# Invasive species

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Non-native organism causing damage to an established environment

[North American beaver](/source/North_American_beaver) (*Castor canadensis*) dam [in Tierra del Fuego](/source/Beavers_in_Southern_Patagonia)

[Kudzu](/source/Kudzu) growing over and smothering trees in [Atlanta](/source/Atlanta), [Georgia](/source/Georgia_(U.S._state))

[Canada goldenrod](/source/Solidago_canadensis) (*Solidago canadensis*) as a roadside weed in Poland

*[Vinca](/source/Vinca)* in a garden[1]

An **invasive species** is an [introduced species](/source/Introduced_species) that harms its new environment.[2] Invasive species adversely affect [habitats](/source/Habitat) and [bioregions](/source/Bioregion), causing [ecological](/source/Ecology), [environmental](/source/Natural_environment), and/or [economic](/source/Economy) damage. Since the 20th century, invasive species have become serious economic, social, and environmental threats worldwide.

Invasion of long-established ecosystems by organisms is a natural phenomenon, but human-facilitated introductions have greatly increased the rate, scale, and geographic range of invasion. For millennia, [humans](/source/Human) have served as both accidental and deliberate dispersal agents, beginning with their [earliest migrations](/source/Early_human_migrations), accelerating in the [Age of Discovery](/source/Age_of_Discovery), and accelerating again with the spread of [international trade](/source/International_trade).

Notable [plant](/source/Plant) species that are invasive in many parts of the world include the [kudzu](/source/Kudzu) vine (*Pueraria*), [water hyacinth](/source/Pontederia_crassipes) *(Pontederia crassipes)*, [common ragweed](/source/Ambrosia_artemisiifolia) (*Ambrosia artemisiifolia*), and [yellow starthistle](/source/Centaurea_solstitialis) (*Centaurea solstitialis*). Notable [animal](/source/Animal) species that are invasive include [European rabbits](/source/European_rabbit) (*Oryctolagus cuniculus*), [domestic cats](/source/Domestic_cat) (*Felis catus*), and [red swamp crayfish](/source/Procambarus_clarkii) (*Procambarus clarkii*).

## Terminology

See also: [Glossary of invasion biology terms](/source/Glossary_of_invasion_biology_terms)

Invasive species are the subset of established non-native species that pose a threat to native species and biodiversity.[3] The term "invasive" is poorly defined and often very subjective.[4] Invasive species may be plants, animals, fungi, and microbes; some include native species that have invaded human habitats such as farms and landscapes.[5] Some broaden the term to include indigenous or "native" species that have [colonized](/source/Colonisation_(biology)) natural areas.[4] Some sources name *[Homo sapiens](/source/Homo_sapiens)* as an invasive species,[6][7] but broad appreciation of human learning capacity and their behavioral potential and [plasticity](/source/Phenotypic_plasticity) may argue against any such fixed categorization.[8]

The definition of "native" can also be controversial. For example, the ancestors of [modern horses](/source/Horse) (*Equus ferus)* [evolved](/source/Evolution_of_the_horse) in [North America](/source/North_America) and radiated to [Eurasia](/source/Eurasia) before becoming extinct in North America. Their reintroduction to North America, by Spanish [conquistadors](/source/Conquistador), led to a 1943 debate over whether the [feral horses](/source/Feral_horse) were native or exotic to the continent of their evolutionary ancestors.[9]

While invasive species can be studied within many subfields of biology, most research on invasive organisms has been in [ecology](/source/Ecology) and [biogeography](/source/Biogeography). Much of the work has been influenced by [Charles Elton's](/source/Charles_Sutherland_Elton) 1958 book *The Ecology of Invasion by Animals and Plants* which creates a generalized picture of biological invasions.[10][11] Studies remained sparse until the 1990s.[11] This research, largely field observational studies, has disproportionately been concerned with [terrestrial plants](/source/Terrestrial_plant).[11] The rapid growth of the field has driven a need to standardize the language used to describe invasive species and events. Despite this, little standard terminology exists. The field lacks any official designation but is commonly referred to as "invasion ecology" or more generally "invasion biology".[10][11] This lack of standard terminology has arisen due to the interdisciplinary nature of the field, which borrows terms from disciplines such as [agriculture](/source/Agriculture), [zoology](/source/Zoology), and [pathology](/source/Pathology), as well as due to studies being performed in isolation.[12][10]

Colautti and MacIsaac nomenclature[4] Stage Characteristic 0 Propagules residing in a donor region I Traveling II Introduced III Localized and numerically rare IVa Widespread but rare IVb Localized but dominant V Widespread and dominant

In an attempt to avoid the ambiguous, subjective, and pejorative vocabulary that so often accompanies discussion of invasive species, even in scientific papers, Colautti and MacIsaac proposed a new nomenclature system. Based on [biogeography](/source/Biogeography) rather than on [taxa](/source/Taxa), [4] the proposed system emphasizes ecology over taxonomy, [human health](/source/Human_health), and economic factors. The model evaluated individual populations rather than entire species. It classified each population based on its success in that environment. This model applied equally to indigenous and to introduced species, and did not automatically categorize successful introductions as harmful.[4]

The USDA's National Invasive Species Information Center defines invasive species very narrowly. According to Executive Order 13112, "'Invasive species' means an alien species whose introduction does or is likely to cause economic or environmental harm or harm to human health."[13]

## Causes

Typically, an introduced species must persist at low [population densities](/source/Population_density) before it can become invasive in a new environment.[14] At low densities, introduced species often face difficulties in reproducing and sustaining a viable population—sometimes requiring multiple introductions before establishment occurs. Patterns of repeated human movement, such as ship traffic between ports or vehicles travelling along major highways, can create recurring opportunities for arrival and establishment, a phenomenon known as high [propagule pressure](/source/Propagule_pressure).[15]

### Ecosystem-based mechanisms

In [ecosystems](/source/Ecosystem), the availability of resources determines the impact of additional species on the ecosystem. Stable ecosystems have a resource equilibrium, which can be changed fundamentally by the arrival of invasive species.[16] When changes such as a [forest fire](/source/Forest_fire) occur, normal [ecological succession](/source/Ecological_succession) favors native [grasses](/source/Graminoid) and [forbs](/source/Forb). An introduced species that can spread faster than natives can outcompete native species for food, squeezing the natives out. [Nitrogen](/source/Nitrogen) and [phosphorus](/source/Phosphorus) are often the limiting factors in these situations.[17] Every species occupies an [ecological niche](/source/Ecological_niche) in its native ecosystem; some species fill large and varied roles, while others are highly specialized. Invading species may occupy unused niches, or create new ones.[18] For example, [edge effects](/source/Edge_effect) describe what happens when part of an ecosystem is disturbed, as in when land is cleared for [agriculture](/source/Agriculture). The boundary between the remaining undisturbed habitat and the newly cleared land itself forms a distinct new habitat, creating new winners and losers, and potentially hosting species that would not otherwise thrive outside the boundary habitat.[19]

In 1958, [Charles S. Elton](/source/Charles_S._Elton) claimed that ecosystems with higher [species diversity](/source/Species_diversity) were less subject to invasive species because fewer niches remained unoccupied.[20] Other ecologists later pointed to highly diverse, but heavily invaded ecosystems, arguing that ecosystems with high species diversity were more susceptible to invasion.[21] This debate hinged on the [spatial scale](/source/Scale_(spatial)) of invasion studies. Small-scale studies tended to show a negative relationship between [diversity](/source/Biodiversity) and invasion, while large-scale studies tended to show the reverse, perhaps a side-effect of invasives' ability to capitalize on increased resource availability and weaker species interactions that are more common when larger samples are considered.[22][23] However, this pattern does not seem to hold true for invasive vertebrates.[24]

The [brown tree snake](/source/Brown_tree_snake) (*Boiga irregularis*) has had an impact on the native bird population of the island ecosystem of [Guam](/source/Guam).

[Island ecosystems](/source/Island_ecosystem) may be more prone to invasion because their species face few strong competitors and predators, and because their distance from colonizing species populations makes them more likely to have "open" niches.[25] For example, native bird populations on [Guam](/source/Guam) have been decimated by the invasive [brown tree snake](/source/Brown_tree_snake) (*Boiga irregularis*).[26] However, the geographical distance and open water characteristic of island ecosystems can also delay colonisation by invasive species. For example, [island nations](/source/Island_country) such as Australia and the Galapagos have a high proportion of their island ecology consisting of endemic species due to immigration of other species being limited by oceans and their geographical isolation.[27]

In [New Zealand](/source/Invasive_species_in_New_Zealand) the first invasive species were the [dogs](/source/Dog) and [rats](/source/Polynesian_rat) brought by [Polynesian](/source/Polynesians) settlers around 1300. These and other introductions devastated endemic New Zealand species.[28][29] The colonization of [Madagascar](/source/Madagascar) brought similar harm to its ecosystems.[30] Logging has caused harm directly by destroying habitat, and has allowed non-native species such as [prickly pear](/source/Opuntia) (*Opuntia*) and [silver wattle](/source/Acacia_dealbata) (*Acacia dealbata*) to invade.[31][32] The [water hyacinth](/source/Eichhornia_crassipes) (*Pontederia crassipes*) forms dense mats on water surfaces, limiting light penetration and hence harming aquatic organisms, and creating substantial management costs.[33][34] The [shrub lantana](/source/Lantana_camara) (*Lantana camara*) is now considered invasive in over 60 countries, and has invaded large geographies in several countries prompting aggressive federal efforts to control it.[35][36]

Along with island ecosystems, intensively managed fenced areas are more prone to invasion. One reason is that species can enter through ways that fences cannot block. In the case of many plant species, dispersal can occur through wind, water, and birds carrying seeds either internally or externally. Small animals or insects are sometimes also able to make it through fenced areas.[37][38][39] Another reason is that intensive management methods create opportunities for these invasive species to thrive. When fenced areas are established, they are intensively managed through clearing vegetation, mowing, and disturbing the soil. This reduces competition from the native plant species in the area and exposes the soil so that invasive species can easily populate the area.[40] The purpose of these fenced areas is often to keep unwanted [herbivores](/source/Herbivore) out of the areas. Because there are no threats to the invasive plants that can take over intensively managed fenced areas, they can flourish without predatory threats.[41] Finally, in general, invasive species have traits that favor their survival. Most invasive species are extremely resilient and have traits that favor their establishment in areas where they are not native. These traits, along with the intensive management of the fenced areas, create an ideal environment in which these invasive species can thrive.[42]

Primary geomorphological effects of invasive plants are bioconstruction and bioprotection. For example, [kudzu](/source/Pueraria_montana) (*Pueraria montana*), a vine native to [Asia](/source/Asia), was widely introduced in the [southeastern United States](/source/Southeastern_US) in the early 20th century to control [soil erosion](/source/Soil_erosion). The primary geomorphological effects of invasive animals are [bioturbation](/source/Bioturbation), [bioerosion](/source/Bioerosion), and bioconstruction. For example, invasions of the [Chinese mitten crab](/source/Chinese_mitten_crab) (*Eriocheir sinensis*) have resulted in higher bioturbation and bioerosion rates.[43]

A native species can also become harmful and effectively invasive to its native environment after human alterations to its [food web](/source/Food_web). This has been the case with the [purple sea urchin](/source/Strongylocentrotus_purpuratus) (*Strongylocentrotus purpuratus*), which has decimated kelp forests along the northern California coast due to overharvesting of its natural predator, the [California sea otter](/source/Sea_otter) (*Enhydra lutris*).[44]

### Species-based mechanisms

[Japanese knotweed](/source/Reynoutria_japonica) (*Reynoutria japonica*) is considered one of the [world's worst invasive species](/source/100_of_the_World's_Worst_Invasive_Alien_Species).

[Cats](/source/Cat) (*Felis catus*) (here, killing a [woodpecker](/source/Woodpecker)) are [considered invasive species](/source/Cats_in_Australia) in Australia and [negatively impact wildlife](/source/Cat_predation_on_wildlife) worldwide.

Invasive species appear to have specific traits or specific combinations of traits that allow them to outcompete [native species](/source/Native_species). In some cases, these characteristics include rates of growth and reproduction. In other cases, invasive species interact with native species other more directly. One study found that 86% of invasive species could be identified from such traits alone.[45] Another study found that invasive species often had only a few of the traits, and that noninvasive species had these also.[45][46][47] Common invasive species traits include fast growth and rapid [reproduction](/source/Reproduction), such as [vegetative reproduction](/source/Vegetative_reproduction) in plants;[45] association with [humans](/source/Human);[48] and prior successful invasions.[49] [Domestic cats](/source/Domestic_cat) (*Felis catus*) are effective predators of [wildlife](/source/Wildlife). They have become feral and invasive in places such as the [Florida Keys](/source/Florida_Keys).[50]

An introduced species might become invasive if it can outcompete native species for resources. If these species evolved under great [competition](/source/Competition_(biology)) or [predation](/source/Predation), then the new environment may host fewer able competitors, allowing the invader to proliferate. [Ecosystems](/source/Ecosystem) used to their fullest capacity by native species can be modeled as [zero-sum](/source/Zero-sum) systems, in which any gain for the invader is a loss for the native. However, such [unilateral](/source/Unilateral) competitive superiority (and extinction of native species with increased populations of the invader) is not the rule.[21][51]

*[Lantana](/source/Lantana)*, abandoned [citrus](/source/Citrus_grove), [Sdei Hemed](/source/Sdei_Hemed)

An invasive species might be able to use resources previously unavailable to native species, such as deep water accessed by a long [taproot](/source/Taproot), or to live on previously uninhabited soil types. For example, [barbed goatgrass](/source/Aegilops_triuncialis) (*Aegilops triuncialis*) was introduced to [California](/source/California) on [serpentine soils](/source/Serpentine_soil), which have low water-retention, low nutrient levels, a high [magnesium](/source/Magnesium)/[calcium](/source/Calcium) ratio, and possible [heavy metal](/source/Heavy_metal_(chemistry)) toxicity. Plant populations on these soils tend to show low density, but goatgrass can form dense stands on these soils and crowd out native species.[52]

Invasive species may also alter their environment by releasing chemical compounds, modifying [abiotic](/source/Abiotic) factors, or affecting the behavior of [herbivores](/source/Herbivore), all of which can impact other species. Some, like [mother of thousands](/source/Kalanchoe_daigremontiana) (*Kalanchoe daigremontana*), produce [allelopathic compounds](/source/Allelopathy) that inhibit competitors.[53] Others like the [toad plant](/source/Stapelia_gigantea) (*Stapelia gigantea)* [facilitate](/source/Ecological_facilitation) the growth of seedlings of other species in arid environments by providing appropriate [microclimates](/source/Microclimate) and preventing herbivores from eating seedlings.[54]

Changes in [fire regimens](/source/Fire_regime) are another form of facilitation. [Cheatgrass](/source/Bromus_tectorum) (*Bromus tectorum*)*,* originally from Eurasia, is highly fire-adapted. It spreads rapidly after burning, and increases the frequency and intensity of fires by providing large amounts of dry [detritus](/source/Detritus) during the fire season in western North America. Where it is widespread, it has altered the local fire regimen so much that native plants cannot survive the frequent fires, allowing it to become dominant in its introduced range.[55]

[Ecological facilitation](/source/Ecological_facilitation) occurs where one species, including invasive species, physically modifies a habitat in ways advantageous to other species. For example, [zebra mussels](/source/Zebra_mussel) (*Dreissena polymorpha*) increase habitat complexity on lake floors, providing crevices in which [invertebrates](/source/Invertebrate) live. This increase in complexity, together with the nutrition provided by the waste products of mussel [filter-feeding](/source/Filter_feeder), increases the density and diversity of [benthic](/source/Benthic_zone) invertebrate communities.[56]

Introduced species may spread rapidly and unpredictably.[57] When [bottlenecks](/source/Population_bottleneck) and [founder effects](/source/Founder_effect) cause a great decrease in the population size and may constrict [genetic variation](/source/Genetic_variation),[58] individuals begin to show additive variance as opposed to epistatic variance. This conversion can lead to increased variance in the founding populations, which permits [rapid evolution](/source/Rapid_evolution).[59] Selection may then act on the capacity to disperse as well as on physiological tolerance to new stressors in the environment, such as changed temperature and different predators and prey.[60]

Rapid adaptive evolution through intraspecific phenotypic plasticity, [pre-adaptation](/source/Exaptation), and post-introduction evolution lead to offspring that have higher fitness. Critically, plasticity permits changes to better suit the individual to its environment. Pre-adaptations and evolution after the introduction reinforce the success of the introduced species.[61]

The [enemy release hypothesis](/source/Enemy_release_hypothesis) states that evolution leads to ecological balance in every ecosystem. No single species can occupy a majority of an ecosystem due to the presences of competitors, predators, and diseases. Introduced species moved to a novel habitat can become invasive, with rapid population growth, when these controls do not exist in the new ecosystem.[62]

## Vectors

Non-native species have many [vectors](/source/Vector_(epidemiology)), but most are associated with human activity. Natural [range](/source/Range_(biology)) extensions are common, but humans often carry specimens faster and over greater distances than natural forces.[63] An early human vector occurred when prehistoric humans introduced the [Pacific rat](/source/Polynesian_rat) (*Rattus exulans*) to [Polynesia](/source/Polynesia).[64]

Humans have historically acted as vectors, both deliberately and accidentally. During the colonial era, there were acclimatization societies that aimed to deliberately establish animals and plants that were recognizable to Europeans. These societies felt that this would be an "improvement" to nature by adding species that were nostalgic and valuable.[65] Another reason for deliberate introduction was that some species are thought to act as biocontrol. The [cane toad](/source/Cane_toad) was introduced in Australia in hopes of controlling the cane beetle population. Cane beetles are pests of the sugar cane crop. While only 100 toads were initially imported, there are now over 200 million present in Australia.[66] Another example of animal introduction is the [Indian mongoose](/source/Indian_grey_mongoose), which was introduced in Hawaii to control the rat population as the rats were damaging sugar cane yields. This, however, ended up being ineffective because rats are nocturnal while the mongoose is [diurnal](/source/Diurnality). Their introduction ended up causing new problems for the island's ecosystems.[67]

The [Chinese mitten crab](/source/Chinese_mitten_crab) (*Eriocheir sinensis*) may have been purposely introduced in [Europe](/source/Europe) and [North America](/source/North_America) because of its commercial value.

Vectors also include plants or seeds imported for [horticulture](/source/Horticulture). The [pet trade](/source/Pet_trade) moves animals across borders, where they can escape and become invasive. Organisms may also stow away on transport vehicles. Incidental human assisted transfer is the main cause of introductions – other than in [polar regions](/source/Polar_regions_of_Earth).[68] Diseases may be vectored by invasive insects: the [Asian citrus psyllid](/source/Diaphorina_citri) (*Diaphorina citri*) carries the bacterial disease [citrus greening](/source/Citrus_greening_disease).[69] The arrival of invasive [propagules](/source/Propagule) to a new site is a function of the site's invasibility.[70] Recreation is one of the most underestimated vectors for invasive species spread. Seeds, spores, larvae, and pathogens can hitchhike on boats, boots, waders, bikes, pets, vehicles, and firewood, moving farther and faster than natural dispersal ever could.[71]

Many invasive species, once they are dominant in the area, become essential to the ecosystem of that area, and their removal could be harmful.[72] Economics plays a major role in exotic species introduction. High demand for the valuable [Chinese mitten crab](/source/Chinese_mitten_crab) is one explanation for the possible intentional release of the species in foreign waters.[73]

### Within the aquatic environment

[Maritime trade](/source/Maritime_transport) has rapidly affected the way marine organisms are transported within the ocean; new means of species transport include hull fouling and ballast water transport. In fact, Molnar et al. 2008 documented the pathways of hundreds of marine invasive species and found that shipping was the dominant mechanism for the transfer of invasive species.[74]

Cargo ship [de-ballasting](/source/Sailing_ballast)

Many marine organisms can attach themselves to vessel hulls. Such organisms are easily transported from one body of water to another, and are a significant risk factor for a biological invasion event.[75] Controlling for vessel hull fouling is voluntary and there are no regulations currently in place to manage hull fouling. However, the governments of [California](/source/California) and [New Zealand](/source/New_Zealand) have announced more stringent control for vessel hull fouling within their respective jurisdictions.[76]

Another vector of non-native aquatic species is [ballast water](/source/Ballast_water_discharge_and_the_environment) taken up at sea and released in port by transoceanic vessels.[77][78] Some 10,000 species are transported via ballast water each day.[79] Many of these are harmful. For example, freshwater [zebra mussels](/source/Zebra_mussel) (*Dreissena polymorpha*) from Eurasia most likely reached the [Great Lakes](/source/Great_Lakes) via ballast water.[80] The mussels outcompete native organisms for oxygen and food, and can be transported in the small puddle left in a supposedly empty ballast tank.[77] Also, three lineages of the Asian clam genus *[Corbicula](/source/Corbicula)* (*C. fluminea*, *C. fluminalis*, *C. leana*) are globally invasive. Molecular data confirm their taxonomic identity and reveal multiple independent introduction events.[81] Regulations attempt to mitigate such risks,[82][83] not always successfully.[84] [Climate change](/source/Climate_change) is causing an increase in [ocean temperature](/source/Ocean_temperature). These changes to the environment in turn cause range shifts in organisms,[85][86] creating new species interactions. For example, organisms in a ballast tank of a ship traveling from the temperate zone through tropical waters may experience temperature fluctuations as much as 20 °C.[87] Heat challenges during transport may enhance the stress tolerance of species in their non-native range, by selecting for genotypes that will survive a second applied heat stress, such as increased ocean temperature in the founder population.[88]

### Effects of wildfire and firefighting

Invasive species often exploit disturbances to an ecosystem ([wildfires](/source/Wildfire), [roads](/source/Road), [foot trails](/source/Foot_trail)) to colonize an area. Large wildfires can [sterilize](/source/Sterilization_(microbiology)) soils, while adding [nutrients](/source/Nutrient).[17] Invasive plants that can regenerate from their roots then have an advantage over natives that rely on seeds for propagation.[55]

## Adverse effects

Invasive species can affect the invaded habitats, communities, and ecosystems adversely, causing ecological, environmental, and/or economic damage.[89][90][91]

### Ecological

The [European Union](/source/European_Union) defines "Invasive Alien Species" as those that are outside their natural distribution area, and that threaten [biological diversity](/source/Biological_diversity).[92][93] Biotic invasion is one of the five top drivers for global [biodiversity loss](/source/Biodiversity_loss), and is increasing because of tourism and [globalization](/source/Globalization).[94][95] This may be particularly true in inadequately regulated [fresh water](/source/Fresh_water) systems, though [quarantines](/source/Quarantine) and [ballast water](/source/Ballast_water) rules have improved the situation.[96]

[American alligator](/source/American_alligator) (*Alligator mississippiensis*) combatting a [Burmese python](/source/Burmese_python_in_Florida) (*Python bivittatus*)  in [Florida](/source/Florida)

Invasive species may drive local native species to extinction via [competitive](/source/Competition_(biology)) exclusion, [niche](/source/Ecological_niche) displacement, or [hybridization](/source/Hybrid_(biology)) with related native species. Therefore, besides their economic ramifications, alien invasions may result in extensive changes in the structure, composition and global distribution of the biota at sites of introduction, leading ultimately to the homogenization of the world's fauna and flora and the [loss of biodiversity](/source/Loss_of_biodiversity).[97][98] It is difficult to unequivocally attribute extinctions to a species invasion, though for example there is strong evidence that the extinction of about 90 [amphibian](/source/Amphibian) species was caused by the [chytrid fungus](/source/Chytridiomycosis) (*Batrachochytrium dendrobatidis*) spread by international trade.[99]

Multiple successive introductions of different non-native species can worsen the total effect, as with the introductions of the [amethyst gem clam](/source/Amethyst_gem_clam) (*Gemma gemma*) and the [European green crab](/source/Carcinus_maenas) (*Carcinus maenas*). The gem clam was introduced into California's [Bodega Harbor](/source/Bodega_Bay) from the US East Coast a century ago. On its own, it did not displace native clams (*Nutricola* spp.). However, in the mid-1990s, the introduction of the European green crab resulted in an increase of the amethyst gem at the expense of the native clams.[100] In [India](/source/India), multiple invasive plants have invaded 66% of natural areas, reducing the densities of native forage plants, declining the habitat-use by wild [herbivores](/source/Herbivore) and threatening the long-term sustenance of dependent [carnivores](/source/Carnivore), including [tigers](/source/Tiger).[101][102]

Invasive species can change the functions of ecosystems. For example, invasive plants can alter the [fire regime](/source/Fire_regime) (e.g., [cheatgrass](/source/Bromus_tectorum), *Bromus tectorum*), [nutrient cycling](/source/Nutrient_cycling) (e.g., [smooth cordgrass](/source/Sporobolus_alterniflorus), *Spartina alterniflora*), and hydrology (e.g.,*[Tamarix](/source/Tamarix)*) in native ecosystems.[103] Invasive species that are closely related to rare native species have the potential to hybridize with the native species. Harmful effects of hybridization have led to a decline and even extinction of native species.[104][105] For example, [hybridization](/source/Hybridization_(biology)) with introduced cordgrass threatens the existence of [California cordgrass](/source/Sporobolus_foliosus) (*Spartina foliosa*) in [San Francisco Bay](/source/San_Francisco_Bay).[106] Invasive species cause competition for native species, and because of this 400 of the 958 endangered species under the [Endangered Species Act](/source/Endangered_Species_Act_of_1973) are at risk.[107]

Poster from the [State of California](/source/Government_of_California) asking campers to not move firewood around, avoiding the spread of invasive species

The unintentional introduction of forest pest species and plant pathogens can change [forest ecology](/source/Forest_ecology) and damage the [timber industry](/source/Timber_industry). Overall, [forest ecosystems](/source/Forest_ecosystem) in the U.S. are widely invaded by exotic pests, plants, and pathogens.[108][109]

The [Asian long-horned beetle](/source/Asian_long-horned_beetle) (*Anoplophora glabripennis*) was first introduced into the U.S. in 1996, and was expected to infect and damage millions of acres of hardwood trees. As of 2005 thirty million dollars had been spent in attempts to eradicate this pest and protect millions of trees in the affected regions.[110] The [woolly adelgid](/source/Woolly_adelgid) (*Adelges tsugae*) has inflicted damage on old-growth spruce, fir and [hemlock](/source/Tsuga) forests and damages the [Christmas tree](/source/Christmas_tree) industry.[111] [Chestnut blight](/source/Chestnut_blight) (*Cryphonectria parasitica*) and [Dutch elm disease](/source/Dutch_elm_disease) ([Ascomycota](/source/Ascomycota)) are plant pathogens with serious impacts.[112][113][114] [Garlic mustard](/source/Alliaria_petiolata) (*Alliaria petiolata)* is one of the most problematic invasive plant species in eastern North American forests, where it is highly invasive of the [understory](/source/Understory), reducing the growth rate of tree seedlings and threatening to modify the forest's tree composition.[115]

The [Kākāpō](/source/K%C4%81k%C4%81p%C5%8D) is a threatened species partially because of [predation](/source/Predation) by invasive species (mostly rats and cats), there are only ≈240 individuals left worldwide [116]

Native [species](/source/Species) can be threatened with [extinction](/source/Extinction)[117] through the process of [genetic pollution](/source/Genetic_pollution). Genetic pollution is unintentional [hybridization](/source/Hybrid_(biology)) and [introgression](/source/Introgression), which leads to homogenization or replacement of local [genotypes](/source/Genotypes) as a result of either a numerical or [fitness](/source/Fitness_(biology)) advantage of the introduced species.[118] Genetic pollution occurs either through introduction or through habitat modification, where previously isolated species are brought into contact with the new genotypes. Invading species have been shown to adapt to their new environments in a remarkably short amount of time.[117] The population size of invading species may remain small for a number of years and then experience an explosion in population, a phenomenon known as "the lag effect".[103]

[Argentine ants](/source/Argentine_ant) (*Linepithema humile*), which form [supercolonies](/source/Ant_supercolony) across continents, are ranked among the [world's 100 worst invasive animal species](/source/100_of_the_World's_Worst_Invasive_Alien_Species).[119]

Hybrids resulting from invasive species interbreeding with native species can incorporate their genotypes into the gene pool over time through [introgression](/source/Introgression). Similarly, in some instances a small invading population can threaten much larger native populations. For example, cordgrass (*Spartina alterniflora)* was introduced in the San Francisco Bay and hybridized with native *California cordgrass (Spartina foliosa).* The higher pollen count and male fitness of the invading species resulted in introgression that threatened the native populations due to lower pollen counts and lower viability of the native species.[120] Reduction in fitness is not always apparent from [morphological](/source/Morphology_(biology)) observations alone. Some degree of [gene flow](/source/Gene_flow) is normal, and preserves constellations of [genes](/source/Gene) and genotypes.[105][121] An example of this is the interbreeding of migrating [coyotes](/source/Coyote) (*Canis latrans*) with the [critically endangered](/source/Critically_endangered) [red wolf](/source/Red_wolf) (*Canis rufus*), in areas of eastern [North Carolina](/source/North_Carolina) where the red wolf was reintroduced, reducing wolf numbers.[122]

### Environmental

In South Africa's [Cape Town](/source/Cape_Town) region, analysis demonstrated that the restoration of priority source water sub-catchments through the removal of thirsty alien plant invasions (such as Australian acacias, [pines](/source/Pine), [eucalyptus](/source/Eucalyptus), and [Australian black wattle](/source/Black_wattle)) would generate expected annual water gains of 50 billion liters within 5 years compared to the business-as-usual scenario (which is important as Cape Town experiences significant [water scarcity](/source/Water_scarcity)). This is the equivalent to one-sixth of the city's current supply needs. These annual gains will double within 30 years. The catchment restoration is significantly more cost-effective then other water augmentation solutions (1/10 the unit cost of alternative options).[123] A water fund has been established, and these exotic species are being eradicated.[124]

### Human health

Invasive species can affect human health. With the alteration in ecosystem functionality (due to homogenization of biota communities), invasive species have resulted in negative effects on human well-being, which includes reduced resource availability, unrestrained spread of human diseases, recreational and educational activities, and tourism.[125][126] Alien species have caused diseases including [human immunodeficiency virus](/source/HIV) (HIV), [monkey pox](/source/Monkeypox), and [severe acute respiratory syndrome](/source/Severe_acute_respiratory_syndrome) (SARS).[126]

Invasive species and accompanying control efforts can have long term [public health](/source/Public_health) implications. For instance, [pesticides](/source/Pesticide) applied to treat a particular pest species could pollute soil and surface water.[110] Encroachment of humans into previously remote ecosystems has exposed exotic diseases such as [HIV](/source/HIV) to the wider population.[110] Introduced [birds](/source/Bird) (e.g. [pigeons](/source/Pigeons)), [rodents](/source/Rodent), and [insects](/source/Insect) (e.g. [mosquito](/source/Mosquito), [flea](/source/Flea), [louse](/source/Louse) and [tsetse fly](/source/Tsetse_fly) pests) can serve as vectors and reservoirs of human afflictions. Throughout recorded history, epidemics of human diseases, such as [malaria](/source/Malaria), [yellow fever](/source/Yellow_fever), [typhus](/source/Typhus), and [bubonic plague](/source/Bubonic_plague), spread via these vectors.[20] A recent example of an introduced disease is the spread of the [West Nile virus](/source/West_Nile_virus), which killed humans, birds, mammals, and reptiles.[127] The introduced [Chinese mitten crabs](/source/Chinese_mitten_crab) (*Eriocheir sinensis*) are carriers of [Asian lung fluke](/source/Paragonimus_westermani).[80] Waterborne disease agents, such as [cholera](/source/Cholera) bacteria (*[Vibrio cholerae](/source/Vibrio_cholerae)*), and causative agents of [harmful algal blooms](/source/Harmful_algal_bloom) are often transported via ballast water.[128]

### Economic

Invasive [water hyacinths](/source/Water_hyacinth) (*Pontederia crassipes*) clog the [Pasig River](/source/Pasig_River) in [Manila](/source/Manila), [Philippines](/source/Philippines) in October 2020.[129]

Globally, invasive species management and control are substantial economic burdens, with expenditures reaching approximately $1.4 trillion annually.[62] The economic impact of invasive species alone was estimated to exceed $423 billion annually as of 2019. This cost has exhibited a significant increase, quadrupling every decade since 1970, underscoring the escalating financial implications of these biological invasions.[130]

Invasive species contribute to [ecological degradation](/source/Environmental_degradation), altering ecosystem functionality and reducing the services ecosystems provide. This necessitates additional expenditures to control the spread of biological invasions, mitigate further impacts, and restore affected ecosystems. For example, the damage caused by 79 invasive species between 1906 and 1991 in the United States has been estimated at US$120 billion. Similarly, in [China](/source/China), invasive species have been reported to reduce the country's gross domestic product (GDP) by 1.36% per year.[126][131]

The management of biological invasions can be costly. In [Australia](/source/Australia), for instance, the expense to monitor, control, manage, and research invasive weed species is approximately AU$116.4 million per year, with costs directed solely to central and local governments.[126]

While, in some cases, invasive species may offer economic benefits—such as the potential for commercial forestry from invasive trees—these benefits are generally overshadowed by the substantial costs associated with biological invasions.[132] In most cases, the [economic returns](/source/Returns_(economics)) from invasive species are far less than the costs they impose.[133][126]

#### United States

In the [Great Lakes region](/source/Great_Lakes_region) the [sea lamprey](/source/Sea_lamprey) (*Petromyzon marinus*) is an invasive species. In its original habitat, it had co-evolved as a [parasite](/source/Parasitism) that did not kill its host. However, in the Great Lakes region, it acts as a predator and can consume up to 40 pounds of fish in its 12–18 month feeding period.[134] Sea lampreys prey on all types of large fish such as [lake trout](/source/Lake_trout) (*Salvelinus namaycush*) and [salmon](/source/Salmon). The sea lampreys' destructive effects on large fish negatively affect the fishing industry and have helped cause the collapse of the population of some species.[134]

[Economic costs](/source/Economic_cost) from invasive species can be separated into direct costs through production loss in agriculture and forestry, and management costs. Estimated damage and control costs of invasive species in the U.S. amount to more than $138 billion annually.[110] Economic losses can occur through loss of [recreational](/source/Recreation) and [tourism](/source/Tourism) revenues.[135] When economic costs of invasions are calculated as production loss and management costs, they are low because they do not consider environmental damage; if monetary values were assigned to the [extinction](/source/Extinction) of species, loss in biodiversity, and loss of [ecosystem services](/source/Ecosystem_services), costs from impacts of invasive species would drastically increase.[110] It is often argued that the key to invasive species management is early detection and rapid response.[136] However, early response only helps when the invasive species is not frequently reintroduced into the managed area, and the cost of response is affordable.[137]

*[Parthenium hysterophorus](/source/Parthenium_hysterophorus)*, [Achanakmar Tiger Reserve](/source/Achanakmar_Tiger_Reserve)

[Weeds](/source/Weed) reduce yield in [agriculture](/source/Agriculture). Many weeds are accidental introductions that accompany imports of commercial seeds and plants. Introduced weeds in pastures compete with native forage plants, threaten young [cattle](/source/Cattle) (e.g., leafy spurge, *[Euphorbia virgata](/source/Euphorbia_virgata)*) or are unpalatable because of [thorns and spines](/source/Thorns%2C_spines%2C_and_prickles) (e.g., [yellow starthistle](/source/Yellow_starthistle), *Centaurea solstitialis*). Forage loss from invasive weeds on pastures amounts to nearly [US$](/source/US%24)1 billion in the U.S.[110] A decline in pollinator services and loss of fruit production has been caused by [honey bees](/source/Western_honeybee) (*Apis mellifera*) infected by the invasive [varroa mite](/source/Varroa_mite) (*Varroa destructor*). Introduced rats (*[Rattus rattus](/source/Rattus_rattus)* and *[R. norvegicus](/source/Rattus_norvegicus)*) have become serious pests[138] on farms, destroying stored grains.[110] The introduction of [leaf miner flies](/source/Agromyzidae) (Agromyzidae), including the [American serpentine leaf miner](/source/Liriomyza_trifolii) (*Liriomyza trifolii*), to California has caused losses in California's [floriculture](/source/Floriculture) industry, as the larvae of these invasive species feed on ornamental plants.[139]

Invasive plant pathogens and insect vectors for plant diseases can suppress agricultural yields and harm nursery stock. [Citrus](/source/Citrus) greening is a [bacterial disease](/source/Bacterial_disease) vectored by the invasive Asian citrus psyllid (*Diaphorina citri*). As a result, citrus is under quarantine and highly regulated in areas where the psyllid has been found.[69]

Invasive species can impact outdoor recreation, such as fishing, [hunting](/source/Hunting), [hiking](/source/Hiking), [wildlife viewing](/source/Wildlife_viewing), and water-based activities. They can damage environmental services including [water quality](/source/Water_quality), plant and animal diversity, and [species abundance](/source/Species_abundance), though the extent of this is under-researched.[140] [Eurasian watermilfoil](/source/Myriophyllum_spicatum) (*Myriophyllum spicatum*) in parts of the US, fills lakes with plants, complicating fishing and boating.[141] The loud call of the introduced [common coqui](/source/Common_coqui) (*Eleutherodactylus coqui*) depresses real estate values in affected neighborhoods of [Hawaii](/source/Hawaii).[142] The large webs of the [orb-weaving spider](/source/Zygiella_x-notata) (*Zygiella x-notata*), invasive in California, disrupts garden work.[143]

#### Europe

The overall economic cost of invasive alien species in Europe between 1960 and 2020 has been estimated at around US$140 billion (including potential costs that may or may not have actually materialized) or US$78 billion (only including observed costs known to have materialized). These estimates are very conservative. Models based on these data suggest a true *annual* cost of around US$140 billion in 2020.[144]

Italy is one of the most invaded countries in [Europe](/source/Europe), with an estimate of more than 3,000 alien species. The impacts of invasive alien species on the economy has been wide-ranging, from management costs, to loss of crops, to infrastructure damage. The overall economic cost of invasions to Italy between 1990 and 2020 was estimated at US$819.76 million (EUR€704.78 million). However, only 15 recorded species have more reliably estimated costs, hence the actual cost may be much larger than the aforementioned sum.[145]

France has an estimated minimum of 2,750 introduced and invasive alien species. Renault et al. (2021) obtained 1,583 cost records for 98 invasive alien species and found that they caused a conservative total cost between US$1.2 billion and 11.5 billion over the period 1993–2018. This study extrapolated costs for species invading France, but for which costs were reported only in other countries but not in France, which yielded an additional cost ranging from US$151 million to $3.03 billion. Damage costs were nearly eight times higher than management expenditure. Insects, and in particular the [Asian tiger mosquito](/source/Aedes_albopictus) (*Aedes albopictus*) and the [yellow fever mosquito](/source/Aedes_aegypti) (*A. aegypti*), created the highest economic costs, followed by non-graminoid terrestrial flowering and aquatic plants (*[Ambrosia artemisiifolia](/source/Ambrosia_artemisiifolia)*, *[Ludwigia](/source/Ludwigia_(plant))* sp. and *[Lagarosiphon major](/source/Lagarosiphon_major)*). Over 90% of alien species currently recorded in France had no costs reported in the literature, resulting in high biases in taxonomic, regional and activity sector coverages. However, the lack of reports does not mean there are no negative consequences or costs.[146]

## Favorable effects

See also: [Introduced species](/source/Introduced_species)

The consensus of the [scientific community](/source/Scientific_community) is that the effects of invasive species on [biodiversity](/source/Biodiversity) are primarily negative, despite the potential for some favorable impacts.[147][148][132] The entomologist [Chris D. Thomas](/source/Chris_D._Thomas) argues that most introduced species are neutral or beneficial with respect to other species[149] but this is a minority opinion.[150][151]

Some invasive species can provide a suitable habitat or food source for other organisms. In areas where a native has become extinct or reached a point that it cannot be restored, non-native species can fill their role despite having negative impacts elsewhere. For instance, in the US, the endangered [southwestern willow flycatcher](/source/Willow_flycatcher) (*Empidonax traillii*) mainly nests in the non-native [tamarisk](/source/Tamarix).[132] The introduced [mesquite](/source/Prosopis_juliflora) (*Neltuma juliflora*) is an aggressive invasive species in [India](/source/India), but is the preferred nesting site of native waterbirds in small cities like [Udaipur](/source/Udaipur) in Rajasthan.[152] Similarly, [Ridgway's rail](/source/Ridgway's_rail) (*Rallus obsoletus*) has adapted to the invasive hybrid of *[Spartina alterniflora](/source/Spartina_alterniflora)* and *[Spartina foliosa](/source/Spartina_foliosa)*, which offers better cover and nesting habitat.[153] In [Australia](/source/Australia), [saltwater crocodiles](/source/Saltwater_crocodile) (*Crocodylus porosus*), which had become endangered, have recovered by feeding on introduced [feral pigs](/source/Feral_pig) (*Sus domesticus*).[154]

Non-native species can provide ecosystem services, functioning as [biocontrol](/source/Biocontrol) agents to limit the effects of invasive agricultural pests.[132] Some species have invaded an area so long ago that they are considered to have [naturalised](/source/Naturalisation_(biology)) there. In the US, the endangered [Taylor's checkerspot](/source/Euphydryas_editha_taylori) butterfly (*Euphydryas editha taylori*) has come to rely on invasive [ribwort plantain](/source/Ribwort_plantain) (*Plantago lanceolata*) as the food plant for its caterpillars.[155]

Some invasions offer potential commercial benefits. For instance, [silver carp](/source/Silver_carp) (*Hypophthalmichthys molitrix*) and [common carp](/source/Common_carp) (*Cyprinus carpio*) can be harvested for human food and exported to markets already familiar with the product, or processed into [pet foods](/source/Pet_food) or [mink](/source/Mink) feed. [Water hyacinth](/source/Water_hyacinth) (*Pontederia crassipes*) can be turned into fuel by [methane digesters](/source/Methane_digesters),[156] and other invasive plants can be harvested and utilized as a source of [bioenergy](/source/Bioenergy).[157]

## Control, eradication, and study

Budget to manage invasive alien species

Humans are versatile enough to remediate adverse effects of species invasions.[158][8][159] The public is motivated by invasive species that impact their local area.[160] The control of alien species populations is important in the conservation of biodiversity in natural ecosystem. Previous studies and control programs that target invasives are the most effective during the earlier phases of invasion, such as eliminating [founder population](/source/Founder_effect) quickly before the introduced species can spread.[161] One of the most promising methods for controlling alien species is genetic.[162]

### Cargo inspection and quarantine

The original motivation was to protect against [agricultural pests](/source/Agricultural_pest) while still allowing the export of agricultural products. In 1994 the first set of global standards were agreed to, including the [Agreement on the Application of Sanitary and Phytosanitary Measures](/source/Agreement_on_the_Application_of_Sanitary_and_Phytosanitary_Measures) (SPS Agreement). These are overseen by the [World Trade Organization](/source/World_Trade_Organization). The [International Maritime Organization](/source/International_Maritime_Organization) oversees the International Convention for the Control and Management of Ships' Ballast Water and Sediments (the [Ballast Water Management Convention](/source/Ballast_Water_Management_Convention)). Although primarily targeted at other, more general environmental concerns, the [Convention on Biological Diversity](/source/Convention_on_Biological_Diversity) does specify some steps that its members should take to control invasive species. The CBD is the most significant international agreement on the environmental consequences of invasive species; most such measures are voluntary and unspecific.[163]

### Slowing spread

[Firefighters](/source/Firefighter) are becoming responsible for decontamination of their own equipment, public water equipment, and private water equipment, due to the risk of aquatic invasive species transfer.[164] In the United States this is especially a concern for [wildland firefighters](/source/Wildfire_suppression) because [quagga](/source/Quagga_mussel) (*Dreissena bugensis*) and [zebra](/source/Zebra_mussel) (*Dreissena polymorpha*) mussel invasion and wildfires co-occur in the American West.[165][166][167][168]

### Reestablishing species

[Takahē](/source/Takah%C4%93) (*Porphyrio hochstetteri*) have bred after translocation to [restored islands](/source/Island_restoration), like these on [Kapiti Island](/source/Kapiti_Island), off New Zealand.

[Island restoration](/source/Island_restoration) deals with the eradication of invasive species on islands. A 2019 study suggests that if eradications of invasive animals were conducted on just 169 islands, the survival prospects of 9.4% of the Earth's most highly threatened terrestrial insular vertebrates would be improved.[169]

Invasive vertebrate eradication on islands aligns with United Nations [Sustainable Development Goal 15](/source/Sustainable_Development_Goal_15) and associated targets.[170][171]

[Rodents](/source/Rodent) were carried to [South Georgia](/source/South_Georgia_Island), an island in the southern Atlantic Ocean with no permanent inhabitants, in the 18th century by sealing and whaling ships. They soon wrought havoc on the island's bird population, eating eggs and attacking chicks. In 2018, the South Georgia Island was declared free of invasive rodents after a multi-year extermination effort. Bird populations have rebounded, including the [South Georgia pipit](/source/South_Georgia_pipit) (*Anthus antarcticus*) and [South Georgia pintail](/source/South_Georgia_pintail) (*Anas georgica georgica*), both endemic to the island.[172][173]

### Taxon substitution

The [Aldabra giant tortoise](/source/Aldabra_giant_tortoise) (*Aldabrachelys gigantea*) has helped to restore ecological equilibrium on two islets off Mauritius, including the [Île aux Aigrettes](/source/%C3%8Ele_aux_Aigrettes) (pictured).

Non-native species can be introduced to fill an ecological engineering role that previously was performed by a native species now extinct. The procedure is known as taxon substitution.[132][174][175] On many islands, tortoise extinction has resulted in dysfunctional ecosystems with respect to seed dispersal and herbivory. On the offshore islets of [Mauritius](/source/Mauritius), tortoises now extinct had served as the keystone herbivores. Introduction of the non-native [Aldabra giant tortoises](/source/Aldabra_giant_tortoise) (*Aldabrachelys gigantea*) on two islets in 2000 and 2007 has begun to restore ecological equilibrium. The introduced tortoises are dispersing seeds of several native plants and are selectively grazing invasive plant species. Grazing and browsing are expected to replace ongoing intensive manual weeding, and the introduced tortoises are already breeding.[176]

### By using them as food

Further information: [List of edible invasive species](/source/List_of_edible_invasive_species)

The practice of eating invasive species to reduce their populations has been explored. In 2005, Chef [Bun Lai](/source/Bun_Lai) of [Miya's Sushi](/source/Miya's) in [New Haven, Connecticut](/source/New_Haven%2C_Connecticut), created the first menu dedicated to invasive species. At that time, half the items on the menu were conceptual because those invasive species were not yet commercially available.[177] By 2013, Miya's offered invasive aquatic species such as Chesapeake [blue catfish](/source/Blue_catfish) (*Ictalurus furcatus*), Florida [lionfish](/source/Lionfish) (*Pterois* *sp.*), Kentucky [silver carp](/source/Silver_carp) (*Hypophthalmichthys molitrix)***,** Georgia [cannonball jellyfish](/source/Cannonball_jellyfish) (*Stomolophus meleagris*), and invasive plants such as [Japanese knotweed](/source/Japanese_knotweed) (*Reynoutria japonica*) and [autumn olive](/source/Autumn_olive) (*Elaeagnus umbellata*).[178][179][180][181] [Joe Roman](/source/Joe_Roman), a [Harvard](/source/Harvard) and [University of Vermont](/source/University_of_Vermont) conservation biologist and recipient of the [Rachel Carson](/source/Rachel_Carson) Environmental award, runs a website named "Eat The Invaders".[182][183][177] In the 21st century, organizations including Reef Environmental Educational Foundation and the Institute for Applied Ecology have published cookbooks and recipes using invasive species as ingredients.[184][185] Invasive plant species have been explored as a sustainable source of beneficial phytochemicals and edible protein.[186][187][188]

Proponents of eating invasive organisms argue that humans have the ability to eat away any species that it has an appetite for, pointing to the many animals which humans have been able to hunt to extinction—such as the [Caribbean monk seal](/source/Caribbean_monk_seal) (*Neomonachus tropicalis*) and the [passenger pigeon](/source/Passenger_pigeon) (*Ectopistes migratorius*). They further point to [Jamaica](/source/Jamaica)'s success in significantly decreasing the population of [lionfish](/source/Lionfish) by encouraging the consumption of the fish.[189] Skeptics point out that once a foreign species has entrenched itself in a new place—such as the [Indo-Pacific](/source/Indo-Pacific) lionfish that has now virtually taken over the waters of the western [Atlantic Ocean](/source/Atlantic_Ocean), [Caribbean](/source/Caribbean) and [Gulf of Mexico](/source/Gulf_of_Mexico)—eradication is almost impossible. Critics argue that encouraging consumption might have the unintended effect of spreading harmful species even more widely.[190]

### Pesticides and herbicides

[Pesticides](/source/Pesticide) are commonly used to control invasives.[191] Herbicides used against invasive plants include [fungal herbicides](/source/Fungal_herbicide).[191] Although the [effective population size](/source/Effective_population_size) of an introduced population is [bottlenecked](/source/Population_bottleneck), some [genetic variation](/source/Genetic_variation) has been known to provide invasive plants with [resistance against these fungal bioherbicides](/source/Plant_disease_resistance).[191] Invasive populations of [cheatgrass](/source/Bromus_tectorum) (*Bromus tectorum*) exist with resistance to *[Ustilago bullata](https://en.wikipedia.org/w/index.php?title=Ustilago_bullata&action=edit&redlink=1)* used as a biocontrol, and a similar problem has been reported in [Japanese stiltgrass](/source/Microstegium_vimineum) *(Microstegium vimineum*) subject to *[Bipolaris microstegii](https://en.wikipedia.org/w/index.php?title=Bipolaris_microstegii&action=edit&redlink=1)* and *[B. drechsleri](https://en.wikipedia.org/w/index.php?title=Bipolaris_drechsleri&action=edit&redlink=1)*.[191] This is not solely a character of invasive plant genetics but is normal for wild plants such as the weed [wild flax](/source/Linum_marginale) (*Linum marginale)* and its fungal pathogen [flax rust](/source/Melampsora_lini) (*Melampsora lini)*.[191] Crops have another disadvantage over any uncontrolled plant – wild native or invasive – namely their greater uptake of nutrients, as they are [deliberately bred](/source/Crop_breeding) to increase nutrient intake to enable increased product output.[191]

### Gene drive

A [gene drive](/source/Gene_drive) could be used to eliminate invasive species and has, for example, been proposed as a way to eliminate [invasive mammal species in New Zealand](/source/Invasive_species_in_New_Zealand).[192] Briefly put, an individual of a species may have two versions of a gene, one with a desired coding outcome and one not, with offspring having a 50:50 chance of inheriting one or the other. Genetic engineering can be used to inhibit inheritance of the non-desired gene, resulting in faster propagation of the desired gene in subsequent generations.[193] Gene drives for biodiversity conservation purposes are being explored as part of The Genetic Biocontrol of Invasive Rodents program because they offer the potential for reduced risk to non-target species and reduced costs when compared to traditional invasive species removal techniques.[194] A wider outreach network for gene drive research exists to raise awareness of the value of gene drive research for the public good.[193] Some scientists are concerned that the technique could wipe out species in their original native habitats.[195] The gene could mutate, causing unforeseen problems,[196] or hybridize with native species.[197]

### Predicting invasive plants

Accurately predicting the impacts of non-native plants can be an especially effective management option because most introductions of non-native plant species are intentional.[198][199][200] Weed risk assessments attempt to predict the chances that a specific plant will have negative effects in a new environment, often using a standardized questionnaire. The resulting total score is associated with a management action such as "prevent introduction".[201][202] Assessments commonly use information about the physiology,[201] life history,[202] native ranges,[203] and phylogenetic relationships of the species evaluated. The effectiveness of the approach is debated.[204][205]

### Predicting invasive animals

Invasive alien animal species can seriously affect human well-being and biodiversity. A hierarchical approach underpins the management measures used to lessen these effects, ranging from invasive species management to invasion prevention through early warning and quick response.[206][207][208] Currently, a small number of research on invading mammals have employed spatially explicit models, and the majority of them only looked at a small number of species. The majority of the research employed climate matching to assess the appropriateness of global geographic regions or the potential for established species to spread farther. For species that are not yet established but are anticipated to do so, modelling techniques may be a helpful tool to evaluate the risk of establishment; nevertheless, there aren't many research of this kind for mammals.[206][209]

### Returning invasive species to origin country

In 2025, for the first time in history, a project to return an invasive species to the country it came from has been started in the [Hebrides](/source/Hebrides) in [Scotland](/source/Scotland). [Hedgehogs](/source/Hedgehog) "native to the UK mainland" were brought to those islands decades ago to fight garden pests, but the introduction caused severe harm to birds which nested on the ground. The authorities decided to move them back to the mainland, helping the birds without harming the hedgehogs.[210]

### Government Regulations

Government regulations offer another way to eradicate invasive species. The Florida Fish and Wildlife Commission (FWC) sponsors competitions aimed at removing [lionfish](/source/Lionfish) from [Florida](/source/Florida) waters.[1] The challenge offers divers with prizes contingent on the number of lionfish each competitor successfully removes. Similarly, the FWC encourages local Floridians to collect invasive [green iguanas](/source/Green_iguana) and turn them into local government facilities.[6] When the state undergoes cold snaps, these non-native lizards often freeze and become stunned allowing for the ease of collecting them. The FWC permits the killing of green iguanas year-round to combat the population.[7]

The state of [Texas](/source/Texas) classifies [feral hogs](/source/Feral_pig) as invasive species. As such, [hunting](/source/Hunting) of these hogs is permitted and highly encouraged by the Texas government year-round. Methods described by the Texas State Government include [trapping](/source/Trapping), corralling, aerial gunning (high-powered rifles discharged from individuals aboard helicopters), and [tracking dogs](/source/Tracking_(dog)).[9] Many local governments across Texas also provide bounties that encourage hunters to increase pressure on the feral hog populations.[12]

The state of [Idaho](/source/Idaho) classifies [walleye](/source/Walleye) as an invasive species in their waters. These predatory fish pose a threat to native aquatic populations. Idaho Fish and Game officials strongly encourage [anglers](/source/Fisherman) to harvest caught walleye year-round with no limits regarding amount or size.[13] Similarly, the Indiana Department of Natural Resources classifies thirteen species of fish, including [bighead carp](/source/Bighead_carp) and [walking catfish](/source/Walking_catfish), as invasive to Indiana waters and finds anglers not in violation of the Exotic Fish Rule only if the fish are killed immediately after being caught.[18]

## Criticisms of the invasive species discourse

The process of removing and classifying animals as invasive species is not without controversy. Several [animal welfare movements](/source/Animal_welfare) have attempted to dissuade the eradication of invasive species. A main point of contention for these activists is the treatment of the invasive species during the eradication process. These proponents argue that the stigma surrounding an invasive species seemingly outweighs the notion that all animals are living beings that deserve respect and care.[211] These activists acknowledge that invasive species contribute to more economic and environmental damage than every [natural disaster](/source/Natural_disaster) combined but find the removal process largely [unethical](/source/Ethics) and [cruel](/source/Cruelty).[211]

Other [animal activists](/source/Animal_rights_movement) reject the idea of labeling a species as invasive altogether. These activists posit the labeling of a species as ‘invasive’ constitutes wrongful [discrimination](/source/Discrimination) that permits systemic violence on the creatures.[212] Proponents of this subset of animal activism profess invasive species labeling is inherently arbitrary and a perpetuator of [speciesism](/source/Speciesism), a term coined by [Peter Singer](/source/Peter_Singer) to describe the “misguided belief that one species is more important than another.”[213] Once a species is determined to be legally invasive, those animals often fail to be protected by anti-cruelty statutes and inhumane killings of the species, like [cane toads](/source/Cane_toads_in_Australia) in [Australia](/source/Australia), become commonplace.[212]

A growing contingent of scientists, [ecologists](/source/Ecology), and animal welfare activists caution utilizing the term ‘invasive’ for species migrating to new [ecosystems](/source/Ecosystem). Scientists disclose that [climate change](/source/Climate_change) has begun pushing species such as [white-tailed deer](/source/White-tailed_deer), [armadillos](/source/Armadillo), [lobsters](/source/Lobster), [unicorn snails](/source/Acanthina), and even [maple trees](/source/Maple) to pursue new ecosystems, a migration necessary to survive hotter temperatures.[214] Rather than labeling these species as invasive, this contingent urges these animals should be welcomed as [refugees](/source/Refugee).[214] The term ‘invasive’ compares these species to enemies in war which warps the public perception of the innocent animals’ livelihood and welfare.

## See also

- [Adventive plant](/source/Adventive_plant)

- [Archaeophyte](/source/Archaeophyte)

- [Climate change and invasive species](/source/Climate_change_and_invasive_species)

- [Ecologically based invasive plant management](/source/Ecologically_based_invasive_plant_management)

- [Escaped plant](/source/Escaped_plant)

- [Hemerochory](/source/Hemerochory)

- [Invasion genetics](/source/Invasion_genetics)

- [Lists of invasive species](/source/Lists_of_invasive_species)

- [Naturalisation (biology)](/source/Naturalisation_(biology))

- [Neophyte (botany)](/source/Neophyte_(botany))

- [Seed dispersal](/source/Seed_dispersal)

- [100 of the World's Worst Invasive Alien Species](/source/100_of_the_World's_Worst_Invasive_Alien_Species)

## References

### Attribution

This article incorporates CC-BY-3.0 text from the reference[98]

### Citations

1. ^ [***a***](#cite_ref-:6_1-0) [***b***](#cite_ref-:6_1-1) ["Challenge"](https://myfwc.com/fishing/saltwater/recreational/lionfish/challenge/). *Florida Fish And Wildlife Conservation Commission*. Retrieved April 30, 2026.

1. **[^](#cite_ref-davis_2-0)** Davis, Mark A.; Thompson, Ken (2000). "Eight Ways to be a Colonizer; Two Ways to be an Invader: A Proposed Nomenclature Scheme for Invasion Ecology". *Bulletin of the Ecological Society of America*. **81** (3): 226–230. [JSTOR](/source/JSTOR_(identifier)) [20168448](https://www.jstor.org/stable/20168448).

1. **[^](#cite_ref-SandlundSchei2001_3-0)** Odd Terje Sandlund; Peter Johan Schei; Åslaug Viken (June 30, 2001). [*Invasive Species and Biodiversity Management*](https://books.google.com/books?id=QHgMqnqaW_YC&pg=PA2). Springer Science & Business Media. pp. 2–. [ISBN](/source/ISBN_(identifier)) [978-0-7923-6876-2](https://en.wikipedia.org/wiki/Special:BookSources/978-0-7923-6876-2). [Archived](https://web.archive.org/web/20211218042441/https://books.google.com/books?id=QHgMqnqaW_YC&pg=PA2) from the original on December 18, 2021. Retrieved November 1, 2020.

1. ^ [***a***](#cite_ref-neutral_4-0) [***b***](#cite_ref-neutral_4-1) [***c***](#cite_ref-neutral_4-2) [***d***](#cite_ref-neutral_4-3) [***e***](#cite_ref-neutral_4-4) Colautti, Robert I.; MacIsaac, Hugh J. (February 24, 2004). ["A neutral terminology to define 'invasive' species: Defining invasive species"](https://doi.org/10.1111%2Fj.1366-9516.2004.00061.x). *Diversity and Distributions*. **10** (2): 135–141. [doi](/source/Doi_(identifier)):[10.1111/j.1366-9516.2004.00061.x](https://doi.org/10.1111%2Fj.1366-9516.2004.00061.x). [S2CID](/source/S2CID_(identifier)) [18971654](https://api.semanticscholar.org/CorpusID:18971654).

1. **[^](#cite_ref-Inderjit2006_5-0)** S. Inderjit (January 16, 2006). [*Invasive Plants: Ecological and Agricultural Aspects*](https://books.google.com/books?id=c-aMIwmis-wC&pg=PA252). Springer Science & Business Media. pp. 252–. [ISBN](/source/ISBN_(identifier)) [978-3-7643-7380-1](https://en.wikipedia.org/wiki/Special:BookSources/978-3-7643-7380-1). [Archived](https://web.archive.org/web/20211218042441/https://books.google.com/books?id=c-aMIwmis-wC&pg=PA252) from the original on December 18, 2021. Retrieved November 1, 2020.

1. ^ [***a***](#cite_ref-:7_6-0) [***b***](#cite_ref-:7_6-1) Luscombe, Richard (February 5, 2026). ["Florida euthanizes 5,000 iguanas after cold snap stuns the invasive reptiles"](https://www.theguardian.com/us-news/2026/feb/05/florida-frozen-iguanas-euthanized). *The Guardian*. [ISSN](/source/ISSN_(identifier)) [0261-3077](https://search.worldcat.org/issn/0261-3077). Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-:9_7-0) [***b***](#cite_ref-:9_7-1) ["Green Iguana"](https://myfwc.com/wildlifehabitats/profiles/reptiles/lizards/green-iguana/). *Florida Fish And Wildlife Conservation Commission*. Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-root_8-0) [***b***](#cite_ref-root_8-1) Root-Bernstein, Meredith; Ladle, Richard (2019). ["Ecology of a widespread large omnivore, Homo sapiens, and its impacts on ecosystem processes"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6802023). *[Ecology and Evolution](/source/Ecology_and_Evolution)*. **9** (19): 10874–94. [Bibcode](/source/Bibcode_(identifier)):[2019EcoEv...910874R](https://ui.adsabs.harvard.edu/abs/2019EcoEv...910874R). [doi](/source/Doi_(identifier)):[10.1002/ece3.5049](https://doi.org/10.1002%2Fece3.5049). [PMC](/source/PMC_(identifier)) [6802023](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6802023). [PMID](/source/PMID_(identifier)) [31641442](https://pubmed.ncbi.nlm.nih.gov/31641442). [S2CID](/source/S2CID_(identifier)) [203370925](https://api.semanticscholar.org/CorpusID:203370925).

1. ^ [***a***](#cite_ref-:10_9-0) [***b***](#cite_ref-:10_9-1) Kinsey, John (2020). ["Nuisance Wildlife in Texas: Wild Pigs"](http://tpwd.texas.gov/huntwild/wild/nuisance/feral_hogs/). *tpwd.texas.gov*. Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-IE_10-0) [***b***](#cite_ref-IE_10-1) [***c***](#cite_ref-IE_10-2) Lockwood, Julie L.; Hoopes, Martha F.; Marchetti, Michael P. (2007). [*Invasion Ecology*](https://web.archive.org/web/20150924074044/http://www.planta.cn/forum/files_planta/invasion_ecology1_208.pdf) (PDF). Blackwell Publishing. p. 7. Archived from [the original](http://www.planta.cn/forum/files_planta/invasion_ecology1_208.pdf) (PDF) on September 24, 2015. Retrieved January 21, 2014.

1. ^ [***a***](#cite_ref-Lowry_11-0) [***b***](#cite_ref-Lowry_11-1) [***c***](#cite_ref-Lowry_11-2) [***d***](#cite_ref-Lowry_11-3) Lowry, E; Rollinson, EJ; Laybourn, AJ; Scott, TE; Aiello-Lammens, ME; Gray, SM; Mickley, J; [Gurevitch, J](/source/Jessica_Gurevitch) (2012). ["Biological invasions: A field synopsis, systematic review, and database of the literature"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3568853). *[Ecology and Evolution](/source/Ecology_and_Evolution)*. **3** (1): 182–96. [doi](/source/Doi_(identifier)):[10.1002/ece3.431](https://doi.org/10.1002%2Fece3.431). [PMC](/source/PMC_(identifier)) [3568853](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3568853). [PMID](/source/PMID_(identifier)) [23404636](https://pubmed.ncbi.nlm.nih.gov/23404636).

1. ^ [***a***](#cite_ref-:11_12-0) [***b***](#cite_ref-:11_12-1) ["Nuisance Wildlife in Texas: Wild Pigs"](http://tpwd.texas.gov/huntwild/wild/nuisance/feral_hogs/). *tpwd.texas.gov*. Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-:12_13-0) [***b***](#cite_ref-:12_13-1) ["Fishing Seasons and Rules | Idaho Fish and Game"](https://idfg.idaho.gov/rules/fish). *idfg.idaho.gov*. Retrieved April 30, 2026.

1. **[^](#cite_ref-tilm_14-0)** Tilman, D. (2004). ["Niche tradeoffs, neutrality, and community structure: A stochastic theory of resource competition, invasion, and community assembly"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC503710). *[Proceedings of the National Academy of Sciences](/source/Proceedings_of_the_National_Academy_of_Sciences)*. **101** (30): 10854–10861. [Bibcode](/source/Bibcode_(identifier)):[2004PNAS..10110854T](https://ui.adsabs.harvard.edu/abs/2004PNAS..10110854T). [doi](/source/Doi_(identifier)):[10.1073/pnas.0403458101](https://doi.org/10.1073%2Fpnas.0403458101). [PMC](/source/PMC_(identifier)) [503710](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC503710). [PMID](/source/PMID_(identifier)) [15243158](https://pubmed.ncbi.nlm.nih.gov/15243158).

1. **[^](#cite_ref-verl_15-0)** Verling, E. (2005). ["Supply-side invasion ecology: characterizing propagule pressure in coastal ecosystems"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1564104). *[Proceedings of the Royal Society B](/source/Proceedings_of_the_Royal_Society_B)*. **272** (1569): 1249–1256. [doi](/source/Doi_(identifier)):[10.1098/rspb.2005.3090](https://doi.org/10.1098%2Frspb.2005.3090). [PMC](/source/PMC_(identifier)) [1564104](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1564104). [PMID](/source/PMID_(identifier)) [16024389](https://pubmed.ncbi.nlm.nih.gov/16024389).

1. **[^](#cite_ref-Byers_2002_16-0)** Byers, James E. (June 2002). "Impact of non-indigenous species on natives enhanced by anthropogenic alteration of selection regimes". *Oikos*. **97** (3): 449–458. [Bibcode](/source/Bibcode_(identifier)):[2002Oikos..97..449B](https://ui.adsabs.harvard.edu/abs/2002Oikos..97..449B). [doi](/source/Doi_(identifier)):[10.1034/j.1600-0706.2002.970316.x](https://doi.org/10.1034%2Fj.1600-0706.2002.970316.x).

1. ^ [***a***](#cite_ref-Davis_2000_528–534_17-0) [***b***](#cite_ref-Davis_2000_528–534_17-1) Davis, M.A.; Grime, J.P.; Thompson, K. (2000). ["Fluctuating resources in plant communities: A general theory of invisibility"](https://doi.org/10.1046%2Fj.1365-2745.2000.00473.x). *[Journal of Ecology](/source/Journal_of_Ecology)*. **88** (3): 528–534. [Bibcode](/source/Bibcode_(identifier)):[2000JEcol..88..528D](https://ui.adsabs.harvard.edu/abs/2000JEcol..88..528D). [doi](/source/Doi_(identifier)):[10.1046/j.1365-2745.2000.00473.x](https://doi.org/10.1046%2Fj.1365-2745.2000.00473.x). [S2CID](/source/S2CID_(identifier)) [14573817](https://api.semanticscholar.org/CorpusID:14573817).

1. ^ [***a***](#cite_ref-:13_18-0) [***b***](#cite_ref-:13_18-1) Resources, Indiana Department of Natural (March 9, 2026). ["Indiana Department of Natural Resources"](https://www.in.gov/dnr/). *Indiana Department of Natural Resources*. Retrieved April 30, 2026.

1. **[^](#cite_ref-19)** Alverson, William S.; Waller, Donald M.; Solheim, Stephen L. (1988). "Forests Too Deer: Edge Effects in Northern Wisconsin". *[Conservation Biology](/source/Conservation_Biology_(journal))*. **2** (4): 348–358. [Bibcode](/source/Bibcode_(identifier)):[1988ConBi...2..348A](https://ui.adsabs.harvard.edu/abs/1988ConBi...2..348A). [doi](/source/Doi_(identifier)):[10.1111/j.1523-1739.1988.tb00199.x](https://doi.org/10.1111%2Fj.1523-1739.1988.tb00199.x). [JSTOR](/source/JSTOR_(identifier)) [2386294](https://www.jstor.org/stable/2386294).

1. ^ [***a***](#cite_ref-elton_20-0) [***b***](#cite_ref-elton_20-1) Elton, C.S. (2000) [1958]. *The Ecology of Invasions by Animals and Plants*. Foreword by Daniel Simberloff. Chicago: University of Chicago Press. p. 196. [ISBN](/source/ISBN_(identifier)) [978-0-226-20638-7](https://en.wikipedia.org/wiki/Special:BookSources/978-0-226-20638-7).

1. ^ [***a***](#cite_ref-Schell_21-0) [***b***](#cite_ref-Schell_21-1) Stohlgren, Thomas J.; Binkley, Dan; Chong, Geneva W.; Kalkhan, Mohammed A.; Schell, Lisa D.; Bull, Kelly A.; et al. (February 1999). "Exotic Plant Species Invade Hot Spots of Native Plant Diversity". *Ecological Monographs*. **69** (1): 25–46. [doi](/source/Doi_(identifier)):[10.1890/0012-9615(1999)069\[0025:EPSIHS\]2.0.CO;2](https://doi.org/10.1890%2F0012-9615%281999%29069%5B0025%3AEPSIHS%5D2.0.CO%3B2).

1. **[^](#cite_ref-22)** Byers, James E.; Noonburg, Erik G. (June 2003). "Scale Dependent Effects of Biotic Resistance to Biological Invasion". *Ecology*. **84** (6): 1428–1433. [Bibcode](/source/Bibcode_(identifier)):[2003Ecol...84.1428B](https://ui.adsabs.harvard.edu/abs/2003Ecol...84.1428B). [doi](/source/Doi_(identifier)):[10.1890/02-3131](https://doi.org/10.1890%2F02-3131).

1. **[^](#cite_ref-23)** Levine, Jonathan M. (May 5, 2000). "Species Diversity and Biological Invasions: Relating Local Process to Community Pattern". *Science*. **288** (5467): 852–854. [Bibcode](/source/Bibcode_(identifier)):[2000Sci...288..852L](https://ui.adsabs.harvard.edu/abs/2000Sci...288..852L). [doi](/source/Doi_(identifier)):[10.1126/science.288.5467.852](https://doi.org/10.1126%2Fscience.288.5467.852). [PMID](/source/PMID_(identifier)) [10797006](https://pubmed.ncbi.nlm.nih.gov/10797006).

1. **[^](#cite_ref-ivey2019_24-0)** Ivey, Matthew R.; Colvin, Michael; Strickland, Bronson K.; Lashley, Marcus A. (June 14, 2019). ["Reduced vertebrate diversity independent of spatial scale following feral swine invasions"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6635915). *[Ecology and Evolution](/source/Ecology_and_Evolution)*. **9** (13): 7761–7767. [Bibcode](/source/Bibcode_(identifier)):[2019EcoEv...9.7761I](https://ui.adsabs.harvard.edu/abs/2019EcoEv...9.7761I). [doi](/source/Doi_(identifier)):[10.1002/ece3.5360](https://doi.org/10.1002%2Fece3.5360). [PMC](/source/PMC_(identifier)) [6635915](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6635915). [PMID](/source/PMID_(identifier)) [31346438](https://pubmed.ncbi.nlm.nih.gov/31346438).

1. **[^](#cite_ref-stach_25-0)** Stachowicz, J.J. (2005). "Species invasions and the relationships between species diversity, community saturation, and ecosystem functioning". In D.F. Sax; J.J. Stachowicz; S.D. Gaines (eds.). [*Species Invasions: Insights into Ecology, Evolution, and Biogeography*](https://archive.org/details/speciesinvasions0000unse). Sunderland, Massachusetts: Sinauer Associates. [ISBN](/source/ISBN_(identifier)) [978-0-87893-811-7](https://en.wikipedia.org/wiki/Special:BookSources/978-0-87893-811-7).

1. **[^](#cite_ref-26)** ["Brown Tree Snake"](https://www.invasivespeciesinfo.gov/profile/brown-tree-snake). *USDA National Invasive Species Information Center*. [Archived](https://web.archive.org/web/20190824120114/https://www.invasivespeciesinfo.gov/profile/brown-tree-snake) from the original on August 24, 2019.

1. **[^](#cite_ref-27)** Unit, Biosafety (May 19, 2010). ["Invasive Alien Species"](https://www.cbd.int/island/invasive.shtml#:~:text=Islands,%20such%20as%20Australia,%20New,and%20public%20health%20of%20inhabitants.). *www.cbd.int*. Retrieved March 2, 2026.

1. **[^](#cite_ref-28)** Howe, K. R. (2003). *The Quest for Origins*. Penguin Books. p. 179. [ISBN](/source/ISBN_(identifier)) [0-14-301857-4](https://en.wikipedia.org/wiki/Special:BookSources/0-14-301857-4).

1. **[^](#cite_ref-29)** ["Rat remains help date New Zealand's colonisation"](https://www.newscientist.com/article/mg19826595-200-rat-remains-help-date-new-zealands-colonisation/?ignored=irrelevant). *New Scientist*. June 4, 2008. [Archived](https://web.archive.org/web/20220611120716/https://www.newscientist.com/article/mg19826595-200-rat-remains-help-date-new-zealands-colonisation/?ignored=irrelevant) from the original on June 11, 2022. Retrieved June 23, 2008.

1. **[^](#cite_ref-30)** Goodman, Steven M. (1997). ["The birds of southeastern Madagascar"](https://doi.org/10.5962%2Fbhl.title.3415). *Fieldiana* (87). [doi](/source/Doi_(identifier)):[10.5962/bhl.title.3415](https://doi.org/10.5962%2Fbhl.title.3415).

1. **[^](#cite_ref-31)** Brown, Kerry A.; Gurevitch, Jessica (April 20, 2004). ["Long-term impacts of logging on forest diversity in Madagascar"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC395920). *Proceedings of the National Academy of Sciences*. **101** (16): 6045–6049. [Bibcode](/source/Bibcode_(identifier)):[2004PNAS..101.6045B](https://ui.adsabs.harvard.edu/abs/2004PNAS..101.6045B). [doi](/source/Doi_(identifier)):[10.1073/pnas.0401456101](https://doi.org/10.1073%2Fpnas.0401456101). [PMC](/source/PMC_(identifier)) [395920](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC395920). [PMID](/source/PMID_(identifier)) [15067121](https://pubmed.ncbi.nlm.nih.gov/15067121).

1. **[^](#cite_ref-32)** Kull, Ca; Tassin, J; Carriere, Sm (February 26, 2015). ["Approaching invasive species in Madagascar"](https://doi.org/10.4314%2Fmcd.v9i2.2). *Madagascar Conservation & Development*. **9** (2): 60. [doi](/source/Doi_(identifier)):[10.4314/mcd.v9i2.2](https://doi.org/10.4314%2Fmcd.v9i2.2).

1. **[^](#cite_ref-33)** Villamagna, A. M.; Murphy, B. R. (February 2010). "Ecological and socio-economic impacts of invasive water hyacinth (*Eichhornia crassipes*): a review". *Freshwater Biology*. **55** (2): 282–298. [Bibcode](/source/Bibcode_(identifier)):[2010FrBio..55..282V](https://ui.adsabs.harvard.edu/abs/2010FrBio..55..282V). [doi](/source/Doi_(identifier)):[10.1111/j.1365-2427.2009.02294.x](https://doi.org/10.1111%2Fj.1365-2427.2009.02294.x).

1. **[^](#cite_ref-Rakotoarisoa_365–379_34-0)** Rakotoarisoa, T. F.; Richter, T.; Rakotondramanana, H.; Mantilla-Contreras, J. (December 2016). "Turning a Problem Into Profit: Using Water Hyacinth (*Eichhornia crassipes*) for Making Handicrafts at Lake Alaotra, Madagascar". *Economic Botany*. **70** (4): 365–379. [Bibcode](/source/Bibcode_(identifier)):[2016EcBot..70..365R](https://ui.adsabs.harvard.edu/abs/2016EcBot..70..365R). [doi](/source/Doi_(identifier)):[10.1007/s12231-016-9362-y](https://doi.org/10.1007%2Fs12231-016-9362-y). [S2CID](/source/S2CID_(identifier)) [255557151](https://api.semanticscholar.org/CorpusID:255557151). [S2CID](/source/S2CID_(identifier)) [18820290](https://api.semanticscholar.org/CorpusID:18820290).

1. **[^](#cite_ref-35)** Bhagwat, Shonil A.; Breman, Elinor; Thekaekara, Tarsh; Thornton, Thomas F.; Willis, Katherine J. (2012). ["A Battle Lost? Report on Two Centuries of Invasion and Management of Lantana camara L. in Australia, India and South Africa"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3293794). *PLOS ONE*. **7** (3) e32407. [Bibcode](/source/Bibcode_(identifier)):[2012PLoSO...732407B](https://ui.adsabs.harvard.edu/abs/2012PLoSO...732407B). [doi](/source/Doi_(identifier)):[10.1371/journal.pone.0032407](https://doi.org/10.1371%2Fjournal.pone.0032407). [PMC](/source/PMC_(identifier)) [3293794](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3293794). [PMID](/source/PMID_(identifier)) [22403653](https://pubmed.ncbi.nlm.nih.gov/22403653).

1. **[^](#cite_ref-36)** Mungi, Ninad Avinash; Qureshi, Qamar; Jhala, Yadvendradev V. (2020). ["Expanding niche and degrading forests: Key to the successful global invasion of Lantana camara (sensu lato)"](https://doi.org/10.1016%2Fj.gecco.2020.e01080). *Global Ecology and Conservation*. **23** e01080. [Bibcode](/source/Bibcode_(identifier)):[2020GEcoC..2301080M](https://ui.adsabs.harvard.edu/abs/2020GEcoC..2301080M). [doi](/source/Doi_(identifier)):[10.1016/j.gecco.2020.e01080](https://doi.org/10.1016%2Fj.gecco.2020.e01080).

1. **[^](#cite_ref-37)** Egawa, Chika (2017). ["Wind dispersal of alien plant species into remnant natural vegetation from adjacent agricultural fields"](https://doi.org/10.1016%2Fj.gecco.2017.04.008). *Global Ecology and Conservation*. **11**: 33–41. [Bibcode](/source/Bibcode_(identifier)):[2017GEcoC..11...33E](https://ui.adsabs.harvard.edu/abs/2017GEcoC..11...33E). [doi](/source/Doi_(identifier)):[10.1016/j.gecco.2017.04.008](https://doi.org/10.1016%2Fj.gecco.2017.04.008).

1. **[^](#cite_ref-38)** Treep, Jelle; De Jager, Monique; Bartumeus, Frederic; Soons, Merel B. (2021). ["Seed dispersal as a search strategy: Dynamic and fragmented landscapes select for multi-scale movement strategies in plants"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7845050). *Movement Ecology*. **9** (1) 4. [Bibcode](/source/Bibcode_(identifier)):[2021MvEco...9....4T](https://ui.adsabs.harvard.edu/abs/2021MvEco...9....4T). [doi](/source/Doi_(identifier)):[10.1186/s40462-020-00239-1](https://doi.org/10.1186%2Fs40462-020-00239-1). [PMC](/source/PMC_(identifier)) [7845050](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7845050). [PMID](/source/PMID_(identifier)) [33514441](https://pubmed.ncbi.nlm.nih.gov/33514441).

1. **[^](#cite_ref-39)** Connolly, Trevor A.; Day, Tim D.; King, Carolyn M. (2009). "Estimating the potential for reinvasion by mammalian pests through pest-exclusion fencing". *Wildlife Research*. **36** (5): 410–421. [Bibcode](/source/Bibcode_(identifier)):[2009WildR..36..410C](https://ui.adsabs.harvard.edu/abs/2009WildR..36..410C). [doi](/source/Doi_(identifier)):[10.1071/WR09021](https://doi.org/10.1071%2FWR09021).

1. **[^](#cite_ref-40)** Halassy, Melinda; Krpán, Emese; Csákvári, Edina; Török, Katalin (2025). ["Mowing increases the vulnerability of sand grassland restoration to secondary invasion despite the sensitivity of invasive species to drought"](https://doi.org/10.1016%2Fj.gecco.2025.e03588). *Global Ecology and Conservation*. **60** e03588. [Bibcode](/source/Bibcode_(identifier)):[2025GEcoC..6003588H](https://ui.adsabs.harvard.edu/abs/2025GEcoC..6003588H). [doi](/source/Doi_(identifier)):[10.1016/j.gecco.2025.e03588](https://doi.org/10.1016%2Fj.gecco.2025.e03588).

1. **[^](#cite_ref-41)** Shen, Xiaoli; Bourg, Norman A.; McShea, William J.; Turner, Benjamin L. (2016). ["Long-Term Effects of White-Tailed Deer Exclusion on the Invasion of Exotic Plants: A Case Study in a Mid-Atlantic Temperate Forest"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4809546). *PLOS ONE*. **11** (3) e0151825. [Bibcode](/source/Bibcode_(identifier)):[2016PLoSO..1151825S](https://ui.adsabs.harvard.edu/abs/2016PLoSO..1151825S). [doi](/source/Doi_(identifier)):[10.1371/journal.pone.0151825](https://doi.org/10.1371%2Fjournal.pone.0151825). [PMC](/source/PMC_(identifier)) [4809546](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4809546). [PMID](/source/PMID_(identifier)) [27019356](https://pubmed.ncbi.nlm.nih.gov/27019356).

1. **[^](#cite_ref-42)** Pepe, Maria; Gratani, Loretta; Fabrini, Giuseppe; Varone, Laura (2020). "Seed germination traits of *Ailanthus altissima* , *Phytolacca americana* and *Robinia pseudoacacia* in response to different thermal and light requirements". *Plant Species Biology*. **35** (4): 300–314. [Bibcode](/source/Bibcode_(identifier)):[2020PSBio..35..300P](https://ui.adsabs.harvard.edu/abs/2020PSBio..35..300P). [doi](/source/Doi_(identifier)):[10.1111/1442-1984.12286](https://doi.org/10.1111%2F1442-1984.12286).

1. **[^](#cite_ref-43)** Fei, Songlin; Phillips, Jonathan; Shouse, Michael (November 23, 2014). ["Biogeomorphic Impacts of Invasive Species"](https://doi.org/10.1146%2Fannurev-ecolsys-120213-091928). *Annual Review of Ecology, Evolution, and Systematics*. **45** (1): 69–87. [doi](/source/Doi_(identifier)):[10.1146/annurev-ecolsys-120213-091928](https://doi.org/10.1146%2Fannurev-ecolsys-120213-091928).

1. **[^](#cite_ref-44)** ["Plague of purple sea urchins ravages California's offshore ecosystem, heads to Oregon"](https://www.latimes.com/california/story/2019-10-24/purple-sea-urchins-california-oregon-coasts). *[Los Angeles Times](/source/Los_Angeles_Times)*. October 24, 2019. [Archived](https://web.archive.org/web/20210714030639/https://www.latimes.com/california/story/2019-10-24/purple-sea-urchins-california-oregon-coasts) from the original on July 14, 2021. Retrieved July 14, 2021.

1. ^ [***a***](#cite_ref-kolar_45-0) [***b***](#cite_ref-kolar_45-1) [***c***](#cite_ref-kolar_45-2) Kolar, C.S. (2001). ["Progress in invasion biology: predicting invaders"](https://figshare.com/articles/journal_contribution/24837945). *[Trends in Ecology & Evolution](/source/Trends_in_Ecology_%26_Evolution)*. **16** (4): 199–204. [doi](/source/Doi_(identifier)):[10.1016/S0169-5347(01)02101-2](https://doi.org/10.1016%2FS0169-5347%2801%2902101-2). [PMID](/source/PMID_(identifier)) [11245943](https://pubmed.ncbi.nlm.nih.gov/11245943). [S2CID](/source/S2CID_(identifier)) [5796978](https://api.semanticscholar.org/CorpusID:5796978).

1. **[^](#cite_ref-theb_46-0)** Thebaud, C. (1996). "Assessing why two introduced Conyza differ in their ability to invade Mediterranean old fields". *Ecology*. **77** (3): 791–804. [Bibcode](/source/Bibcode_(identifier)):[1996Ecol...77..791T](https://ui.adsabs.harvard.edu/abs/1996Ecol...77..791T). [doi](/source/Doi_(identifier)):[10.2307/2265502](https://doi.org/10.2307%2F2265502). [JSTOR](/source/JSTOR_(identifier)) [2265502](https://www.jstor.org/stable/2265502).

1. **[^](#cite_ref-reichard_47-0)** Reichard, S.H. (1997). ["Predicting invasions of woody plants introduced into North America"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7162396). *[Conservation Biology](/source/Conservation_Biology_(journal))*. **11** (1): 193–203. [Bibcode](/source/Bibcode_(identifier)):[1997ConBi..11..193R](https://ui.adsabs.harvard.edu/abs/1997ConBi..11..193R). [doi](/source/Doi_(identifier)):[10.1046/j.1523-1739.1997.95473.x](https://doi.org/10.1046%2Fj.1523-1739.1997.95473.x). [PMC](/source/PMC_(identifier)) [7162396](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7162396). [S2CID](/source/S2CID_(identifier)) [29816498](https://api.semanticscholar.org/CorpusID:29816498).

1. **[^](#cite_ref-Williams_48-0)** Williams, J. D. (1998). "Non-indigenous Species". *Status and Trends of the Nation's Biological Resources*. [Reston, Virginia](/source/Reston%2C_Virginia): [United States Geological Survey](/source/United_States_Geological_Survey). pp. 117–29. [ISBN](/source/ISBN_(identifier)) [978-0-16-053285-6](https://en.wikipedia.org/wiki/Special:BookSources/978-0-16-053285-6). [DTIC](/source/DTIC_(identifier)) [ADA368849](https://apps.dtic.mil/sti/citations/tr/ADA368849).

1. **[^](#cite_ref-ewell_49-0)** Ewell, J.J. (1999). ["Deliberate introductions of species: Research needs – Benefits can be reaped, but risks are high"](https://doi.org/10.2307%2F1313438). *[BioScience](/source/BioScience)*. **49** (8): 619–630. [Bibcode](/source/Bibcode_(identifier)):[1999BiSci..49..619E](https://ui.adsabs.harvard.edu/abs/1999BiSci..49..619E). [doi](/source/Doi_(identifier)):[10.2307/1313438](https://doi.org/10.2307%2F1313438). [JSTOR](/source/JSTOR_(identifier)) [1313438](https://www.jstor.org/stable/1313438).

1. **[^](#cite_ref-Cove_Gardner_2018_50-0)** Cove, Michael V.; Gardner, Beth; Simons, Theodore R.; Kays, Roland; O'Connell, Allan F. (February 1, 2018). "Free-ranging domestic cats (*Felis catus*) on public lands: estimating density, activity, and diet in the Florida Keys". *[Biological Invasions](/source/Biological_Invasions)*. **20** (2): 333–344. [Bibcode](/source/Bibcode_(identifier)):[2018BiInv..20..333C](https://ui.adsabs.harvard.edu/abs/2018BiInv..20..333C). [doi](/source/Doi_(identifier)):[10.1007/s10530-017-1534-x](https://doi.org/10.1007%2Fs10530-017-1534-x). [S2CID](/source/S2CID_(identifier)) [3536174](https://api.semanticscholar.org/CorpusID:3536174).

1. **[^](#cite_ref-sax_51-0)** Sax, Dov F.; Gaines, Steven D.; Brown, James H. (December 2002). "Species Invasions Exceed Extinctions on Islands Worldwide: A Comparative Study of Plants and Birds". *The American Naturalist*. **160** (6): 766–783. [Bibcode](/source/Bibcode_(identifier)):[2002ANat..160..766S](https://ui.adsabs.harvard.edu/abs/2002ANat..160..766S). [doi](/source/Doi_(identifier)):[10.1086/343877](https://doi.org/10.1086%2F343877). [PMID](/source/PMID_(identifier)) [18707464](https://pubmed.ncbi.nlm.nih.gov/18707464). [S2CID](/source/S2CID_(identifier)) [8628360](https://api.semanticscholar.org/CorpusID:8628360).

1. **[^](#cite_ref-52)** Huenneke, Laura Foster; Hamburg, Steven P.; Koide, Roger; Mooney, Harold A.; Vitousek, Peter M. (1990). "Effects of Soil Resources on Plant Invasion and Community Structure in Californian Serpentine Grassland". *Ecology*. **71** (2): 478–491. [Bibcode](/source/Bibcode_(identifier)):[1990Ecol...71..478H](https://ui.adsabs.harvard.edu/abs/1990Ecol...71..478H). [doi](/source/Doi_(identifier)):[10.2307/1940302](https://doi.org/10.2307%2F1940302). [JSTOR](/source/JSTOR_(identifier)) [1940302](https://www.jstor.org/stable/1940302).

1. **[^](#cite_ref-HERRERAFERRER-PARIS2018_53-0)** Herrera, Ileana; Ferrer-Paris, José R.; Benzo, Diana; Flores, Saúl; García, Belkis; Nassar, Jafet M. (2018). ["An Invasive Succulent Plant (Kalanchoe daigremontiana) Influences Soil Carbon and Nitrogen Mineralization in a Neotropical Semiarid Zone"](https://unsworks.unsw.edu.au/bitstreams/12acbbad-8af9-444b-a478-e7dc9d941d9b/download). *[Pedosphere](/source/Pedosphere)*. **28** (4): 632–643. [Bibcode](/source/Bibcode_(identifier)):[2018Pedos..28..632H](https://ui.adsabs.harvard.edu/abs/2018Pedos..28..632H). [doi](/source/Doi_(identifier)):[10.1016/S1002-0160(18)60029-3](https://doi.org/10.1016%2FS1002-0160%2818%2960029-3). [hdl](/source/Hdl_(identifier)):[1959.4/unsworks_64013](https://hdl.handle.net/1959.4%2Funsworks_64013). [S2CID](/source/S2CID_(identifier)) [104843296](https://api.semanticscholar.org/CorpusID:104843296).

1. **[^](#cite_ref-54)** Herrera, Ileana; Ferrer-Paris, José R.; Hernández-Rosas, José I.; Nassar, Jafet M. (2016). "Impact of two invasive succulents on native-seedling recruitment in Neotropical arid environments". *[Journal of Arid Environments](/source/Journal_of_Arid_Environments)*. **132**: 15–25. [Bibcode](/source/Bibcode_(identifier)):[2016JArEn.132...15H](https://ui.adsabs.harvard.edu/abs/2016JArEn.132...15H). [doi](/source/Doi_(identifier)):[10.1016/j.jaridenv.2016.04.007](https://doi.org/10.1016%2Fj.jaridenv.2016.04.007).

1. ^ [***a***](#cite_ref-Brooks_2004_677–688_55-0) [***b***](#cite_ref-Brooks_2004_677–688_55-1) Brooks, Matthew L.; D'Antonio, Carla M.; Richardson, David M.; Grace, James B.; Keeley, Jon E.; DiTOMASO, Joseph M.; Hobbs, Richard J.; Pellant, Mike; Pyke, David (2004). ["Effects of Invasive Alien Plants on Fire Regimes"](https://doi.org/10.1641%2F0006-3568%282004%29054%5B0677%3AEOIAPO%5D2.0.CO%3B2). *BioScience*. **54** (7): 677. [doi](/source/Doi_(identifier)):[10.1641/0006-3568(2004)054\[0677:EOIAPO\]2.0.CO;2](https://doi.org/10.1641%2F0006-3568%282004%29054%5B0677%3AEOIAPO%5D2.0.CO%3B2). [S2CID](/source/S2CID_(identifier)) [13769125](https://api.semanticscholar.org/CorpusID:13769125).

1. **[^](#cite_ref-silv_56-0)** Silver Botts, P.; Patterson, B.A.; Schlosser, D. (1996). "Zebra mussel effects on benthic invertebrates: Physical or biotic?". *[Journal of the North American Benthological Society](/source/Journal_of_the_North_American_Benthological_Society)*. **15** (2): 179–184. [Bibcode](/source/Bibcode_(identifier)):[1996JNABS..15..179B](https://ui.adsabs.harvard.edu/abs/1996JNABS..15..179B). [doi](/source/Doi_(identifier)):[10.2307/1467947](https://doi.org/10.2307%2F1467947). [JSTOR](/source/JSTOR_(identifier)) [1467947](https://www.jstor.org/stable/1467947). [S2CID](/source/S2CID_(identifier)) [84660670](https://api.semanticscholar.org/CorpusID:84660670).

1. **[^](#cite_ref-57)** Keddy, Paul A. (2017). [*Plant Ecology*](https://books.google.com/books?id=ncloDgAAQBAJ&q=Plant+Ecology). Cambridge University Press. p. 343. [ISBN](/source/ISBN_(identifier)) [978-1-107-11423-4](https://en.wikipedia.org/wiki/Special:BookSources/978-1-107-11423-4). [Archived](https://web.archive.org/web/20210816224645/https://books.google.com/books?id=ncloDgAAQBAJ&q=Plant+Ecology) from the original on August 16, 2021. Retrieved October 6, 2020.

1. **[^](#cite_ref-58)** Xu, Cheng-Yuan; Tang, Shaoqing; Fatemi, Mohammad; Gross, Caroline L.; Julien, Mic H.; Curtis, Caitlin; van Klinken, Rieks D. (September 1, 2015). ["Population structure and genetic diversity of invasive Phyla canescens: implications for the evolutionary potential"](https://doi.org/10.1890%2FES14-00374.1). *[Ecosphere](/source/Ecosphere_(journal))*. **6** (9): art162. [Bibcode](/source/Bibcode_(identifier)):[2015Ecosp...6..374X](https://ui.adsabs.harvard.edu/abs/2015Ecosp...6..374X). [doi](/source/Doi_(identifier)):[10.1890/ES14-00374.1](https://doi.org/10.1890%2FES14-00374.1).

1. **[^](#cite_ref-Prentis_2008_288-294_59-0)** Prentis, Peter (2008). "Adaptive evolution in invasive species". *[Trends in Plant Science](/source/Trends_in_Plant_Science)*. **13** (6): 288–294. [Bibcode](/source/Bibcode_(identifier)):[2008TPS....13..288P](https://ui.adsabs.harvard.edu/abs/2008TPS....13..288P). [doi](/source/Doi_(identifier)):[10.1016/j.tplants.2008.03.004](https://doi.org/10.1016%2Fj.tplants.2008.03.004). [hdl](/source/Hdl_(identifier)):[10019.1/112332](https://hdl.handle.net/10019.1%2F112332). [PMID](/source/PMID_(identifier)) [18467157](https://pubmed.ncbi.nlm.nih.gov/18467157).

1. **[^](#cite_ref-Eunmi_2002_386-391_60-0)** Lee, Carol Eunmi (2002). "Evolutionary genetics of invasive species". *[Trends in Ecology & Evolution](/source/Trends_in_Ecology_%26_Evolution)*. **17** (8): 386–391. [doi](/source/Doi_(identifier)):[10.1016/s0169-5347(02)02554-5](https://doi.org/10.1016%2Fs0169-5347%2802%2902554-5).

1. **[^](#cite_ref-Zenni_2013_635-644_61-0)** Zenni, R.D. (2013). "Adaptive Evolution and Phenotypic Plasticity During Naturalization and Spread of Invasive Species: Implications for Tree Invasion Biology". *[Biological Invasions](/source/Biological_Invasions)*. **16** (3): 635–644. [doi](/source/Doi_(identifier)):[10.1007/s10530-013-0607-8](https://doi.org/10.1007%2Fs10530-013-0607-8). [S2CID](/source/S2CID_(identifier)) [82590](https://api.semanticscholar.org/CorpusID:82590).

1. ^ [***a***](#cite_ref-amstutz2018_62-0) [***b***](#cite_ref-amstutz2018_62-1) Amstutz, Lisa J (2018). *Invasive Species*. Minneapolis, MN: Abdo Publishing. pp. 8–10. [ISBN](/source/ISBN_(identifier)) [978-1-5321-1024-5](https://en.wikipedia.org/wiki/Special:BookSources/978-1-5321-1024-5).

1. **[^](#cite_ref-63)** Cassey, P (2005). "Concerning Invasive Species: Reply to Brown and Sax". *[Austral Ecology](/source/Austral_Ecology)*. **30** (4): 475–480. [Bibcode](/source/Bibcode_(identifier)):[2005AusEc..30..475C](https://ui.adsabs.harvard.edu/abs/2005AusEc..30..475C). [doi](/source/Doi_(identifier)):[10.1111/j.1442-9993.2005.01505.x](https://doi.org/10.1111%2Fj.1442-9993.2005.01505.x). [hdl](/source/Hdl_(identifier)):[10019.1/119884](https://hdl.handle.net/10019.1%2F119884).

1. **[^](#cite_ref-64)** Matisoo-Smith, E. (1998). ["Patterns of prehistoric human mobility in Polynesia indicated by mtDNA from the Pacific rat"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC24590). *[Proceedings of the National Academy of Sciences of the United States of America](/source/Proceedings_of_the_National_Academy_of_Sciences_of_the_United_States_of_America)*. **95** (25): 15145–15150. [Bibcode](/source/Bibcode_(identifier)):[1998PNAS...9515145M](https://ui.adsabs.harvard.edu/abs/1998PNAS...9515145M). [doi](/source/Doi_(identifier)):[10.1073/pnas.95.25.15145](https://doi.org/10.1073%2Fpnas.95.25.15145). [PMC](/source/PMC_(identifier)) [24590](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC24590). [PMID](/source/PMID_(identifier)) [9844030](https://pubmed.ncbi.nlm.nih.gov/9844030).

1. **[^](#cite_ref-65)** ["Going Forth and Multiplying: Animal Acclimatization and Invasion"](https://dspace.mit.edu/bitstream/handle/1721.1/72380/goingforth.ehversion.dec11.pdf?isAllowed=y&sequence=1) (PDF). *dspace.mit.edu*.

1. **[^](#cite_ref-66)** ["The cane toad (bufo marinus)"](https://web.archive.org/web/20221008121115/https://www.dcceew.gov.au/sites/default/files/documents/cane-toad-fs.pdf) (PDF). Archived from [the original](https://www.dcceew.gov.au/sites/default/files/documents/cane-toad-fs.pdf) (PDF) on October 8, 2022.

1. **[^](#cite_ref-67)** Fineran, Katie E. "1.5 Non-Native Mongoose (Herpestes javanicus): A Threat to Hawaiian Islands". In Lower, Brian H.; Shaul, Travis R.; Shaul, Kylienne A.; Weaver, Ella M. (eds.). [*Environmental ScienceBites Volume 2*](https://ohiostate.pressbooks.pub/sciencebitesvolume2/chapter/1-5-non-native-mongoose-herpestes-javanicus-a-threat-to-hawaiian-islands/). Retrieved January 14, 2026.

1. **[^](#cite_ref-Essl-et-al-2020_68-0)** Essl, Franz; Lenzner, Bernd; Bacher, Sven; Bailey, Sarah; Capinha, Cesar; Daehler, Curtis; et al. (September 2020). ["Drivers of future alien species impacts: An expert-based assessment"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7496498). *Global Change Biology*. **26** (9): 4880–4893. [Bibcode](/source/Bibcode_(identifier)):[2020GCBio..26.4880E](https://ui.adsabs.harvard.edu/abs/2020GCBio..26.4880E). [doi](/source/Doi_(identifier)):[10.1111/gcb.15199](https://doi.org/10.1111%2Fgcb.15199). [PMC](/source/PMC_(identifier)) [7496498](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7496498). [PMID](/source/PMID_(identifier)) [32663906](https://pubmed.ncbi.nlm.nih.gov/32663906).

1. ^ [***a***](#cite_ref-r2_69-0) [***b***](#cite_ref-r2_69-1) ["Citrus Greening"](http://www.clemson.edu/public/regulatory/plant_industry/invasive_exotic_programs/Pest%20Alerts/citrus_greening.html). *Clemson Public Service Activities - The Department of Plant Industry*.{{[cite web](https://en.wikipedia.org/wiki/Template:Cite_web)}}: CS1 maint: deprecated archival service ([link](https://en.wikipedia.org/wiki/Category:CS1_maint:_deprecated_archival_service))

1. **[^](#cite_ref-70)** Leung, B. (2007). ["The risk of establishment of aquatic invasive species: joining invasibility and propagule pressure"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2275890). *[Proceedings of the Royal Society B](/source/Proceedings_of_the_Royal_Society_B)*. **274** (1625): 2733–2739. [doi](/source/Doi_(identifier)):[10.1098/rspb.2007.0841](https://doi.org/10.1098%2Frspb.2007.0841). [PMC](/source/PMC_(identifier)) [2275890](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2275890). [PMID](/source/PMID_(identifier)) [17711834](https://pubmed.ncbi.nlm.nih.gov/17711834).

1. **[^](#cite_ref-:17_71-0)** ["Recreation & Travel: How Outdoor Activities Spread Invasive Species"](https://rewildingreport.com/invasive-species-recreation-pathway). *rewildingreport.com*. Retrieved May 22, 2026.

1. **[^](#cite_ref-72)** Zavaleta, Erika S.; Hobbs, Richard J.; Mooney, Harold A. (August 2001). "Viewing invasive species removal in a whole-ecosystem context". *Trends in Ecology & Evolution*. **16** (8): 454–459. [doi](/source/Doi_(identifier)):[10.1016/s0169-5347(01)02194-2](https://doi.org/10.1016%2Fs0169-5347%2801%2902194-2).

1. **[^](#cite_ref-73)** Seinfeld, John H. (2016). Arias, Andres Hugo; Marcovecchio, Jorge Eduardo (eds.). *Marine Pollution and Climate Change*. [John Wiley & Sons](/source/John_Wiley_%26_Sons). [ISBN](/source/ISBN_(identifier)) [978-1-4822-9944-1](https://en.wikipedia.org/wiki/Special:BookSources/978-1-4822-9944-1).

1. **[^](#cite_ref-74)** Molnar, Jennifer L.; Gamboa, Rebecca L.; Revenga, Carmen; Spalding, Mark D. (November 2008). "Assessing the global threat of invasive species to marine biodiversity". *Frontiers in Ecology and the Environment*. **6** (9): 485–492. [Bibcode](/source/Bibcode_(identifier)):[2008FrEE....6..485M](https://ui.adsabs.harvard.edu/abs/2008FrEE....6..485M). [doi](/source/Doi_(identifier)):[10.1890/070064](https://doi.org/10.1890%2F070064).

1. **[^](#cite_ref-75)** Drake, John (2007). ["Hull fouling is a risk factor for intercontinental species exchange in aquatic ecosystems"](https://doi.org/10.3391%2Fai.2007.2.2.7). *[Aquatic Invasions](https://en.wikipedia.org/w/index.php?title=Aquatic_Invasions&action=edit&redlink=1)*. **2** (2): 121–131. [doi](/source/Doi_(identifier)):[10.3391/ai.2007.2.2.7](https://doi.org/10.3391%2Fai.2007.2.2.7).

1. **[^](#cite_ref-76)** ["Biofouling moves up the regulatory agenda – GARD"](https://www.gard.no/web/updates/content/24305557/biofouling-moves-up-the-regulatory-agenda.). *www.gard.no*. [Archived](https://web.archive.org/web/20200113231807/http://www.gard.no/web/updates/content/24305557/biofouling-moves-up-the-regulatory-agenda.) from the original on January 13, 2020. Retrieved September 19, 2018.

1. ^ [***a***](#cite_ref-cargo_77-0) [***b***](#cite_ref-cargo_77-1) Egan, Dan (October 31, 2005). ["Noxious cargo"](https://web.archive.org/web/20111021122316/http://www.jsonline.com/news/wisconsin/98880204.html). *[Journal Sentinel](/source/Journal_Sentinel)*. Archived from [the original](http://www.jsonline.com/news/wisconsin/98880204.html) on October 21, 2011. Retrieved April 22, 2017.

1. **[^](#cite_ref-78)** Xu, Jian; Wickramarathne, Thanuka L.; Chawla, Nitesh V.; Grey, Erin K.; Steinhaeuser, Karsten; Keller, Reuben P.; Drake, John M.; Lodge, David M. (2014). "Improving management of aquatic invasions by integrating shipping network, ecological, and environmental data". *Proceedings of the 20th ACM SIGKDD international conference on Knowledge discovery and data mining*. pp. 1699–1708. [doi](/source/Doi_(identifier)):[10.1145/2623330.2623364](https://doi.org/10.1145%2F2623330.2623364). [ISBN](/source/ISBN_(identifier)) [978-1-4503-2956-9](https://en.wikipedia.org/wiki/Special:BookSources/978-1-4503-2956-9). [S2CID](/source/S2CID_(identifier)) [2371978](https://api.semanticscholar.org/CorpusID:2371978).

1. **[^](#cite_ref-79)** Streftaris, N; Zenetos, Argyro; Papathanassiou, Enangelos (2005). ["Globalisation in marine ecosystems: The story of non-indigenous marine species across European seas"](https://www.researchgate.net/publication/253862066). *[Oceanography and Marine Biology](/source/Oceanography_and_Marine_Biology)*. **43**: 419–453. [Archived](https://web.archive.org/web/20180920011308/https://www.researchgate.net/publication/253862066) from the original on September 20, 2018. Retrieved September 19, 2018.

1. ^ [***a***](#cite_ref-pnwaquaticinv_80-0) [***b***](#cite_ref-pnwaquaticinv_80-1) Aquatic invasive species. A Guide to Least-Wanted Aquatic Organisms of the Pacific Northwest. 2001. University of Washington

1. **[^](#cite_ref-Bespalaya2026_81-0)** Bespalaya YV, Kropotin AV, Aksenova OV, Gofarov MY, Kondakov AV, Konopleva ES, Khrebtova IS, Mabrouki Y, Palatov DM, Prokin AA, Karabanov DP, Travina OV, Taybi AF, Izzatulaev ZI, Boymurodov KT, Tahirova EN, Snegovaya NY, Tien TV, Vikhrev IV, Vinarski MV, Kyi CT, Chan N, Lunn Z, Unnikrishnan SK, Pattnaik S, Raut D, Subba Rao NV, Bolotov IN (2026). ["Global Molecular Taxonomy, Phylogeny and Biogeography of the Clam Genus Corbicula (Bivalvia: Cyrenidae)"](https://onlinelibrary.wiley.com/doi/10.1111/zsc.70022). *Zoologica Scripta*. **55** (1): 74–101. [doi](/source/Doi_(identifier)):[10.1111/zsc.70022](https://doi.org/10.1111%2Fzsc.70022).

1. **[^](#cite_ref-82)** Great Lake Commission. ["Status of Ballast Water Discharge Regulations in the Great Lakes Region"](https://www.glc.org/wp-content/uploads/GLC-BW-Reg-Summary-11.14.16.pdf) (PDF). [Archived](https://web.archive.org/web/20200212212419/http://www.glc.org/wp-content/uploads/GLC-BW-Reg-Summary-11.14.16.pdf) (PDF) from the original on February 12, 2020. Retrieved September 19, 2018.

1. **[^](#cite_ref-83)** USCG. ["Ballast Water Management for Control of Non-Indigenous Species in Waters of the United States"](https://www.dco.uscg.mil/Portals/9/DCO%20Documents/5p/5ps/NVIC/2018/NVIC-01_18.pdf) (PDF). [Archived](https://web.archive.org/web/20200511133624/https://www.dco.uscg.mil/Portals/9/DCO%20Documents/5p/5ps/NVIC/2018/NVIC-01_18.pdf) (PDF) from the original on May 11, 2020. Retrieved September 19, 2018.

1. **[^](#cite_ref-trainer2012_84-0)** Trainer, Vera L.; Bates, Stephen S.; Lundholm, Nina; Thessen, Anne E.; Cochlan, William P.; Adams, Nicolaus G.; Trick, Charles G. (2012). "Pseudo-nitzschia physiological ecology, phylogeny, toxicity, monitoring and impacts on ecosystem health". *Harmful Algae*. **14**: 271–300. [Bibcode](/source/Bibcode_(identifier)):[2012HAlga..14..271T](https://ui.adsabs.harvard.edu/abs/2012HAlga..14..271T). [doi](/source/Doi_(identifier)):[10.1016/j.hal.2011.10.025](https://doi.org/10.1016%2Fj.hal.2011.10.025). [hdl](/source/Hdl_(identifier)):[1912/5118](https://hdl.handle.net/1912%2F5118).

1. **[^](#cite_ref-85)** Occhipinti-Ambrogi, Anna (2007). "Global change and marine communities: Alien species and climate change". *[Marine Pollution Bulletin](/source/Marine_Pollution_Bulletin)*. **55** (7–9): 342–352. [Bibcode](/source/Bibcode_(identifier)):[2007MarPB..55..342O](https://ui.adsabs.harvard.edu/abs/2007MarPB..55..342O). [doi](/source/Doi_(identifier)):[10.1016/j.marpolbul.2006.11.014](https://doi.org/10.1016%2Fj.marpolbul.2006.11.014). [PMID](/source/PMID_(identifier)) [17239404](https://pubmed.ncbi.nlm.nih.gov/17239404).

1. **[^](#cite_ref-86)** Rahel, Frank J.; Olden, Julian D. (June 2008). ["Assessing the Effects of Climate Change on Aquatic Invasive Species"](https://doi.org/10.1111%2Fj.1523-1739.2008.00950.x). *Conservation Biology*. **22** (3): 521–533. [Bibcode](/source/Bibcode_(identifier)):[2008ConBi..22..521R](https://ui.adsabs.harvard.edu/abs/2008ConBi..22..521R). [doi](/source/Doi_(identifier)):[10.1111/j.1523-1739.2008.00950.x](https://doi.org/10.1111%2Fj.1523-1739.2008.00950.x). [PMID](/source/PMID_(identifier)) [18577081](https://pubmed.ncbi.nlm.nih.gov/18577081). [S2CID](/source/S2CID_(identifier)) [313824](https://api.semanticscholar.org/CorpusID:313824).

1. **[^](#cite_ref-87)** Hua, J.; Hwang, W.H. (2012). "Effects of voyage routing on the survival of microbes in ballast water". *[Ocean Engineering](/source/Ocean_Engineering)*. **42**: 165–175. [Bibcode](/source/Bibcode_(identifier)):[2012OcEng..42..165H](https://ui.adsabs.harvard.edu/abs/2012OcEng..42..165H). [doi](/source/Doi_(identifier)):[10.1016/j.oceaneng.2012.01.013](https://doi.org/10.1016%2Fj.oceaneng.2012.01.013).

1. **[^](#cite_ref-88)** Lenz, Mark; Ahmed, Yasser; Canning-Clode, João; Díaz, Eliecer; Eichhorn, Sandra; Fabritzek, Armin G.; da Gama, Bernardo A. P.; Garcia, Marie; von Juterzenka, Karen (May 24, 2018). "Heat challenges can enhance population tolerance to thermal stress in mussels: a potential mechanism by which ship transport can increase species invasiveness". *[Biological Invasions](/source/Biological_Invasions)*. **20** (11): 3107–3122. [Bibcode](/source/Bibcode_(identifier)):[2018BiInv..20.3107L](https://ui.adsabs.harvard.edu/abs/2018BiInv..20.3107L). [doi](/source/Doi_(identifier)):[10.1007/s10530-018-1762-8](https://doi.org/10.1007%2Fs10530-018-1762-8). [S2CID](/source/S2CID_(identifier)) [53082967](https://api.semanticscholar.org/CorpusID:53082967).

1. **[^](#cite_ref-ehrenfeld2010_89-0)** Ehrenfeld, Joan G. (December 1, 2010). "Ecosystem Consequences of Biological Invasions". *Annual Review of Ecology, Evolution, and Systematics*. **41** (1): 59–80. [doi](/source/Doi_(identifier)):[10.1146/annurev-ecolsys-102209-144650](https://doi.org/10.1146%2Fannurev-ecolsys-102209-144650).

1. **[^](#cite_ref-90)** Simberloff, Daniel (2013). *Invasive species: what everyone needs to know*. Oxford: Oxford University Press. [ISBN](/source/ISBN_(identifier)) [978-0-19-992201-7](https://en.wikipedia.org/wiki/Special:BookSources/978-0-19-992201-7).

1. **[^](#cite_ref-91)** ["Environmental and Ecological Impacts | National Invasive Species Information Center"](https://www.invasivespeciesinfo.gov/subject/environmental-and-ecological-impacts). *www.invasivespeciesinfo.gov*. Retrieved December 11, 2025.

1. **[^](#cite_ref-92)** ["Communication From The Commission To The Council, The European Parliament, The European Economic And Social Committee And The Committee Of The Regions Towards An EU Strategy On Invasive Species"](http://ec.europa.eu/environment/nature/invasivealien/docs/1_EN_resume_impact_assesment_part1_v3.pdf) (PDF). [Archived](https://web.archive.org/web/20160305033628/http://ec.europa.eu/environment/nature/invasivealien/docs/1_EN_resume_impact_assesment_part1_v3.pdf) (PDF) from the original on March 5, 2016. Retrieved May 17, 2011.

1. **[^](#cite_ref-93)** Lakicevic, Milena; Mladenovic, Emina (2018). ["Non-native and invasive tree species - their impact on biodiversity loss"](https://doi.org/10.2298%2FZMSPN1834019L). *[Zbornik Matice Srpske za Prirodne Nauke](https://en.wikipedia.org/w/index.php?title=Zbornik_Matice_Srpske_za_Prirodne_Nauke&action=edit&redlink=1)* (134): 19–26. [doi](/source/Doi_(identifier)):[10.2298/ZMSPN1834019L](https://doi.org/10.2298%2FZMSPN1834019L).

1. **[^](#cite_ref-94)** National Research Council (US) Committee on the Scientific Basis for Predicting the Invasive Potential of Nonindigenous Plants Plant Pests in the United States (2002). [*Predicting Invasions of Nonindigenous Plants and Plant Pests*](https://www.nap.edu/read/10259/chapter/1). [Bibcode](/source/Bibcode_(identifier)):[2002nap..book10259N](https://ui.adsabs.harvard.edu/abs/2002nap..book10259N). [doi](/source/Doi_(identifier)):[10.17226/10259](https://doi.org/10.17226%2F10259). [ISBN](/source/ISBN_(identifier)) [978-0-309-08264-8](https://en.wikipedia.org/wiki/Special:BookSources/978-0-309-08264-8). [PMID](/source/PMID_(identifier)) [25032288](https://pubmed.ncbi.nlm.nih.gov/25032288). [Archived](https://web.archive.org/web/20191117204337/https://www.nap.edu/read/10259/chapter/1) from the original on November 17, 2019. Retrieved November 17, 2019.

1. **[^](#cite_ref-95)** Lewis, Simon L.; Maslin, Mark A. (2015). ["Defining the Anthropocene"](https://zotero.org/groups/5435545/items/CF2PI4C2). *[Nature](/source/Nature_(journal))*. **519** (7542): 171–180. [Bibcode](/source/Bibcode_(identifier)):[2015Natur.519..171L](https://ui.adsabs.harvard.edu/abs/2015Natur.519..171L). [doi](/source/Doi_(identifier)):[10.1038/nature14258](https://doi.org/10.1038%2Fnature14258). [PMID](/source/PMID_(identifier)) [25762280](https://pubmed.ncbi.nlm.nih.gov/25762280). [S2CID](/source/S2CID_(identifier)) [205242896](https://api.semanticscholar.org/CorpusID:205242896).

1. **[^](#cite_ref-96)** [Millennium Ecosystem Assessment](/source/Millennium_Ecosystem_Assessment) (2005). ["Ecosystems and Human Well-being: Biodiversity Synthesis"](http://www.millenniumassessment.org/documents/document.354.aspx.pdf) (PDF). [World Resources Institute](/source/World_Resources_Institute). [Archived](https://web.archive.org/web/20191014033601/http://www.millenniumassessment.org/documents/document.354.aspx.pdf) (PDF) from the original on October 14, 2019. Retrieved September 18, 2007.

1. **[^](#cite_ref-97)** Baiser, Benjamin; Olden, Julian D.; Record, Sydne; Lockwood, Julie L.; McKinney, Michael L. (2012). ["Pattern and process of biotic homogenization in the New Pangaea"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3497087). *[Proceedings of the Royal Society B: Biological Sciences](/source/Proceedings_of_the_Royal_Society_B%3A_Biological_Sciences)*. **279** (1748): 4772–4777. [doi](/source/Doi_(identifier)):[10.1098/rspb.2012.1651](https://doi.org/10.1098%2Frspb.2012.1651). [PMC](/source/PMC_(identifier)) [3497087](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3497087). [PMID](/source/PMID_(identifier)) [23055062](https://pubmed.ncbi.nlm.nih.gov/23055062).

1. ^ [***a***](#cite_ref-Odendaal_2008_98-0) [***b***](#cite_ref-Odendaal_2008_98-1) Odendaal, L. J.; Haupt, T. M.; Griffiths, C. L. (2008). ["The alien invasive land snail *Theba pisana* in the West Coast National Park: Is there cause for concern?"](https://doi.org/10.4102%2Fkoedoe.v50i1.153). *[Koedoe](/source/Koedoe)*. **50** (1): 93–98. [doi](/source/Doi_(identifier)):[10.4102/koedoe.v50i1.153](https://doi.org/10.4102%2Fkoedoe.v50i1.153).

1. **[^](#cite_ref-99)** Fisher, Matthew C.; Garner, Trenton W. J. (2020). ["Chytrid fungi and global amphibian declines"](https://discovery.ucl.ac.uk/id/eprint/10092667/1/NRMICRO-19-244_FINAL_ACCEPTED.pdf) (PDF). *[Nature Reviews Microbiology](/source/Nature_Reviews_Microbiology)*. **18** (6): 332–343. [doi](/source/Doi_(identifier)):[10.1038/s41579-020-0335-x](https://doi.org/10.1038%2Fs41579-020-0335-x). [hdl](/source/Hdl_(identifier)):[10044/1/78596](https://hdl.handle.net/10044%2F1%2F78596). [PMID](/source/PMID_(identifier)) [32099078](https://pubmed.ncbi.nlm.nih.gov/32099078). [S2CID](/source/S2CID_(identifier)) [211266075](https://api.semanticscholar.org/CorpusID:211266075). [Archived](https://web.archive.org/web/20201107202307/https://discovery.ucl.ac.uk/id/eprint/10092667/1/NRMICRO-19-244_FINAL_ACCEPTED.pdf) (PDF) from the original on November 7, 2020. Retrieved September 28, 2020.

1. **[^](#cite_ref-100)** Grosholz, E.D. (2005). ["Recent biological invasion may hasten invasional meltdown by accelerating historical introductions"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC545825). *[Proceedings of the National Academy of Sciences](/source/Proceedings_of_the_National_Academy_of_Sciences)*. **102** (4): 1088–1091. [Bibcode](/source/Bibcode_(identifier)):[2005PNAS..102.1088G](https://ui.adsabs.harvard.edu/abs/2005PNAS..102.1088G). [doi](/source/Doi_(identifier)):[10.1073/pnas.0308547102](https://doi.org/10.1073%2Fpnas.0308547102). [PMC](/source/PMC_(identifier)) [545825](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC545825). [PMID](/source/PMID_(identifier)) [15657121](https://pubmed.ncbi.nlm.nih.gov/15657121).

1. **[^](#cite_ref-101)** Mungi, Ninad Avinash (2023). ["Distribution, drivers and restoration priorities of plant invasions in India"](https://doi.org/10.1111%2F1365-2664.14506). *Journal of Applied Ecology*. **60** (11): 2400–2412. [Bibcode](/source/Bibcode_(identifier)):[2023JApEc..60.2400M](https://ui.adsabs.harvard.edu/abs/2023JApEc..60.2400M). [doi](/source/Doi_(identifier)):[10.1111/1365-2664.14506](https://doi.org/10.1111%2F1365-2664.14506).

1. **[^](#cite_ref-102)** Rastogi, Rajat (2023). ["Multiple invasions exert combined magnified effects on native plants, soil nutrients and alters the plant-herbivore interaction in dry tropical forest"](https://www.sciencedirect.com/science/article/pii/S0378112723000142). *Forest Ecology and Management*. **531** 120781. [Bibcode](/source/Bibcode_(identifier)):[2023ForEM.53120781R](https://ui.adsabs.harvard.edu/abs/2023ForEM.53120781R). [doi](/source/Doi_(identifier)):[10.1016/j.foreco.2023.120781](https://doi.org/10.1016%2Fj.foreco.2023.120781).

1. ^ [***a***](#cite_ref-causesepidemiology_103-0) [***b***](#cite_ref-causesepidemiology_103-1) Mack, Richard N.; [Simberloff, Daniel](/source/Daniel_Simberloff); Mark Lonsdale, W.; Evans, Harry; Clout, Michael; Bazzaz, Fakhri A. (June 2000). "Biotic Invasions: Causes, Epidemiology, Global Consequences, and Control". *Ecological Applications*. **10** (3): 689–710. [doi](/source/Doi_(identifier)):[10.1890/1051-0761(2000)010\[0689:BICEGC\]2.0.CO;2](https://doi.org/10.1890%2F1051-0761%282000%29010%5B0689%3ABICEGC%5D2.0.CO%3B2). [S2CID](/source/S2CID_(identifier)) [711038](https://api.semanticscholar.org/CorpusID:711038).

1. **[^](#cite_ref-104)** Hawkes, C.V. (2005). "Plant invasion alters nitrogen cycling by modifying the soil nitrifying community". *[Ecology Letters](/source/Ecology_Letters)*. **8** (9): 976–985. [Bibcode](/source/Bibcode_(identifier)):[2005EcolL...8..976H](https://ui.adsabs.harvard.edu/abs/2005EcolL...8..976H). [doi](/source/Doi_(identifier)):[10.1111/j.1461-0248.2005.00802.x](https://doi.org/10.1111%2Fj.1461-0248.2005.00802.x). [PMID](/source/PMID_(identifier)) [34517683](https://pubmed.ncbi.nlm.nih.gov/34517683).

1. ^ [***a***](#cite_ref-rhymer_105-0) [***b***](#cite_ref-rhymer_105-1) Rhymer, J. M.; Simberloff, D. (1996). "Extinction by hybridization and introgression". *[Annual Review of Ecology and Systematics](/source/Annual_Review_of_Ecology_and_Systematics)*. **27** (1): 83–109. [Bibcode](/source/Bibcode_(identifier)):[1996AnRES..27...83R](https://ui.adsabs.harvard.edu/abs/1996AnRES..27...83R). [doi](/source/Doi_(identifier)):[10.1146/annurev.ecolsys.27.1.83](https://doi.org/10.1146%2Fannurev.ecolsys.27.1.83).

1. **[^](#cite_ref-ayres_106-0)** Ayres, D.; et al. (2004). "Spread of exotic cordgrasses and hybrids (*Spartina* sp.) in the tidal marshes of San Francisco Bay, California, USA". *[Biological Invasions](/source/Biological_Invasions)*. **6** (2): 221–231. [Bibcode](/source/Bibcode_(identifier)):[2004BiInv...6..221A](https://ui.adsabs.harvard.edu/abs/2004BiInv...6..221A). [doi](/source/Doi_(identifier)):[10.1023/B:BINV.0000022140.07404.b7](https://doi.org/10.1023%2FB%3ABINV.0000022140.07404.b7). [S2CID](/source/S2CID_(identifier)) [24732543](https://api.semanticscholar.org/CorpusID:24732543).

1. **[^](#cite_ref-107)** Primtel, David (2005). "Update on the environmental and economic costs associated with alien-invasive species in the United States". *[Ecological Economics](/source/Ecological_Economics_(journal))*. **52** (3): 273–288. [Bibcode](/source/Bibcode_(identifier)):[2005EcoEc..52..273P](https://ui.adsabs.harvard.edu/abs/2005EcoEc..52..273P). [doi](/source/Doi_(identifier)):[10.1016/j.ecolecon.2004.10.002](https://doi.org/10.1016%2Fj.ecolecon.2004.10.002).

1. **[^](#cite_ref-108)** Liebhold, S.; et al. (2013). ["A highly aggregated geographical distribution of forest pest invasions in the USA"](https://doi.org/10.1111%2Fddi.12112). *[Diversity and Distributions](/source/Diversity_and_Distributions)*. **19** (9): 1208–1216. [Bibcode](/source/Bibcode_(identifier)):[2013DivDi..19.1208L](https://ui.adsabs.harvard.edu/abs/2013DivDi..19.1208L). [doi](/source/Doi_(identifier)):[10.1111/ddi.12112](https://doi.org/10.1111%2Fddi.12112). [S2CID](/source/S2CID_(identifier)) [85799394](https://api.semanticscholar.org/CorpusID:85799394).

1. **[^](#cite_ref-109)** Oswalt, C.; et al. (2015). ["A subcontinental view of forest plant invasions"](https://doi.org/10.3897%2Fneobiota.24.8378). *[NeoBiota](/source/NeoBiota)*. **24**: 49–54. [doi](/source/Doi_(identifier)):[10.3897/neobiota.24.8378](https://doi.org/10.3897%2Fneobiota.24.8378).

1. ^ [***a***](#cite_ref-pimental_110-0) [***b***](#cite_ref-pimental_110-1) [***c***](#cite_ref-pimental_110-2) [***d***](#cite_ref-pimental_110-3) [***e***](#cite_ref-pimental_110-4) [***f***](#cite_ref-pimental_110-5) [***g***](#cite_ref-pimental_110-6) Pimentel, D.; R., Zuniga; Morrison, D (2005). "Update on the environmental and economic costs associated with alien-invasive species in the United States". *[Ecological Economics](/source/Ecological_Economics)*. **52** (3): 273–288. [Bibcode](/source/Bibcode_(identifier)):[2005EcoEc..52..273P](https://ui.adsabs.harvard.edu/abs/2005EcoEc..52..273P). [doi](/source/Doi_(identifier)):[10.1016/j.ecolecon.2004.10.002](https://doi.org/10.1016%2Fj.ecolecon.2004.10.002).

1. **[^](#cite_ref-111)** ["South/Adelges piceae - Bugwoodwiki"](https://wiki.bugwood.org/Archive:South/Balsam_Woolly_Aphid). *wiki.bugwood.org*. [Archived](https://web.archive.org/web/20110722063118/http://wiki.bugwood.org/Archive:South/Balsam_Woolly_Aphid) from the original on July 22, 2011. Retrieved June 26, 2022.

1. **[^](#cite_ref-112)** Schlarbaum, Scott E., Frederick Hebard, Pauline C. Spaine, and Joseph C. Kamalay. (1998) ["Three American Tragedies: Chestnut Blight, Butternut Canker, and Dutch Elm Disease'](https://research.fs.usda.gov/treesearch/745) . In: Britton, Kerry O., Ed. Exotic Pests of Eastern Forests Conference Proceedings; 1997 April 8–10; Nashville, TN. U.S. Forest Service and Tennessee Exotic Pest Plant Council., pp. 45–54.

1. **[^](#cite_ref-USDA-Forest_Service-Schlarbaum-1997_113-0)** Schlarbaum, Scott E.; Hebard, Frederick; Spaine, Pauline C.; Kamalay, Joseph C. (1997). ["Three American Tragedies: Chestnut Blight, Butternut Canker and Dutch Elm Disease"](https://www.srs.fs.usda.gov/pubs/ja/ja_schlarbaum002.htm). *(originally published via: Proceedings: Exotic Pests of Eastern Forests; (1997 April 8–10); Nashville, TN. Tennessee Exotic Pest Plant Council: 45–54.)*. Southern Research Station, [Forest Service](/source/United_States_Forest_Service), [United States Department of Agriculture](/source/United_States_Department_of_Agriculture). [Archived](https://web.archive.org/web/20120424101943/http://www.srs.fs.usda.gov/pubs/ja/ja_schlarbaum002.htm) from the original on April 24, 2012. Retrieved June 22, 2012.

1. **[^](#cite_ref-114)** Schlarbaum, Scott E.; Hebard, Frederick; Spaine, Pauline C.; Kamalay, Joseph C.; Schlarbaum, Scott E.; Hebard, Frederick; Spaine, Pauline C.; Kamalay, Joseph C. ["Southern Research Station"](https://web.archive.org/web/20121123093613/http://www.treesearch.fs.fed.us/pubs/745). *Tree Search*. US Forest Service, USDA. Archived from [the original](http://www.treesearch.fs.fed.us/pubs/745) on November 23, 2012.

1. **[^](#cite_ref-115)** Rodger, Vikki; Stinson, Kristin; Finzi, Adrian (2008). ["Ready or Not, Garlic Mustard Is Moving In: *Alliaria petiolata* as a Member of Eastern North American Forests"](https://doi.org/10.1641%2Fb580510). *[BioScience](/source/BioScience)*. **58** (5): 5. [Bibcode](/source/Bibcode_(identifier)):[2008BiSci..58..426R](https://ui.adsabs.harvard.edu/abs/2008BiSci..58..426R). [doi](/source/Doi_(identifier)):[10.1641/b580510](https://doi.org/10.1641%2Fb580510).

1. **[^](#cite_ref-116)** ["Kākāpō Recovery Programme"](https://www.meridianenergy.co.nz/community-support/kakapo-recovery-programme). *Meridian Energy*. July 10, 2024. Retrieved February 26, 2026.

1. ^ [***a***](#cite_ref-:1_117-0) [***b***](#cite_ref-:1_117-1) Mooney, HA; Cleland, EE (2001). ["The evolutionary impact of invasive species"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC33232). *[Proceedings of the National Academy of Sciences of the United States of America](/source/Proceedings_of_the_National_Academy_of_Sciences_of_the_United_States_of_America)*. **98** (10): 5446–51. [Bibcode](/source/Bibcode_(identifier)):[2001PNAS...98.5446M](https://ui.adsabs.harvard.edu/abs/2001PNAS...98.5446M). [doi](/source/Doi_(identifier)):[10.1073/pnas.091093398](https://doi.org/10.1073%2Fpnas.091093398). [PMC](/source/PMC_(identifier)) [33232](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC33232). [PMID](/source/PMID_(identifier)) [11344292](https://pubmed.ncbi.nlm.nih.gov/11344292).

1. **[^](#cite_ref-118)** ["Glossary: definitions from the following publication: Aubry, C., R. Shoal and V. Erickson. 2005. Grass cultivars: their origins, development, and use on national forests and grasslands in the Pacific Northwest. USDA Forest Service. 44 pages, plus appendices.; Native Seed Network (NSN), Institute for Applied Ecology, 563 SW Jefferson Ave, Corvallis, OR 97333, USA"](https://web.archive.org/web/20060222092651/http://www.nativeseednetwork.org/article_view?id=13). Nativeseednetwork.org. Archived from [the original](http://www.nativeseednetwork.org/article_view?id=13) on February 22, 2006. Retrieved May 17, 2011.

1. **[^](#cite_ref-119)** Boudjelas, Souyad (2000). [100 of the world's worst invasive alien species](https://portals.iucn.org/library/sites/library/files/documents/2000-126.pdf) (PDF). *iucn.org* (Report). [International Union for Conservation of Nature](/source/International_Union_for_Conservation_of_Nature). Retrieved July 20, 2018.

1. **[^](#cite_ref-120)** Anttila, C. K.; King, R. A.; Ferris, C.; Ayres, D. R.; Strong, D. R. (2000). "Reciprocal hybrid formation of Spartina in San Francisco Bay". *[Molecular Ecology](/source/Molecular_Ecology)*. **9** (6): 765–770. [Bibcode](/source/Bibcode_(identifier)):[2000MolEc...9..765A](https://ui.adsabs.harvard.edu/abs/2000MolEc...9..765A). [doi](/source/Doi_(identifier)):[10.1046/j.1365-294x.2000.00935.x](https://doi.org/10.1046%2Fj.1365-294x.2000.00935.x). [PMID](/source/PMID_(identifier)) [10849292](https://pubmed.ncbi.nlm.nih.gov/10849292). [S2CID](/source/S2CID_(identifier)) [32865913](https://api.semanticscholar.org/CorpusID:32865913).

1. **[^](#cite_ref-121)** [*Genetic Pollution from Farm Forestry using eucalypt species and hybrids; A report for the RIRDC/L&WA/FWPRDC\]; Joint Venture Agroforestry Program; by Brad M. Potts, Robert C. Barbour, Andrew B. Hingston; September 2001; RIRDC Publication No 01/114; RIRDC Project No CPF – 3A;*](https://web.archive.org/web/20040102175403/http://www.rirdc.gov.au/reports/AFT/01-114.pdf) (PDF). Australian Government, Rural Industrial Research and Development Corporation. 2001. [ISBN](/source/ISBN_(identifier)) [978-0-642-58336-9](https://en.wikipedia.org/wiki/Special:BookSources/978-0-642-58336-9). Archived from [the original](http://www.rirdc.gov.au/reports/AFT/01-114.pdf) (PDF) on January 2, 2004. Retrieved April 22, 2017.

1. **[^](#cite_ref-122)** Bohling, Justin H.; Waits, Lisette P. (2015). "Factors influencing red wolf–coyote hybridization in eastern North Carolina, USA". *[Biological Conservation](/source/Biological_Conservation)*. **184**: 108–116. [Bibcode](/source/Bibcode_(identifier)):[2015BCons.184..108B](https://ui.adsabs.harvard.edu/abs/2015BCons.184..108B). [doi](/source/Doi_(identifier)):[10.1016/j.biocon.2015.01.013](https://doi.org/10.1016%2Fj.biocon.2015.01.013).

1. **[^](#cite_ref-123)** ["Cape Town is Facing Day Zero"](https://www.nature.org/en-us/about-us/where-we-work/africa/stories-in-africa/cape-town-faces--day-zero-/). *The Nature Conservancy*. Retrieved November 6, 2023.

1. **[^](#cite_ref-124)** ["Greater cape town water fund"](https://www.nature.org/content/dam/tnc/nature/en/documents/GCTWF-summary-11.14.18.pdf) (PDF). [Archived](https://web.archive.org/web/20210228175854/https://www.nature.org/content/dam/tnc/nature/en/documents/GCTWF-summary-11.14.18.pdf) (PDF) from the original on February 28, 2021. Retrieved November 16, 2020.

1. **[^](#cite_ref-:8_125-0)** Mazza, G.; Tricarico, E.; Genovesi, P.; Gherardi, F. (December 19, 2013). "Biological invaders are threats to human health: an overview". *Ethology Ecology & Evolution*. **26** (2–3): 112–129. [doi](/source/Doi_(identifier)):[10.1080/03949370.2013.863225](https://doi.org/10.1080%2F03949370.2013.863225). [S2CID](/source/S2CID_(identifier)) [58888740](https://api.semanticscholar.org/CorpusID:58888740).

1. ^ [***a***](#cite_ref-:22_126-0) [***b***](#cite_ref-:22_126-1) [***c***](#cite_ref-:22_126-2) [***d***](#cite_ref-:22_126-3) [***e***](#cite_ref-:22_126-4) Pyšek, P.; Richardson, D.M. (2010). ["Invasive Species, Environmental Change and Management, and Health"](https://doi.org/10.1146%2Fannurev-environ-033009-095548). *Annual Review of Environment and Resources*. **35** (1): 25–55. [doi](/source/Doi_(identifier)):[10.1146/annurev-environ-033009-095548](https://doi.org/10.1146%2Fannurev-environ-033009-095548). [hdl](/source/Hdl_(identifier)):[11104/0189317](https://hdl.handle.net/11104%2F0189317).

1. **[^](#cite_ref-127)** Lanciotti, R. S.; Roehrig, J. T.; Deubel, V.; Smith, J.; Parker, M.; Steele, K.; et al. (December 17, 1999). "Origin of the West Nile Virus Responsible for an Outbreak of Encephalitis in the Northeastern United States". *Science*. **286** (5448): 2333–2337. [doi](/source/Doi_(identifier)):[10.1126/science.286.5448.2333](https://doi.org/10.1126%2Fscience.286.5448.2333). [PMID](/source/PMID_(identifier)) [10600742](https://pubmed.ncbi.nlm.nih.gov/10600742).

1. **[^](#cite_ref-128)** Hallegraeff, G.M. (1998). ["Transport of toxic dinoflagellates via ships' ballast water: Bioeconomic risk assessment and efficacy of possible ballast water management strategies"](https://doi.org/10.3354%2Fmeps168297). *[Marine Ecology Progress Series](/source/Marine_Ecology_Progress_Series)*. **168**: 297–309. [Bibcode](/source/Bibcode_(identifier)):[1998MEPS..168..297H](https://ui.adsabs.harvard.edu/abs/1998MEPS..168..297H). [doi](/source/Doi_(identifier)):[10.3354/meps168297](https://doi.org/10.3354%2Fmeps168297).

1. **[^](#cite_ref-129)** Dela Cruz, Raymond Carl (October 7, 2020). ["Water hyacinths ground Pasig River Ferry ops"](https://web.archive.org/web/20231028150214/https://www.pna.gov.ph/articles/1117832). *[Philippine News Agency](/source/Philippine_News_Agency)*. Archived from [the original](https://www.pna.gov.ph/articles/1117832) on October 28, 2023. Retrieved August 10, 2024.

1. **[^](#cite_ref-130)** Environment, U. N. (September 4, 2023). ["Invasive Alien Species Report"](http://www.unep.org/resources/report/invasive-alien-species-report). *www.unep.org*. Retrieved May 29, 2024.

1. **[^](#cite_ref-131)** Xu, Haigen; Ding, Hui; Li, Mingyan; Qiang, Sheng; Guo, Jianying; Han, Zhengmin; Huang, Zongguo; Sun, Hongying; He, Shunping; Wu, Hairong; Wan, Fanghao (2006). "The distribution and economic losses of alien species invasion to China". *Biological Invasions*. **8** (7): 1495–1500. [Bibcode](/source/Bibcode_(identifier)):[2006BiInv...8.1495X](https://ui.adsabs.harvard.edu/abs/2006BiInv...8.1495X). [doi](/source/Doi_(identifier)):[10.1007/s10530-005-5841-2](https://doi.org/10.1007%2Fs10530-005-5841-2). [S2CID](/source/S2CID_(identifier)) [25890246](https://api.semanticscholar.org/CorpusID:25890246).

1. ^ [***a***](#cite_ref-potential_132-0) [***b***](#cite_ref-potential_132-1) [***c***](#cite_ref-potential_132-2) [***d***](#cite_ref-potential_132-3) [***e***](#cite_ref-potential_132-4) Schlaepfer, Martin A.; Sax, Dov F.; Olden, Julian D. (June 2011). "The Potential Conservation Value of Non-Native Species: Conservation Value of Non-Native Species". *Conservation Biology*. **25** (3): 428–437. [doi](/source/Doi_(identifier)):[10.1111/j.1523-1739.2010.01646.x](https://doi.org/10.1111%2Fj.1523-1739.2010.01646.x). [PMID](/source/PMID_(identifier)) [21342267](https://pubmed.ncbi.nlm.nih.gov/21342267). [S2CID](/source/S2CID_(identifier)) [2947682](https://api.semanticscholar.org/CorpusID:2947682).

1. **[^](#cite_ref-:32_133-0)** Molnar, Jennifer L; Gamboa, Rebecca L; Revenga, Carmen; Spalding, Mark D (2008). "Assessing the global threat of invasive species to marine biodiversity". *Frontiers in Ecology and the Environment*. **6** (9): 485–492. [Bibcode](/source/Bibcode_(identifier)):[2008FrEE....6..485M](https://ui.adsabs.harvard.edu/abs/2008FrEE....6..485M). [doi](/source/Doi_(identifier)):[10.1890/070064](https://doi.org/10.1890%2F070064).

1. ^ [***a***](#cite_ref-:0_134-0) [***b***](#cite_ref-:0_134-1) ["Great Lakes Fishery Commission – Sea Lamprey"](http://www.glfc.org/sea-lamprey.php). *www.glfc.org*. [Archived](https://web.archive.org/web/20171025023343/http://www.glfc.org/sea-lamprey.php) from the original on October 25, 2017. Retrieved October 24, 2017.

1. **[^](#cite_ref-simb_135-0)** Simberloff, D. (2001). "Biological invasions – How are they affecting us, and what can we do about them?". *[Western North American Naturalist](/source/Western_North_American_Naturalist)*. **61** (3): 308–315. [JSTOR](/source/JSTOR_(identifier)) [41717176](https://www.jstor.org/stable/41717176).

1. **[^](#cite_ref-136)** [*2008–2012 National Invasive Species Management Plan*](https://www.doi.gov/sites/doi.gov/files/migrated/invasivespecies/upload/2008-2012-National-Invasive-Species-Management-Plan.pdf) (PDF). Washington, DC.: National Invasive Species Council, Department of the Interior. 2008. [Archived](https://web.archive.org/web/20150929011911/https://www.doi.gov/sites/doi.gov/files/migrated/invasivespecies/upload/2008-2012-National-Invasive-Species-Management-Plan.pdf) (PDF) from the original on September 29, 2015.

1. **[^](#cite_ref-137)** Holden, Matthew H.; Nyrop, Jan P.; Ellner, Stephen P. (June 1, 2016). ["The economic benefit of time-varying surveillance effort for invasive species management"](https://doi.org/10.1111%2F1365-2664.12617). *[Journal of Applied Ecology](/source/Journal_of_Applied_Ecology)*. **53** (3): 712–721. [Bibcode](/source/Bibcode_(identifier)):[2016JApEc..53..712H](https://ui.adsabs.harvard.edu/abs/2016JApEc..53..712H). [doi](/source/Doi_(identifier)):[10.1111/1365-2664.12617](https://doi.org/10.1111%2F1365-2664.12617).

1. **[^](#cite_ref-138)** Gougherty, Andrew V.; Davies, T. Jonathan (November 8, 2021). ["Towards a phylogenetic ecology of plant pests and pathogens"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8450633). *Philosophical Transactions of the Royal Society B: Biological Sciences*. **376** (1837) 20200359. [doi](/source/Doi_(identifier)):[10.1098/rstb.2020.0359](https://doi.org/10.1098%2Frstb.2020.0359). [PMC](/source/PMC_(identifier)) [8450633](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8450633). [PMID](/source/PMID_(identifier)) [34538142](https://pubmed.ncbi.nlm.nih.gov/34538142).

1. **[^](#cite_ref-139)** ["American serpentine leafminer – Liriomyza trifolii (Burgess)"](http://entnemdept.ufl.edu/creatures/veg/leaf/a_serpentine_leafminer.htm). *entnemdept.ufl.edu*. [Archived](https://web.archive.org/web/20191125043634/http://entnemdept.ufl.edu/Creatures/veg/leaf/a_serpentine_leafminer.htm) from the original on November 25, 2019. Retrieved November 20, 2019.

1. **[^](#cite_ref-140)** Eiswerth, M.E.; Darden, Tim D.; Johnson, Wayne S.; Agapoff, Jeanmarie; Harris, Thomas R. (2005). "Input-output modeling, outdoor recreation, and the economic impacts of weeds". *[Weed Science](/source/Weed_Science)*. **53** (1): 130–137. [Bibcode](/source/Bibcode_(identifier)):[2005WeedS..53..130E](https://ui.adsabs.harvard.edu/abs/2005WeedS..53..130E). [doi](/source/Doi_(identifier)):[10.1614/WS-04-022R](https://doi.org/10.1614%2FWS-04-022R). [S2CID](/source/S2CID_(identifier)) [85608607](https://api.semanticscholar.org/CorpusID:85608607).

1. **[^](#cite_ref-141)** ["Eurasian Watermilfoil in the Great Lakes Region"](https://web.archive.org/web/20080725034837/http://great-lakes.net/envt/flora-fauna/invasive/milfoil.html#overview). *Great Lakes Information Network*. November 1, 2006. Archived from [the original](http://great-lakes.net/envt/flora-fauna/invasive/milfoil.html) on July 25, 2008.

1. **[^](#cite_ref-142)** Sin, Hans; Radford, Adam (2007). "Coqui frog research and management efforts in Hawaii". [*Managing Vertebrate Invasive Species: Proceedings of an International Symposium (G. W. Witmer, W. C. Pitt, K. A. Fagerstone, Eds)*](https://web.archive.org/web/20170525103353/http://www.aphis.usda.gov/wildlife_damage/nwrc/symposia/invasive_symposium/content/Sin157_167_MVIS.pdf) (PDF). Fort Collins, Colorado: USDA/APHIS/WS, National Wildlife Research Center. Archived from [the original](https://www.aphis.usda.gov/wildlife_damage/nwrc/symposia/invasive_symposium/content/Sin157_167_MVIS.pdf) (PDF) on May 25, 2017. Retrieved June 26, 2013.

1. **[^](#cite_ref-143)** ["Spider Invaders"](https://www.kqed.org/quest/9595/spider-invaders). *KQED*. October 18, 2010. [Archived](https://web.archive.org/web/20201105170805/https://www.kqed.org/quest/9595/spider-invaders) from the original on November 5, 2020. Retrieved December 13, 2020.

1. **[^](#cite_ref-144)** Haubrock, Phillip J.; Turbelin, Anna J.; Cuthbert, Ross N.; Novoa, Ana; Taylor, Nigel G.; Angulo, Elena; et al. (2021). ["Economic costs of invasive alien species across Europe"](https://doi.org/10.3897%2Fneobiota.67.58196). *[Neobiota](/source/Neobiota)*. **67**: 153–190. [Bibcode](/source/Bibcode_(identifier)):[2021NeoBi..67..153H](https://ui.adsabs.harvard.edu/abs/2021NeoBi..67..153H). [doi](/source/Doi_(identifier)):[10.3897/neobiota.67.58196](https://doi.org/10.3897%2Fneobiota.67.58196). [hdl](/source/Hdl_(identifier)):[10138/333320](https://hdl.handle.net/10138%2F333320). [S2CID](/source/S2CID_(identifier)) [237460752](https://api.semanticscholar.org/CorpusID:237460752).

1. **[^](#cite_ref-145)** Haubrock, Phillip J.; Cuthbert, Ross N.; Tricarico, Elena; Diagne, Christophe; Courchamp, Franck; Gozlan, Rodolphe E. (July 29, 2021). ["The recorded economic costs of alien invasive species in Italy"](https://hal.archives-ouvertes.fr/hal-03410329/file/CostsinItaly.pdf) (PDF). *NeoBiota*. **67**: 247–266. [Bibcode](/source/Bibcode_(identifier)):[2021NeoBi..67..247H](https://ui.adsabs.harvard.edu/abs/2021NeoBi..67..247H). [doi](/source/Doi_(identifier)):[10.3897/neobiota.67.57747](https://doi.org/10.3897%2Fneobiota.67.57747). [hdl](/source/Hdl_(identifier)):[2158/1262519](https://hdl.handle.net/2158%2F1262519). [S2CID](/source/S2CID_(identifier)) [238819772](https://api.semanticscholar.org/CorpusID:238819772).

1. **[^](#cite_ref-146)** Renault, David; Manfrini, Eléna; Leroy, Boris; Diagne, Christophe; Ballesteros-Mejia, Liliana; Angulo, Elena; Courchamp, Franck (July 29, 2021). ["Biological invasions in France: Alarming costs and even more alarming knowledge gaps"](https://doi.org/10.3897%2Fneobiota.67.59134). *NeoBiota*. **67**: 191–224. [Bibcode](/source/Bibcode_(identifier)):[2021NeoBi..67..191R](https://ui.adsabs.harvard.edu/abs/2021NeoBi..67..191R). [doi](/source/Doi_(identifier)):[10.3897/neobiota.67.59134](https://doi.org/10.3897%2Fneobiota.67.59134). [S2CID](/source/S2CID_(identifier)) [237462170](https://api.semanticscholar.org/CorpusID:237462170).

1. **[^](#cite_ref-doubtingthomas_147-0)** Halley, John (2019). ["Doubting Thomas and the Love of Invasive Species"](https://doi.org/10.1111%2Fcobi.13413). Book Review. *[Conservation Biology](/source/Conservation_Biology_(journal))*. **33** (6): 1451–1453. [Bibcode](/source/Bibcode_(identifier)):[2019ConBi..33.1451H](https://ui.adsabs.harvard.edu/abs/2019ConBi..33.1451H). [doi](/source/Doi_(identifier)):[10.1111/cobi.13413](https://doi.org/10.1111%2Fcobi.13413).

1. **[^](#cite_ref-148)** Cadotte, Marc W. (October 1, 2015). ["Taming the 'New Wild': tackling the good and the bad with invasive species"](https://doi.org/10.1007/s10530-015-0933-0). *Biological Invasions*. **17** (10): 3067–3072. [doi](/source/Doi_(identifier)):[10.1007/s10530-015-0933-0](https://doi.org/10.1007%2Fs10530-015-0933-0). [ISSN](/source/ISSN_(identifier)) [1573-1464](https://search.worldcat.org/issn/1573-1464).

1. **[^](#cite_ref-149)** [Thomas, Chris](/source/Chris_D._Thomas) (2017). *Inheritors of the Earth: How Nature Is Thriving in an Age of Extinction*. [PublicAffairs](/source/PublicAffairs). [ISBN](/source/ISBN_(identifier)) [978-1-61039-727-8](https://en.wikipedia.org/wiki/Special:BookSources/978-1-61039-727-8).

1. **[^](#cite_ref-150)** Jr, Tom Zeller (May 30, 2017). ["The Fascist History of De-Extinction"](https://undark.org/2017/05/30/nazis-aurochs-deextinction-mammoth/). *Undark Magazine*. Retrieved April 27, 2025.

1. **[^](#cite_ref-:5_151-0)** Fletcher, Rebecca A.; Brooks, Rachel K.; Lakoba, Vasiliy T.; Sharma, Gourav; Heminger, Ariel R.; Dickinson, Christopher C.; Barney, Jacob N. (2019). ["Invasive plants negatively impact native, but not exotic, animals"](https://onlinelibrary.wiley.com/doi/abs/10.1111/gcb.14752). *Global Change Biology*. **25** (11): 3694–3705. [Bibcode](/source/Bibcode_(identifier)):[2019GCBio..25.3694F](https://ui.adsabs.harvard.edu/abs/2019GCBio..25.3694F). [doi](/source/Doi_(identifier)):[10.1111/gcb.14752](https://doi.org/10.1111%2Fgcb.14752). [ISSN](/source/ISSN_(identifier)) [1365-2486](https://search.worldcat.org/issn/1365-2486). [PMID](/source/PMID_(identifier)) [31389131](https://pubmed.ncbi.nlm.nih.gov/31389131).

1. **[^](#cite_ref-152)** Mehta, Kanishka; Koli, Vijay K.; Kittur, Swati; Sundar, K. S. Gopi (February 21, 2024). "Can you nest where you roost? Waterbirds use different sites but similar cues to locate roosting and breeding sites in a small Indian city". *Urban Ecosystems*. **27** (4): 1279–1290. [Bibcode](/source/Bibcode_(identifier)):[2024UrbEc..27.1279M](https://ui.adsabs.harvard.edu/abs/2024UrbEc..27.1279M). [doi](/source/Doi_(identifier)):[10.1007/s11252-023-01454-5](https://doi.org/10.1007%2Fs11252-023-01454-5). [S2CID](/source/S2CID_(identifier)) [267973120](https://api.semanticscholar.org/CorpusID:267973120).

1. **[^](#cite_ref-153)** McBroom, Jen (December 2012). [Clapper Rail Surveys for the San Francisco Estuary Invasive Spartina Project](https://spartina.org/project_documents/revegetation_program/CLRA%20Report%202012.pdf) (PDF) (Report). Oakland, California: State Coastal Conservancy. [Archived](https://web.archive.org/web/20170305070303/http://spartina.org/project_documents/revegetation_program/CLRA%20Report%202012.pdf) (PDF) from the original on March 5, 2017. Retrieved November 30, 2020.

1. **[^](#cite_ref-154)** Ham, Anthony (August 15, 2022). ["Pigs to the Rescue: An Invasive Species Helped Save Australia's Crocodiles"](https://www.nytimes.com/2022/08/15/science/invasive-species-pigs-crocodiles.html). *The New York Times*.

1. **[^](#cite_ref-155)** Thomas, Chris D.. Inheritors of the Earth (p. 148). PublicAffairs. Kindle Edition.

1. **[^](#cite_ref-wolv_156-0)** Wolverton, B. C.; McDonald, Rebecca C. (1981). "Energy from vascular plant wastewater treatment systems". *[Economic Botany](/source/Economic_Botany)*. **35** (2): 224–232. [Bibcode](/source/Bibcode_(identifier)):[1981EcBot..35..224W](https://ui.adsabs.harvard.edu/abs/1981EcBot..35..224W). [doi](/source/Doi_(identifier)):[10.1007/BF02858689](https://doi.org/10.1007%2FBF02858689). [S2CID](/source/S2CID_(identifier)) [24217507](https://api.semanticscholar.org/CorpusID:24217507).. Cited in Duke, J. (1983) [*Handbook of Energy Crops*](http://www.hort.purdue.edu/newcrop/duke_energy/dukeindex.html) [Archived](https://web.archive.org/web/20130212071515/http://www.hort.purdue.edu/newcrop/duke_energy/dukeindex.html) February 12, 2013, at the [Wayback Machine](/source/Wayback_Machine). Purdue University, Center for New Crops & Plants Products

1. **[^](#cite_ref-157)** Van Meerbeek, Koenraad; Appels, Lise; Dewil, Raf; Calmeyn, Annelies; Lemmens, Pieter; Muys, Bart; Hermy, Martin (May 1, 2015). "Biomass of invasive plant species as a potential feedstock for bioenergy production". *[Biofuels, Bioproducts and Biorefining](/source/Biofuels%2C_Bioproducts_and_Biorefining)*. **9** (3): 273–282. [doi](/source/Doi_(identifier)):[10.1002/bbb.1539](https://doi.org/10.1002%2Fbbb.1539). [S2CID](/source/S2CID_(identifier)) [83918875](https://api.semanticscholar.org/CorpusID:83918875).

1. **[^](#cite_ref-158)** Roelvink, Gerda; Martin, Kevin St; Gibson-Graham, J. K. (2015). *Making Other Worlds Possible: Performing Diverse Economies*. University of Minnesota Press. [ISBN](/source/ISBN_(identifier)) [978-0-8166-9329-0](https://en.wikipedia.org/wiki/Special:BookSources/978-0-8166-9329-0).

1. **[^](#cite_ref-Garrido_159-0)** Garrido-Pérez, Edgardo I.; Tella Ruiz, David (2016). ["Homo sapiens (Primates: Hominidae): ¿una especie invasora o aún peor? Un reto para potenciar la Ecología y la Biología de la conservación"](https://www.researchgate.net/publication/317083729). *Puente Biológico*. **8**: 43–55. Translated as Garrido-Pérez, Edgardo I.; Tella Ruiz, David (2016). ["*Homo sapiens* (Primates: Hominidae): an invasive species or even worse? A challenge for strengthening ecology and conservation biology"](https://www.researchgate.net/publication/319234497). [Archived](https://web.archive.org/web/20220611120711/https://www.researchgate.net/publication/319234497_Homo_sapiens_Primates_Hominidae_an_invasive_species_or_even_worse_A_challenge_for_strengthening_ecology_and_conservation_biology_Translated_from_Spanish) from the original on June 11, 2022. Retrieved August 19, 2020 – via ResearchGate.

1. **[^](#cite_ref-160)** Hakam, Lara (February 2013). ["Invasive Species: Public Awareness and Education"](http://depts.washington.edu/oldenlab/wordpress/wp-content/uploads/2013/02/Invasive-Species-Survey_Hakam_August2016.pdf#page=8) (PDF). *University of Washington*. [Archived](https://web.archive.org/web/20211105154129/http://depts.washington.edu/oldenlab/wordpress/wp-content/uploads/2013/02/Invasive-Species-Survey_Hakam_August2016.pdf#page=8) (PDF) from the original on November 5, 2021. Retrieved September 30, 2020.

1. **[^](#cite_ref-161)** Williams, Peter A. (November 1997). [*Ecology and Management of Invasive Weeds*](https://www.doc.govt.nz/Documents/science-and-technical/csp07-entire.pdf) (PDF). Dept. of Conservation, Wellington, N.Z. p. 22. [ISBN](/source/ISBN_(identifier)) [0-478-01956-4](https://en.wikipedia.org/wiki/Special:BookSources/0-478-01956-4).{{[cite book](https://en.wikipedia.org/wiki/Template:Cite_book)}}: CS1 maint: date and year ([link](https://en.wikipedia.org/wiki/Category:CS1_maint:_date_and_year))

1. **[^](#cite_ref-162)** Makhrov, A. A.; Karabanov, D. P.; Koduhova, Yu. V. (July 2014). "Genetic methods for the control of alien species". *Russian Journal of Biological Invasions*. **5** (3): 194–202. [Bibcode](/source/Bibcode_(identifier)):[2014RuJBI...5..194M](https://ui.adsabs.harvard.edu/abs/2014RuJBI...5..194M). [doi](/source/Doi_(identifier)):[10.1134/S2075111714030096](https://doi.org/10.1134%2FS2075111714030096). [S2CID](/source/S2CID_(identifier)) [256073288](https://api.semanticscholar.org/CorpusID:256073288).

1. **[^](#cite_ref-Lodge-et-al-2016_163-0)** Lodge, David M.; Simonin, Paul W.; Burgiel, Stanley W.; Keller, Reuben P.; Bossenbroek, Jonathan M.; Jerde, Christopher L.; et al. (November 1, 2016). ["Risk Analysis and Bioeconomics of Invasive Species to Inform Policy and Management"](https://doi.org/10.1146%2Fannurev-environ-110615-085532). *Annual Review of Environment and Resources*. **41** (1): 453–488. [doi](/source/Doi_(identifier)):[10.1146/annurev-environ-110615-085532](https://doi.org/10.1146%2Fannurev-environ-110615-085532).

1. **[^](#cite_ref-Sea-Grant-NY_164-0)** O'Neill, Jr., Charles R. (2002). ["Zebra Mussels and Fire Control Equipment"](http://www.slocounty.ca.gov/Departments/Public-Works/Forms-Documents/Water-Resources/Invasive-Mussels/NY-Fire-Control-Equipment.pdf) (PDF). [SUNY College at Brockport](/source/SUNY_College_at_Brockport): [Sea Grant](/source/National_Sea_Grant_College_Program). [Archived](https://web.archive.org/web/20211105212840/https://www.slocounty.ca.gov/Departments/Public-Works/Forms-Documents/Water-Resources/Invasive-Mussels/NY-Fire-Control-Equipment.pdf) (PDF) from the original on November 5, 2021. Retrieved May 23, 2021.

1. **[^](#cite_ref-wildland-firefight-NPR_165-0)** Ouellet, Nicky (August 23, 2017). ["Wildland Firefighters Try to Combat Spread of Invasive Species"](http://www.npr.org/2017/08/23/545617315/wildland-firefighters-try-to-combat-spread-of-invasive-species). *[All Things Considered](/source/All_Things_Considered)*. [NPR](/source/NPR). [Archived](https://web.archive.org/web/20210613005731/https://www.npr.org/2017/08/23/545617315/wildland-firefighters-try-to-combat-spread-of-invasive-species) from the original on June 13, 2021. Retrieved May 23, 2021.

1. **[^](#cite_ref-Montana-firefight-MtPR_166-0)** Ouellet, Nicky (July 27, 2017). ["How Montana Is Fighting Invasive Hitchhikers On Firefighting Aircraft"](http://www.mtpr.org/post/how-montana-fighting-invasive-hitchhikers-firefighting-aircraft). *[Montana Public Radio](/source/Montana_Public_Radio)*. [Archived](https://web.archive.org/web/20210523185157/https://www.mtpr.org/post/how-montana-fighting-invasive-hitchhikers-firefighting-aircraft) from the original on May 23, 2021. Retrieved May 23, 2021.

1. **[^](#cite_ref-NWCG_167-0)** [National Wildfire Coordinating Group](/source/National_Wildfire_Coordinating_Group) (January 2017). ["Guide to Preventing Aquatic Invasive Species Transport by Wildland Fire Operations"](https://www.nwcg.gov/sites/default/files/publications/pms444.pdf) (PDF). [Archived](https://web.archive.org/web/20210419034257/https://www.nwcg.gov/sites/default/files/publications/pms444.pdf) (PDF) from the original on April 19, 2021. Retrieved May 23, 2021.

1. **[^](#cite_ref-Mt-DNRC_168-0)** [National Wildfire Coordinating Group](/source/National_Wildfire_Coordinating_Group) (June 11, 2018). ["Decontaminating Firefighting Equipment to Reduce the Spread of Aquatic Invasive Species"](http://dnrc.mt.gov/divisions/forestry/docs/fire-and-aviation/2018-nrcg-how-to-guide_aquatic-invasive-species_final.pdf) (PDF). [Archived](https://web.archive.org/web/20210428072947/http://dnrc.mt.gov/divisions/forestry/docs/fire-and-aviation/2018-nrcg-how-to-guide_aquatic-invasive-species_final.pdf) (PDF) from the original on April 28, 2021. Retrieved May 23, 2021.

1. **[^](#cite_ref-169)** Holmes, Nick (March 27, 2019). ["Globally important islands where eradicating invasive mammals will benefit highly threatened vertebrates"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6436766). *[PLOS ONE](/source/PLOS_ONE)*. **14** (3) e0212128. [Bibcode](/source/Bibcode_(identifier)):[2019PLoSO..1412128H](https://ui.adsabs.harvard.edu/abs/2019PLoSO..1412128H). [doi](/source/Doi_(identifier)):[10.1371/journal.pone.0212128](https://doi.org/10.1371%2Fjournal.pone.0212128). [PMC](/source/PMC_(identifier)) [6436766](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6436766). [PMID](/source/PMID_(identifier)) [30917126](https://pubmed.ncbi.nlm.nih.gov/30917126).

1. **[^](#cite_ref-de_WitZilliacus2020_170-0)** de Wit, Luz A.; Zilliacus, Kelly M.; Quadri, Paulo; Will, David; Grima, Nelson; Spatz, Dena; et al. (September 2020). ["Invasive vertebrate eradications on islands as a tool for implementing global Sustainable Development Goals"](https://doi.org/10.1017%2FS0376892920000211). *Environmental Conservation*. **47** (3): 139–148. [Bibcode](/source/Bibcode_(identifier)):[2020EnvCo..47..139D](https://ui.adsabs.harvard.edu/abs/2020EnvCo..47..139D). [doi](/source/Doi_(identifier)):[10.1017/S0376892920000211](https://doi.org/10.1017%2FS0376892920000211). [S2CID](/source/S2CID_(identifier)) [221990256](https://api.semanticscholar.org/CorpusID:221990256).

1. **[^](#cite_ref-171)** ["Pursuing Sustainable Development for Island Communities by Removing Invasive Species"](https://www.islandconservation.org/sustainable-development-communities-removing-invasive-species/). Island Conservation. August 13, 2020. [Archived](https://web.archive.org/web/20200926042621/https://www.islandconservation.org/sustainable-development-communities-removing-invasive-species/) from the original on September 26, 2020. Retrieved August 13, 2020.

1. **[^](#cite_ref-172)** Warren, Matt (May 8, 2018). ["Rat begone: Record eradication effort rids sub-Antarctic island of invasive rodents"](https://www.science.org/content/article/rat-begone-record-eradication-effort-rids-sub-antarctic-island-invasive-rodents). *Science*. [Archived](https://web.archive.org/web/20180509035618/http://www.sciencemag.org/news/2018/05/rat-begone-record-eradication-effort-rids-sub-antarctic-island-invasive-rodents) from the original on May 9, 2018. Retrieved May 9, 2018.

1. **[^](#cite_ref-173)** Hester, Jessica Leight (May 17, 2018). ["The Intrepid Rat-Sniffing Terriers of South Georgia Island"](https://www.atlasobscura.com/articles/how-to-exterminate-rats-on-an-island). *[Atlas Obscura](/source/Atlas_Obscura)*. [Archived](https://web.archive.org/web/20180522041931/https://www.atlasobscura.com/articles/how-to-exterminate-rats-on-an-island) from the original on May 22, 2018. Retrieved June 6, 2018.

1. **[^](#cite_ref-174)** ["Invasive plants can create positive ecological change"](https://www.sciencedaily.com/releases/2011/02/110211095555.htm). *Science Daily*. February 14, 2011. [Archived](https://web.archive.org/web/20170525102601/https://www.sciencedaily.com/releases/2011/02/110211095555.htm) from the original on May 25, 2017. Retrieved June 22, 2017. Invasive species could fill niches in degraded ecosystems and help restore native biodiversity....

1. **[^](#cite_ref-175)** Searcy, Christopher A.; Rollins, Hilary B.; Shaffer, H. Bradley (2016). ["Ecological equivalency as a tool for endangered species management"](https://doi.org/10.1890%2F14-1674). *[Ecological Applications](/source/Ecological_Applications)*. **26** (1): 94–103. [Bibcode](/source/Bibcode_(identifier)):[2016EcoAp..26...94S](https://ui.adsabs.harvard.edu/abs/2016EcoAp..26...94S). [doi](/source/Doi_(identifier)):[10.1890/14-1674](https://doi.org/10.1890%2F14-1674). [PMID](/source/PMID_(identifier)) [27039512](https://pubmed.ncbi.nlm.nih.gov/27039512).

1. **[^](#cite_ref-176)** Hansen, Dennis M.; Donlan, C. Josh; Griffiths, Christine J.; Campbell, Karl J. (2010). ["Ecological history and latent conservation potential: Large and giant tortoises as a model for taxon substitutions"](https://doi.org/10.1111%2Fj.1600-0587.2010.06305.x). *[Ecography](/source/Ecography)*. **33** (2): 272–284. [Bibcode](/source/Bibcode_(identifier)):[2010Ecogr..33..272H](https://ui.adsabs.harvard.edu/abs/2010Ecogr..33..272H). [doi](/source/Doi_(identifier)):[10.1111/j.1600-0587.2010.06305.x](https://doi.org/10.1111%2Fj.1600-0587.2010.06305.x).

1. ^ [***a***](#cite_ref-r1_177-0) [***b***](#cite_ref-r1_177-1) Jacobsen, Rowan (March 24, 2014). ["The Invasivore's Dilemma"](https://www.outsideonline.com/1922351/invasivores-dilemma). *Outside*. [Archived](https://web.archive.org/web/20190528211949/https://www.outsideonline.com/1922351/invasivores-dilemma) from the original on May 28, 2019. Retrieved May 28, 2019.

1. **[^](#cite_ref-178)** Lai, Bun (September 1, 2013). "Invasive Species Menu of a World-Class Chef". *[Scientific American](/source/Scientific_American)*. **309** (3): 40–43. [Bibcode](/source/Bibcode_(identifier)):[2013SciAm.309c..40L](https://ui.adsabs.harvard.edu/abs/2013SciAm.309c..40L). [doi](/source/Doi_(identifier)):[10.1038/scientificamerican0913-40](https://doi.org/10.1038%2Fscientificamerican0913-40). [PMID](/source/PMID_(identifier)) [24003552](https://pubmed.ncbi.nlm.nih.gov/24003552).

1. **[^](#cite_ref-179)** Billock, Jennifer (February 9, 2016). ["Bite Back Against Invasive Species at Your Next Meal"](https://www.smithsonianmag.com/travel/indulge-invasive-species-restaurants-across-country-180957899/). *Smithsonian Magazine*. [Archived](https://web.archive.org/web/20190322193509/https://www.smithsonianmag.com/travel/indulge-invasive-species-restaurants-across-country-180957899/) from the original on March 22, 2019. Retrieved May 28, 2019.

1. **[^](#cite_ref-180)** Snyder, Michael (May 19, 2017). ["Can We Really Eat Invasive Species into Submission?"](https://www.scientificamerican.com/article/can-we-really-eat-invasive-species-into-submission/?redirect=1). *Scientific American*. [Archived](https://web.archive.org/web/20200801204428/https://www.scientificamerican.com/article/can-we-really-eat-invasive-species-into-submission/?redirect=1) from the original on August 1, 2020. Retrieved May 28, 2019.

1. **[^](#cite_ref-181)** Kolbert, Elizabeth (December 2, 2012). ["Alien Entrées"](https://www.newyorker.com/magazine/2012/12/10/alien-entrees). *New Yorker*. [Archived](https://web.archive.org/web/20191018202041/https://www.newyorker.com/magazine/2012/12/10/alien-entrees) from the original on October 18, 2019. Retrieved February 13, 2020.

1. **[^](#cite_ref-182)** ["Bio"](http://www.joeroman.com/new/bio/). *Joe Roman*. March 12, 2015. [Archived](https://web.archive.org/web/20190528213031/http://www.joeroman.com/bio/) from the original on May 28, 2019. Retrieved June 26, 2022.

1. **[^](#cite_ref-183)** ["Eat The Invaders — Fighting Invasive Species, One Bite At A Time!"](http://eattheinvaders.org/). *eattheinvaders.org*. [Archived](https://web.archive.org/web/20190519015542/http://eattheinvaders.org/) from the original on May 19, 2019. Retrieved June 26, 2022.

1. **[^](#cite_ref-184)** Parks, Mary; Thanh, Thai (2019). *The Green Crab Cookbook*. Green Crab R&d. [ISBN](/source/ISBN_(identifier)) [978-0-578-42794-2](https://en.wikipedia.org/wiki/Special:BookSources/978-0-578-42794-2).

1. **[^](#cite_ref-185)** ["Lionfish Cookbook 2nd Edition | Reef Environmental Education Foundation"](https://www.reef.org/products/lionfish-cookbook-2nd-edition). *www.reef.org*. [Archived](https://web.archive.org/web/20190528212729/https://www.reef.org/products/lionfish-cookbook-2nd-edition) from the original on May 28, 2019. Retrieved May 28, 2019.

1. **[^](#cite_ref-186)** Iyer, Ajay; Bestwick, Charles S.; Duncan, Sylvia H.; Russell, Wendy R. (February 15, 2021). ["Invasive Plants Are a Valuable Alternate Protein Source and Can Contribute to Meeting Climate Change Targets"](https://doi.org/10.3389%2Ffsufs.2021.575056). *Frontiers in Sustainable Food Systems*. **5** 575056. [Bibcode](/source/Bibcode_(identifier)):[2021FrSFS...575056I](https://ui.adsabs.harvard.edu/abs/2021FrSFS...575056I). [doi](/source/Doi_(identifier)):[10.3389/fsufs.2021.575056](https://doi.org/10.3389%2Ffsufs.2021.575056). [hdl](/source/Hdl_(identifier)):[2164/15875](https://hdl.handle.net/2164%2F15875).

1. **[^](#cite_ref-187)** Iyer, Ajay; Guerrier, Lisa; Leveque, Salomé; Bestwick, Charles S.; Duncan, Sylvia H.; Russell, Wendy R. (2022). ["High throughput method development and optimised production of leaf protein concentrates with potential to support the agri-industry"](https://link.springer.com/10.1007/s11694-021-01136-w). *Journal of Food Measurement and Characterization*. **16** (1): 49–65. [doi](/source/Doi_(identifier)):[10.1007/s11694-021-01136-w](https://doi.org/10.1007%2Fs11694-021-01136-w). [hdl](/source/Hdl_(identifier)):[2164/19275](https://hdl.handle.net/2164%2F19275). [S2CID](/source/S2CID_(identifier)) [244407388](https://api.semanticscholar.org/CorpusID:244407388).

1. **[^](#cite_ref-188)** Nuñez, Martin A.; Kuebbing, Sara; Dimarco, Romina D.; Simberloff, Daniel (December 2012). ["Invasive Species: to eat or not to eat, that is the question"](https://conbio.onlinelibrary.wiley.com/doi/10.1111/j.1755-263X.2012.00250.x). *Conservation Letters*. **5** (5): 334–341. [Bibcode](/source/Bibcode_(identifier)):[2012ConL....5..334N](https://ui.adsabs.harvard.edu/abs/2012ConL....5..334N). [doi](/source/Doi_(identifier)):[10.1111/j.1755-263X.2012.00250.x](https://doi.org/10.1111%2Fj.1755-263X.2012.00250.x). [hdl](/source/Hdl_(identifier)):[11336/198362](https://hdl.handle.net/11336%2F198362).

1. **[^](#cite_ref-189)** Conniff, Richard (January 24, 2014). ["Invasive Lionfish, the Kings of the Caribbean, May Have Met Their Match"](https://web.archive.org/web/20140127033047/https://news.yahoo.com/invasive-lionfish-kings-caribbean-may-met-match-011600208.html). *Yahoo News*. Archived from [the original](https://news.yahoo.com/invasive-lionfish-kings-caribbean-may-met-match-011600208.html) on January 27, 2014.

1. **[^](#cite_ref-190)** Bryce, Emma (February 6, 2015). ["Cooking can't solve the threat of invasive species"](https://www.theguardian.com/environment/world-on-a-plate/2015/feb/06/cooking-cant-solve-the-invasive-threat). *[The Guardian](/source/The_Guardian)*. [Archived](https://web.archive.org/web/20171017201839/https://www.theguardian.com/environment/world-on-a-plate/2015/feb/06/cooking-cant-solve-the-invasive-threat) from the original on October 17, 2017. Retrieved October 16, 2017.

1. ^ [***a***](#cite_ref-Goss2020_191-0) [***b***](#cite_ref-Goss2020_191-1) [***c***](#cite_ref-Goss2020_191-2) [***d***](#cite_ref-Goss2020_191-3) [***e***](#cite_ref-Goss2020_191-4) [***f***](#cite_ref-Goss2020_191-5) Goss, EM; Kendig, AE; Adhikari, A; Lane, B; Kortessis, N; Holt, RD; Clay, K; Harmon, PF; Flory, SL (August 2020). "Disease in Invasive Plant Populations". *Annual Review of Phytopathology*. **58** (1): 97–117. [Bibcode](/source/Bibcode_(identifier)):[2020AnRvP..58...97G](https://ui.adsabs.harvard.edu/abs/2020AnRvP..58...97G). [doi](/source/Doi_(identifier)):[10.1146/annurev-phyto-010820-012757](https://doi.org/10.1146%2Fannurev-phyto-010820-012757). [PMID](/source/PMID_(identifier)) [32516034](https://pubmed.ncbi.nlm.nih.gov/32516034). [S2CID](/source/S2CID_(identifier)) [219563975](https://api.semanticscholar.org/CorpusID:219563975).

1. **[^](#cite_ref-Kalmakoff2016_192-0)** Kalmakoff, James (October 11, 2016). ["CRISPR for pest-free NZ"](https://web.archive.org/web/20161019152322/http://www.merlinnz.com/blog/crispr-pest-free-nz/). Archived from [the original](http://www.merlinnz.com/blog/crispr-pest-free-nz/) on October 19, 2016. Retrieved October 19, 2016.

1. ^ [***a***](#cite_ref-GeneDrive_193-0) [***b***](#cite_ref-GeneDrive_193-1) ["Mission & Principles Statement"](https://genedrivenetwork.org). July 1, 2018. Retrieved November 14, 2018.

1. **[^](#cite_ref-194)** ["GBIRd Fact Sheet"](http://www.geneticbiocontrol.org/wp-content/uploads/2018/05/GBIRD-FactSheet-April-2018.pdf) (PDF). April 1, 2018. Retrieved November 14, 2018.

1. **[^](#cite_ref-195)** ["'Gene drives' could wipe out whole populations of pests in one fell swoop"](https://theconversation.com/gene-drives-could-wipe-out-whole-populations-of-pests-in-one-fell-swoop-81681). *The Conversation*. August 8, 2017.

1. **[^](#cite_ref-196)** ["An Argument Against Gene Drives to Extinguish New Zealand Mammals: Life Finds a Way"](http://blogs.plos.org/dnascience/2017/11/30/an-argument-against-gene-drives-to-extinguish-new-zealand-mammals-life-finds-a-way/). *Plos blogs*. November 30, 2017.

1. **[^](#cite_ref-NZ_gene_drive_risks_197-0)** Campbell, Colin (October 17, 2016). ["Risks may accompany gene drive technology"](https://www.odt.co.nz/opinion/risks-may-accompany-gene-drive-technology#comment-1086). [Otago Daily Times](/source/Otago_Daily_Times). Retrieved October 19, 2016.

1. **[^](#cite_ref-198)** Ööpik, Merle; Kukk, Toomas; Kull, Kalevi; Kull, Tiiu (2008). "The importance of human mediation in species establishment: analysis of the alien flora of Estonia". *Boreal Environment Research*. **13** (Supplement A): 53–67. [hdl](/source/Hdl_(identifier)):[10138/235238](https://hdl.handle.net/10138%2F235238).

1. **[^](#cite_ref-199)** Lehan, Nora E.; Murphy, Julia R.; Thorburn, Lukas P.; Bradley, Bethany A. (July 2013). "Accidental introductions are an important source of invasive plants in the continental United States". *American Journal of Botany*. **100** (7): 1287–1293. [doi](/source/Doi_(identifier)):[10.3732/ajb.1300061](https://doi.org/10.3732%2Fajb.1300061). [PMID](/source/PMID_(identifier)) [23825135](https://pubmed.ncbi.nlm.nih.gov/23825135).

1. **[^](#cite_ref-200)** Virtue, J.G.; Bennett, Sarita; Randall, R.P. (2004). "Plant introductions in Australia: how can we resolve 'weedy' conflicts of interest?: Plant introductions in Australia: how can we resolve 'weedy' conflicts of interest?". In Sindel, Brian Mark; Johnson, Stephen Barry (eds.). *Weed Management: Balancing People, Planet, Profit: 14th Australian Weeds Conference: Papers & Proceedings*. Weed Society of New South Wales. pp. 42–48. [ISBN](/source/ISBN_(identifier)) [978-0-9752488-0-5](https://en.wikipedia.org/wiki/Special:BookSources/978-0-9752488-0-5). [S2CID](/source/S2CID_(identifier)) [82300163](https://api.semanticscholar.org/CorpusID:82300163).

1. ^ [***a***](#cite_ref-:2_201-0) [***b***](#cite_ref-:2_201-1) Pheloung, P.C.; Williams, P.A.; Halloy, S.R. (December 1999). "A weed risk assessment model for use as a biosecurity tool evaluating plant introductions". *Journal of Environmental Management*. **57** (4): 239–251. [Bibcode](/source/Bibcode_(identifier)):[1999JEnvM..57..239P](https://ui.adsabs.harvard.edu/abs/1999JEnvM..57..239P). [doi](/source/Doi_(identifier)):[10.1006/jema.1999.0297](https://doi.org/10.1006%2Fjema.1999.0297).

1. ^ [***a***](#cite_ref-:3_202-0) [***b***](#cite_ref-:3_202-1) Koop, Anthony L.; Fowler, Larry; Newton, Leslie P.; Caton, Barney P. (February 2012). "Development and validation of a weed screening tool for the United States". *Biological Invasions*. **14** (2): 273–294. [Bibcode](/source/Bibcode_(identifier)):[2012BiInv..14..273K](https://ui.adsabs.harvard.edu/abs/2012BiInv..14..273K). [doi](/source/Doi_(identifier)):[10.1007/s10530-011-0061-4](https://doi.org/10.1007%2Fs10530-011-0061-4). [S2CID](/source/S2CID_(identifier)) [254280051](https://api.semanticscholar.org/CorpusID:254280051).

1. **[^](#cite_ref-203)** Pfadenhauer, William G.; Nelson, Michael F.; Laginhas, Brit B.; Bradley, Bethany A. (January 2023). ["Remember your roots: Biogeographic properties of plants' native habitats can inform invasive plant risk assessments"](https://scholarworks.umass.edu/cgi/viewcontent.cgi?article=1002&context=oeb_grad_pubs). *Diversity and Distributions*. **29** (1): 4–18. [Bibcode](/source/Bibcode_(identifier)):[2023DivDi..29....4P](https://ui.adsabs.harvard.edu/abs/2023DivDi..29....4P). [doi](/source/Doi_(identifier)):[10.1111/ddi.13639](https://doi.org/10.1111%2Fddi.13639). [S2CID](/source/S2CID_(identifier)) [253220107](https://api.semanticscholar.org/CorpusID:253220107).

1. **[^](#cite_ref-204)** Gordon, Doria R.; Flory, S. Luke; Lieurance, Deah; Hulme, Philip E.; Buddenhagen, Chris; Caton, Barney; et al. (March 2016). "Weed Risk Assessments Are an Effective Component of Invasion Risk Management". *Invasive Plant Science and Management*. **9** (1): 81–83. [Bibcode](/source/Bibcode_(identifier)):[2016IPSM....9...81G](https://ui.adsabs.harvard.edu/abs/2016IPSM....9...81G). [doi](/source/Doi_(identifier)):[10.1614/IPSM-D-15-00053.1](https://doi.org/10.1614%2FIPSM-D-15-00053.1). [hdl](/source/Hdl_(identifier)):[10019.1/116927](https://hdl.handle.net/10019.1%2F116927). [S2CID](/source/S2CID_(identifier)) [86276601](https://api.semanticscholar.org/CorpusID:86276601).

1. **[^](#cite_ref-205)** Hulme, Philip E. (February 2012). ["Weed risk assessment: a way forward or a waste of time?: Weed risk assessment: a way forward or waste of time?"](https://doi.org/10.1111%2Fj.1365-2664.2011.02069.x). *Journal of Applied Ecology*. **49** (1): 10–19. [doi](/source/Doi_(identifier)):[10.1111/j.1365-2664.2011.02069.x](https://doi.org/10.1111%2Fj.1365-2664.2011.02069.x).

1. ^ [***a***](#cite_ref-:4_206-0) [***b***](#cite_ref-:4_206-1) Bertolino, Sandro; Sciandra, Chiara; Bosso, Luciano; Russo, Danilo; Lurz, Peter W.W.; Di Febbraro, Mirko (2020). ["Spatially explicit models as tools for implementing effective management strategies for invasive alien mammals"](https://onlinelibrary.wiley.com/doi/abs/10.1111/mam.12185). *Mammal Review*. **50** (2): 187–199. [Bibcode](/source/Bibcode_(identifier)):[2020MamRv..50..187B](https://ui.adsabs.harvard.edu/abs/2020MamRv..50..187B). [doi](/source/Doi_(identifier)):[10.1111/mam.12185](https://doi.org/10.1111%2Fmam.12185). [hdl](/source/Hdl_(identifier)):[2318/1737199](https://hdl.handle.net/2318%2F1737199). [ISSN](/source/ISSN_(identifier)) [1365-2907](https://search.worldcat.org/issn/1365-2907).

1. **[^](#cite_ref-207)** Shackleton, Ross T.; Shackleton, Charlie M.; Kull, Christian A. (January 1, 2019). ["The role of invasive alien species in shaping local livelihoods and human well-being: A review"](https://www.sciencedirect.com/science/article/pii/S0301479718305267). *Journal of Environmental Management*. The human and social dimensions of invasion science and management. **229**: 145–157. [Bibcode](/source/Bibcode_(identifier)):[2019JEnvM.229..145S](https://ui.adsabs.harvard.edu/abs/2019JEnvM.229..145S). [doi](/source/Doi_(identifier)):[10.1016/j.jenvman.2018.05.007](https://doi.org/10.1016%2Fj.jenvman.2018.05.007). [hdl](/source/Hdl_(identifier)):[10019.1/122761](https://hdl.handle.net/10019.1%2F122761). [ISSN](/source/ISSN_(identifier)) [0301-4797](https://search.worldcat.org/issn/0301-4797). [PMID](/source/PMID_(identifier)) [30049620](https://pubmed.ncbi.nlm.nih.gov/30049620).

1. **[^](#cite_ref-208)** Pejchar, Liba; Mooney, Harold A. (September 1, 2009). ["Invasive species, ecosystem services and human well-being"](https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(09)00176-1). *Trends in Ecology & Evolution*. **24** (9): 497–504. [Bibcode](/source/Bibcode_(identifier)):[2009TEcoE..24..497P](https://ui.adsabs.harvard.edu/abs/2009TEcoE..24..497P). [doi](/source/Doi_(identifier)):[10.1016/j.tree.2009.03.016](https://doi.org/10.1016%2Fj.tree.2009.03.016). [ISSN](/source/ISSN_(identifier)) [0169-5347](https://search.worldcat.org/issn/0169-5347). [PMID](/source/PMID_(identifier)) [19577817](https://pubmed.ncbi.nlm.nih.gov/19577817).

1. **[^](#cite_ref-209)** Bender, Emily M.; Gebru, Timnit; McMillan-Major, Angelina; Shmitchell, Shmargaret (March 1, 2021). "On the Dangers of Stochastic Parrots: Can Language Models be Too Big? 🦜". *Proceedings of the 2021 ACM Conference on Fairness, Accountability, and Transparency*. FAccT '21. New York, NY, USA: Association for Computing Machinery. pp. 610–623. [doi](/source/Doi_(identifier)):[10.1145/3442188.3445922](https://doi.org/10.1145%2F3442188.3445922). [ISBN](/source/ISBN_(identifier)) [978-1-4503-8309-7](https://en.wikipedia.org/wiki/Special:BookSources/978-1-4503-8309-7).

1. **[^](#cite_ref-210)** Elliards, Xander (April 9, 2025). ["Hedgehogs to be evicted from Scottish island in world-first trial"](https://www.thenational.scot/news/25075393.hedgehogs-evicted-scottish-island-world-first-trial/). The National. Retrieved April 10, 2025.

1. ^ [***a***](#cite_ref-:14_211-0) [***b***](#cite_ref-:14_211-1) ["Detailed Discussion of the Ethical Treatment of Invasive Species | Animal Legal & Historical Center"](https://www.animallaw.info/article/detailed-discussion-ethical-treatment-invasive-species). *www.animallaw.info*. Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-:15_212-0) [***b***](#cite_ref-:15_212-1) Abbate, C.E.; Fischer, Bob (October 27, 2019). ["Don't Demean "Invasives": Conservation and Wrongful Species Discrimination"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6912556). *Animals*. **9** (11): 871. [doi](/source/Doi_(identifier)):[10.3390/ani9110871](https://doi.org/10.3390%2Fani9110871). [ISSN](/source/ISSN_(identifier)) [2076-2615](https://search.worldcat.org/issn/2076-2615). [PMC](/source/PMC_(identifier)) [6912556](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6912556). [PMID](/source/PMID_(identifier)) [31717868](https://pubmed.ncbi.nlm.nih.gov/31717868).

1. **[^](#cite_ref-213)** ["What Is Speciesism and How You Can Overcome It"](https://www.peta.org/features/what-is-speciesism/). *PETA*. April 9, 2019. Retrieved April 30, 2026.

1. ^ [***a***](#cite_ref-:16_214-0) [***b***](#cite_ref-:16_214-1) Bolotnikova, Marina (November 28, 2021). ["It's time to stop demonizing "invasive" species"](https://www.vox.com/down-to-earth/22796160/invasive-species-climate-change-range-shifting). *Vox*. Retrieved April 30, 2026.

[1]

## Further reading

- Mitchell, Heidi J.; Bartsch, Detlef (January 21, 2020). ["Regulation of GM Organisms for Invasive Species Control"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6985037). *Frontiers in Bioengineering and Biotechnology*. **7** 454. [doi](/source/Doi_(identifier)):[10.3389/fbioe.2019.00454](https://doi.org/10.3389%2Ffbioe.2019.00454). [PMC](/source/PMC_(identifier)) [6985037](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6985037). [PMID](/source/PMID_(identifier)) [32039172](https://pubmed.ncbi.nlm.nih.gov/32039172).

- [Removing Threat from Invasive Species with Genetic Engineering](https://sitn.hms.harvard.edu/flash/2014/removing-threat-from-invasive-species-with-genetic-engineering/)—*Science in the News*

- Sheppard, Andy; et al. (September 4, 2023). ["The true damage of invasive alien species was just revealed in a landmark report. Here's how we must act"](https://theconversation.com/the-true-damage-of-invasive-alien-species-was-just-revealed-in-a-landmark-report-heres-how-we-must-act-211893). *[The Conversation](/source/The_Conversation_(website))*.

- White, Michael (13 Dec 2017) [6 Dec 2017]. ["Should We Fight Invasive Species with Genetic Engineering?"](https://psmag.com/environment/should-we-fight-invasive-species-with-genetic-engineering) *Pacific Standard*.

## External links

- [North American Invasive Species Network](https://www.naisn.org/), a consortium that uses a coordinated network to advance science-based understanding and enhance management of non-native, invasive species.

- [Great Britain Non-native Species Secretariat (NNSS) website](https://www.nonnativespecies.org/)

- [CABI Invasive Species Compendium](https://www.cabi.org/ISC/), an encyclopaedic resource of scientific information

- [Invasive Species](https://www.invasivespeciesinfo.gov/), National Invasive Species Information Center, [United States National Agricultural Library](/source/United_States_National_Agricultural_Library)

- [Invasive Species Specialist Group](https://web.archive.org/web/20110406095350/http://issg.org/database/welcome/) – Global Invasive Species Database

- [Pacific Island Ecosystems at Risk](http://www.hear.org/pier/) project

- [invadingspecies.com](http://www.invadingspecies.com/) of the [Ontario Ministry of Natural Resources](/source/Ontario_Ministry_of_Natural_Resources) and [Ontario Federation of Anglers and Hunters](/source/Ontario_Federation_of_Anglers_and_Hunters)

- [Aquatic invasive species in Ireland](https://www.fisheriesireland.ie/what-we-do/research/research-theme-invasive-species), Inland Fisheries Ireland

- [Invasive alien species in Belgium](http://ias.biodiversity.be/) Belgian Forum on Invasive Species (BFIS)

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v t e Ecology: Modelling ecosystems: Trophic components General Abiotic component Abiotic stress Behaviour Biogeochemical cycle Biomass Biotic component Biotic stress Carrying capacity Competition Ecosystem Ecosystem ecology Ecosystem model Green world hypothesis Keystone species List of feeding behaviours Metabolic theory of ecology Productivity Resource Restoration Producers Autotrophs Chemosynthesis Chemotrophs Foundation species Kinetotrophs Mixotrophs Myco-heterotrophy Mycotroph Organotrophs Photoheterotrophs Photosynthesis Photosynthetic efficiency Phototrophs Primary nutritional groups Primary production Consumers Apex predator Bacterivore Carnivores Chemoorganotroph Foraging Generalist and specialist species Intraguild predation Herbivores Heterotroph Heterotrophic nutrition Insectivore Mesopredators Mesopredator release hypothesis Omnivores Optimal foraging theory Planktivore Predation Prey switching Decomposers Chemoorganoheterotrophy Decomposition Detritivores Detritus Microorganisms Archaea Bacteriophage Lithoautotroph Lithotrophy Marine Microbial cooperation Microbial ecology Microbial food web Microbial intelligence Microbial loop Mycoloop Microbial mat Microbial metabolism Phage ecology Food webs Biomagnification Ecological efficiency Ecological pyramid Energy flow Food chain Trophic level Example webs Lakes Rivers Soil Tritrophic interactions in plant defense Marine food webs cold seeps hydrothermal vents intertidal kelp forests North Pacific Gyre San Francisco Estuary tide pool Processes Ascendency Bioaccumulation Cascade effect Climax community Competitive exclusion principle Consumer–resource interactions Copiotrophs Dominance Ecological network Ecological succession Energy quality Energy systems language f-ratio Feed conversion ratio Feeding frenzy Mesotrophic soil Nutrient cycle Oligotroph Paradox of the plankton Trophic cascade Trophic mutualism Trophic state index Defense, counter Animal coloration Anti-predator adaptations Camouflage Deimatic behaviour Herbivore adaptations to plant defense Mimicry Plant defense against herbivory Predator avoidance in schooling fish

v t e Ecology: Modelling ecosystems: Other components Population ecology Abundance Allee effect Consumer-resource model Depensation Ecological yield Effective population size Intraspecific competition Logistic function Malthusian growth model Maximum sustainable yield Overpopulation Overexploitation Population cycle Population dynamics Population modeling Population size Predator–prey (Lotka–Volterra) equations Recruitment Small population size Stability Resilience Resistance Random generalized Lotka–Volterra model Species Biodiversity Density-dependent inhibition Ecological effects of biodiversity Ecological extinction Endemic species Flagship species Gradient analysis Indicator species Introduced species Invasive species / Native species Latitudinal gradients in species diversity Minimum viable population Neutral theory Occupancy–abundance relationship Population viability analysis Priority effect Rapoport's rule Relative abundance distribution Relative species abundance Species diversity Species homogeneity Species richness Species distribution Species–area curve Umbrella species Species interaction Antibiosis Biological interaction Commensalism Community ecology Ecological facilitation Interspecific competition Mutualism Parasitism Storage effect Symbiosis Spatial ecology Biogeography Cross-boundary subsidy Ecocline Ecotone Ecotype Disturbance Edge effects Foster's rule Habitat fragmentation Ideal free distribution Intermediate disturbance hypothesis Insular biogeography Land change modeling Landscape ecology Landscape epidemiology Landscape limnology Metapopulation Patch dynamics r/K selection theory Resource selection function Source–sink dynamics Niche Ecological trap Ecosystem engineer Environmental niche modelling Guild Habitat Marine Semiaquatic Terrestrial Limiting similarity Niche apportionment models Niche construction Niche differentiation Ontogenetic niche shift Other networks Assembly rules Bateman's principle Bioluminescence Ecological collapse Ecological debt Ecological deficit Ecological energetics Ecological indicator Ecological threshold Ecosystem diversity Emergence Extinction debt Kleiber's law Liebig's law of the minimum Marginal value theorem Thorson's rule Xerosere Other Allometry Alternative stable state Balance of nature Biological data visualization Ecological economics Ecological footprint Ecological forecasting Ecological humanities Ecological stoichiometry Ecopath Ecosystem based fisheries Endolith Evolutionary ecology Functional ecology Industrial ecology Macroecology Microecosystem Natural environment Regime shift Sexecology Systems ecology Urban ecology Theoretical ecology Outline of ecology

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Authority control databases International GND National United States Japan Latvia

1. **[^](#cite_ref-:17_215-0)** Cite error: The named reference :17 was invoked but never defined (see the [help page](https://en.wikipedia.org/wiki/Help:Cite_errors/Cite_error_references_no_text)).

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Adapted from the Wikipedia article [Invasive species](https://en.wikipedia.org/wiki/Invasive_species) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Invasive_species?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.