblotania](/source/Humblotania)'' <small>Lacroix, 1993</small>
**''[Joziratia](/source/Joziratia)'' <small>Lacroix, 1993</small>
**''[Komrina](/source/Komrina)'' <small>Lacroix, 1993</small>
**''[Lebbea](/source/Lebbea)'' <small>Lacroix, 1993</small>
**''[Machala](/source/Machala_(beetle))'' <small>Lacroix, 1993</small>
**''[Mayataia](/source/Mayataia)'' <small>Lacroix, 1993</small>
**''[Mucrencya](/source/Mucrencya)'' <small>Lacroix, 1993</small>
**''[Pentaphylla](/source/Pentaphylla_(beetle))'' <small>Dewailly, 1950</small>
**''[Polyenaria](/source/Polyenaria)'' <small>Dewailly, 1950</small>
**''[Pseudenaria](/source/Pseudenaria)'' <small>Fairmaire, 1901</small>
**''[Pseudencya](/source/Pseudencya)'' <small>Lacroix, 1991</small>
**''[Ravautiana](/source/Ravautiana)'' <small>Lacroix, 1991</small>
**''[Renaudiana](/source/Renaudiana)'' <small>Lacroix, 1993</small>
**''[Semienaria](/source/Semienaria)'' <small>Dewailly, 1950</small>
**''[Synenaria](/source/Synenaria)'' <small>Lacroix, 1993</small>
**''[Vadonaria](/source/Vadonaria)'' <small>Dewailly, 1950</small>
**''[Varencya](/source/Varencya)'' <small>Lacroix, 1993</small>
*subtribe Melolonthina <small>Leach, 1819</small>
**''[Melolontha](/source/Melolontha)'' <small>Fabricius, 1775</small>  (cockchafers)
**''[Polyphylla](/source/Polyphylla)'' <small>Harris, 1841</small> (lined June beetles)
**''[Psilopholis](/source/Psilopholis)'' <small>Brenske, 1892</small>
*subtribe Pegylina <small>Lacroix, 1989</small>
**''[Eupegylis](/source/Eupegylis)'' <small>Duvivier, 1892</small>
**''[Neopegylis](/source/Neopegylis)'' <small>Lacroix, 2015</small>
**''[Pegylis](/source/Pegylis)'' <small>Erichson, 1847</small>
*Furthermore, the following genera have not been assigned to a subtribe:
**''[Achranoxia](/source/Achranoxia)'' <small>Kraatz, 1888</small>
**''[Adoretops](/source/Adoretops)'' <small>Kraatz, 1883</small>
**''[Alepida](/source/Alepida)'' <small>Allsopp, 2018</small>
**''[Allophyllus](/source/Allophyllus)'' <small>Fauvel, 1903</small>
**''[Allothnonius](/source/Allothnonius)'' <small>Britton, 1978</small>
**''[Amblonoxia](/source/Amblonoxia)'' <small>Reitter, 1902</small>  (dusty June beetles)
**''[Anisopholis](/source/Anisopholis)'' <small>Moser, 1913</small>
**''[Anoxia](/source/Anoxia_(beetle))'' <small>Laporte, 1832</small>
**''[Anoxoides](/source/Anoxoides)'' <small>Petrovitz, 1971</small>
**''[Antitrogus](/source/Antitrogus)'' <small>Burmeister, 1855</small>
**''[Barryfilius](/source/Barryfilius)'' <small>Allsopp, 2022</small>
**''[Beriqua](/source/Beriqua)'' <small>Péringuey, 1904</small>
**''[Brenskecnemus](/source/Brenskecnemus)'' <small>Allsopp & Schoolmeesters, 2024</small>
**''[Carlschoenherria](/source/Carlschoenherria)'' <small>Bezděk, 2016</small>
**''[Cephaloschiza](/source/Cephaloschiza)'' <small>Moser, 1920</small>
**''[Cheronia](/source/Cheronia_(beetle))'' <small>Croissandeau, 1893</small>
**''[Chiloschiza](/source/Chiloschiza)'' <small>Moser, 1924</small>
**''[Clypeolontha](/source/Clypeolontha)'' <small>Li & Yang, 1999</small>
**''[Cocherellus](/source/Cocherellus)'' <small>Paulian, 1991</small>
**''[Cochliotodes](/source/Cochliotodes)'' <small>Burgeon, 1946</small>
**''[Conebius](/source/Conebius)'' <small>Fauvel, 1903</small>
**''[Cryptotrogus](/source/Cryptotrogus)'' <small>Kraatz, 1888</small>
**''[Cyphonotus](/source/Cyphonotus)'' <small>Fischer von Waldheim, 1823</small>
**''[Dermolepida](/source/Dermolepida)'' <small>Arrow, 1941</small>
**''[Dinacoma](/source/Dinacoma)'' <small>Casey, 1889</small>
**''[Engistanoxia](/source/Engistanoxia)'' <small>Lacroix, 2002</small>
**''[Entyposis](/source/Entyposis)'' <small>Kolbe, 1895</small>
**''[Euranoxia](/source/Euranoxia)'' <small>Semenov, 1890</small>
**''[Euryschiza](/source/Euryschiza)'' <small>Brenske, 1898</small>
**''[Exolontha](/source/Exolontha)'' <small>Reitter, 1902</small>
**''[Fossocarus](/source/Fossocarus)'' <small>Howden, 1961</small>
**''[Globulischiza](/source/Globulischiza)'' <small>Frey, 1974</small>
**''[Gnaphalostetha](/source/Gnaphalostetha)'' <small>Reiche & Saulcy, 1856</small>
**''[Gronocarus](/source/Gronocarus)'' <small>Schaeffer, 1927</small>
**''[Heptelia](/source/Heptelia)'' <small>Brenske, 1898</small>
**''[Holisonycha](/source/Holisonycha)'' <small>Péringuey, 1904</small>
**''[Holomelia](/source/Holomelia)'' <small>Brenske, 1891</small>
**''[Holopycnia](/source/Holopycnia)'' <small>Brenske, 1896</small>
**''[Holorhopaea](/source/Holorhopaea)'' <small>Britton, 1978</small>
**''[Homoeoschiza](/source/Homoeoschiza)'' <small>Kolbe, 1894</small>
**''[Hoplosternodes](/source/Hoplosternodes)'' <small>Burgeon, 1946</small>
**''[Howdenocarus](/source/Howdenocarus)'' <small>Hardy, 1978</small>
**''[Hypolepida](/source/Hypolepida)'' <small>Britton, 1978</small>
**''[Hypothyce](/source/Hypothyce)'' <small>Howden, 1968</small>
**''[Hypotrichia](/source/Hypotrichia_(beetle))'' <small>LeConte, 1861</small>
**''[Lecanotrogus](/source/Lecanotrogus)'' <small>Kolbe, 1894</small>
**''[Megacoryne](/source/Megacoryne)'' <small>Britton, 1987</small>
**''[Megarhopaea](/source/Megarhopaea)'' <small>Britton, 1978</small>
**†''Melolonthites'' <small>Heer, 1847</small>
**''[Menigionyx](/source/Menigionyx)'' <small>Arrow, 1941</small>
**''[Metatrogus](/source/Metatrogus)'' <small>Britton, 1978</small>
**''[Microphylla](/source/Microphylla)'' <small>Kraatz, 1890</small>
**''[Microrhopaea](/source/Microrhopaea)'' <small>Lea, 1920</small>
**†''Miolachnosterna'' <small>Wickham, 1914</small>
**''[Nanorhopaea](/source/Nanorhopaea)'' <small>Britton, 1978</small>
**''[Nematophylla](/source/Nematophylla)'' <small>Arrow, 1906</small>
**''[Nesadmiralitia](/source/Nesadmiralitia)'' <small>Prokofiev, 2016</small>
**''[Nuiba](/source/Nuiba)'' <small>Prokofiev, 2016</small>
**''[Octoplasia](/source/Octoplasia)'' <small>Brenske, 1892</small>
**''[Oligophylla](/source/Oligophylla)'' <small>Kraatz, 1894</small>
**''[Oreotrogus](/source/Oreotrogus)'' <small>Kolbe, 1897</small>
**''[Pararhopaea](/source/Pararhopaea)'' <small>Blackburn, 1911</small>
**''[Pentelia](/source/Pentelia)'' <small>Brenske, 1891</small>
**''[Perissosoma](/source/Perissosoma)'' <small>Waterhouse, 1875</small>
**''[Phiara](/source/Phiara_(beetle))'' <small>Brenske, 1897</small>
**''[Plectrodes](/source/Plectrodes)'' <small>Horn, 1867</small>
**''[Polyphyllum](/source/Polyphyllum)'' <small>Blanchard, 1851</small>
**''[Proteroschiza](/source/Proteroschiza)'' <small>Brenske, 1896</small>
**''[Pseudholophylla](/source/Pseudholophylla)'' <small>Blackburn, 1911</small>
**''[Pseudosymmachia](/source/Pseudosymmachia)'' <small>Dalla Torre, 1913</small>
**''[Pseudotocama](/source/Pseudotocama)'' <small>Keith, 2008</small>
**''[Ramilia](/source/Ramilia_(beetle))'' <small>Kolbe, 1894</small>
**''[Rhadinolontha](/source/Rhadinolontha)'' <small>Arrow, 1930</small>
**''[Rhizoproctus](/source/Rhizoproctus)'' <small>Kolbe, 1894</small>
**''[Rhopaea](/source/Rhopaea)'' <small>Erichson, 1847</small>
**''[Rhyxicephalus](/source/Rhyxicephalus)'' <small>Moser, 1921</small>
**''[Rhyxiderma](/source/Rhyxiderma)'' <small>Moser, 1924</small>
**''[Schismatocera](/source/Schismatocera)'' <small>Gautier Des Cottes, 1872</small>
**''[Sebakue](/source/Sebakue)'' <small>Péringuey, 1904</small>
**''[Spathoschiza](/source/Spathoschiza)'' <small>Arrow, 1902</small>
**''[Sphodroxia](/source/Sphodroxia)'' <small>Kraatz, 1890</small>
**''[Stenopegylis](/source/Stenopegylis)'' <small>Arrow, 1943</small>
**''[Suntemnonycha](/source/Suntemnonycha)'' <small>Péringuey, 1904</small>
**''[Syngeneschiza](/source/Syngeneschiza)'' <small>Brenske, 1898</small>
**''[Thyce](/source/Thyce)'' <small>LeConte, 1856</small>
**''[Tocama](/source/Tocama)'' <small>Reitter, 1902</small>
**''[Tricholontha](/source/Tricholontha)'' <small>Nomura, 1952</small>
**''[Trinoxia](/source/Trinoxia)'' <small>Brenske, 1894</small>
**''[Xenotrogus](/source/Xenotrogus)'' <small>Britton, 1978</small>
**''[Zietzia](/source/Zietzia)'' <small>Blackburn, 1894</small>

==Formerly placed here==
*''[Amphimallon](/source/Amphimallon)'' <small>Latreille, 1825</small> (European chafers)
*''[Brahmina](/source/Brahmina)'' <small>Blanchard, 1850</small>
*''[Holotrichia](/source/Holotrichia)'' <small>Hope, 1837</small>
*''[Lasiopsis](/source/Lasiopsis)'' <small>Erichson, 1847</small>
*''[Lepidiota](/source/Lepidiota)'' <small>Kirby, 1828</small>
*''[Leucopholis](/source/Leucopholis)'' <small>Dejean, 1833</small>
*''[Megistophylla](/source/Megistophylla)'' <small>Burmeister, 1855</small>
*''[Miridiba](/source/Miridiba)'' <small>Reitter, 1901</small>
*''[Phyllophaga](/source/Phyllophaga)'' <small>Harris, 1827</small> (May beetles)
*''[Rhizotrogus](/source/Rhizotrogus)'' <small>Lepeletier & Serville, 1825</small>

==References==
<references>

<ref name=Jia2021>
{{Cite journal| title = Morphological description of the white grub Melolontha incana (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini)
|last=Jia|first1=Zhi‐Chao|last2=Fang|first2=Hong|last3=Jiang|first3=Lu
|journal=Microscopy Research and Technique|volume=84|issue=5
|date=2021
}}</ref>

</references>
{{refbegin}}
* Bouchard, P., Y. Bousquet, A. Davies, M. Alonso-Zarazaga, J. Lawrence, C. Lyal, A. Newton, et al. (2011). "Family-group names in Coleoptera (Insecta)". ''ZooKeys, vol. 88'', 1–972.
{{refend}}

==Further reading==
{{refbegin}}
* Arnett, R. H. Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). (21 June 2002). ''American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea''. CRC Press LLC, Boca Raton, Florida {{ISBN|978-0-8493-0954-0}}.
* {{Cite book| author = Ross H. Arnett| date   = 30 July 2000| title  = American Insects: A Handbook of the Insects of America North of Mexico| publisher = CRC Press| isbn     = 978-0-8493-0212-1}}
* Richard E. White. (1983). ''Peterson Field Guides: Beetles''. Houghton Mifflin Company.
{{refend}}
{{-}}

{{Taxonbar|from=Q1175811}}
{{Use dmy dates|date=February 2018}}

Category:Melolonthinae
Category:Melolonthini

{{Melolonthinae-stub}}

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Adapted from the Wikipedia article [Melolonthini](https://en.wikipedia.org/wiki/Melolonthini) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Melolonthini?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.