# Iostephane

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***Iostephane*** is a [genus](/source/Genus) of [Mexican](/source/Mexico) flowering plants in the family [Asteraceae](/source/Asteraceae).[1][2][3] They were first published in G.Bentham & J.D.Hooker, Genera Plantarum Vol.2 on page 368 in 1873.[4]

The species are rosette-forming herbaceous [perennials](/source/Perennial_plants) that produce relatively large heads with yellow or purple (*I. heterophylla*) rays. They have large, somewhat quadrate cypselae ([achenes](/source/Achenes)), that may either have or lack a [pappus](/source/Pappus_(botany)).

Molecular phylogenetic studies by Schilling in 1991,[5] suggested that there is an unexpectedly close relationship between *Iostephane* and *[Dendroviguiera](/source/Dendroviguiera)* (formerly *Viguiera* sect. *Maculatae*), a genus of shrubs and trees that is also [endemic](/source/Endemic) to [Mexico](/source/Mexico).[6]

## Species

As accepted by [Plants of the World Online](/source/Plants_of_the_World_Online),[4] and Global Compositae Checklist;[7]

- *[Iostephane heterophylla](/source/Iostephane_heterophylla)* (Cav.) Benth. – widespread from [Chihuahua](/source/Chihuahua_(state)) to [Oaxaca](/source/Oaxaca)
- *[Iostephane madrensis](/source/Iostephane_madrensis)* (S.Wats.) Strother – [Chihuahua](/source/Chihuahua_(state))
- *[Iostephane papposa](/source/Iostephane_papposa)* J. J. Fay – [Oaxaca](/source/Oaxaca)
- *[Iostephane trilobata](/source/Iostephane_trilobata)* Hemsl. – [Chiapas](/source/Chiapas), [Oaxaca](/source/Oaxaca), [Puebla](/source/Puebla), [México State](/source/M%C3%A9xico_State)

## References

1. [Bentham, George. 1873. Genera Plantarum 2: 368](https://www.biodiversitylibrary.org/page/657503#page/371/mode/1up) in Latin

1. Strother, J. L. 1983. *Pionocarpus* becomes *Iostephane* (Compositae: Heliantheae): a synopsis. Madroño 30: 34-48.

1. [Tropicos, *Iostephane* Benth.](http://www.tropicos.org/Name/40031309)

1. ["*Iostephane* Benth. | Plants of the World Online | Kew Science"](https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:9496-1). *Plants of the World Online*. Retrieved 2 November 2023.

1. Schilling, E. E. and J. L. Panero. 1991. Evidence for a close relationship between *Iostephane* and *Viguiera* sect. *Maculatae* (Asteraceae: Heliantheae). American Journal of Botany 78: 1054-1062.

1. Schilling, E, E. and J. L. Panero 2011. A revised classification of subtribe Helianthinae (Asteraceae: Heliantheae). II. Derived lineages. Botanical Journal of the Linnean Society 167: 311-331.

1. ["Flann, C (ed) 2009+ Global Compositae Checklist"](https://archive.today/20141209155630/http://dixon.iplantcollaborative.org/CompositaeWeb/default.aspx?Page=NameDetails&TabNum=0&NameId=2ed3518e-4091-441d-9e4c-19f2409e14da). Archived from [the original](http://dixon.iplantcollaborative.org/CompositaeWeb/default.aspx?Page=NameDetails&TabNum=0&NameId=2ed3518e-4091-441d-9e4c-19f2409e14da) on 2014-12-09. Retrieved 2014-12-09.

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Adapted from the Wikipedia article [Iostephane](https://en.wikipedia.org/wiki/Iostephane) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Iostephane?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.