# Hyperolius

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{{short description|Genus of frogs}}
{{automatic taxobox
| image = Hyperolius argus Al-ReedFrogs 02.jpg
| image_caption = ''[Hyperolius argus](/source/Hyperolius_argus)''
| taxon = Hyperolius
| subdivision_ranks = Species
| subdivision = 145 species (but see text)
| authority = {{interlanguage link|Wilhelm Ludwig Rapp|fr|Wilhelm Ludwig Rapp|de|Wilhelm von Rapp|lt=Rapp}}, 1842
| type_species = ''[Hyla horstocki](/source/Hyperolius_horstockii)''
| type_species_authority = [Schlegel](/source/Hermann_Schlegel), 1837
| synonyms = {{genus list
 | Eucnemis | Tschudi, 1838 – [homonym](/source/Homonym_(biology)) of ''Eucnemis'' Ahrens, 1812 (Insecta)
 | Hyperolius | Rapp, 1842 – replacement name
 | Epipole | Gistel, 1848
 | Crumenifera | Cope, 1862
 | Rappia | Günther, 1865
 | Nesionixalus | Perret, 1976
 | Chlorolius | Perret, 1988
 | Alexteroon | Perret, 1988 }}
| synonyms_ref = <ref name="Frost-2022"/>
}}

'''''Hyperolius''''' (commonly known as the '''African reed frogs''' or '''reed frogs''') is a large [genus](/source/genus) of [frog](/source/frog)s in the family [Hyperoliidae](/source/Hyperoliidae) from [Sub-Saharan Africa](/source/Sub-Saharan_Africa).<ref name="Frost-2022"/><ref name="Channing and Rödel 2019"/> The genus is known to exhibit [sexual dichromatism](/source/sexual_dichromatism), a trait that is otherwise rare in frogs.<ref name="Portik-2019">{{cite web |last1=Portik |first1=D.M. |title=Sexual Dichromatism Drives Diversification within a Major Radiation of African Amphibians |url=https://academic.oup.com/sysbio/article/68/6/859/5477408|journal=Systematic Biology |volume=68 |issue=6 |date= November 2019 |pages= 859–875 |doi=10.1093/sysbio/syz023}}</ref>

==Species==
Different sources may delimit species differently, and as new species are still being described, different number of species can be found. {{As of|March 2026}}, ''Amphibian Species of the World'' lists 145 species<ref name="Frost-2022"/> and ''AmphibiaWeb'' 155 species.<ref name=AmphibiaWeb>{{cite web |url=https://amphibiaweb.org/lists/Hyperoliidae.shtml |title=Hyperoliidae |year=2022 |work=AmphibiaWeb: Information on amphibian biology and conservation. [web application] |publisher=Berkeley, California: AmphibiaWeb |access-date=2026-03-01}}</ref> The following list follows the ''Amphibian Species of the World'':<ref name="Frost-2022"/>
{{columns-list|colwidth=25em|
* ''[Hyperolius acuticephalus](/source/Hyperolius_acuticephalus)'' <small>Ahl, 1931</small>
* ''[Hyperolius acutirostris](/source/Hyperolius_acutirostris)'' <small>Buchholz and Peters, 1875</small>
* ''[Hyperolius ademetzi](/source/Hyperolius_ademetzi)'' <small>Ahl, 1931</small>
* ''[Hyperolius adspersus](/source/Hyperolius_adspersus)'' <small>Peters, 1877</small>
* ''[Hyperolius albofrenatus](/source/Hyperolius_albofrenatus)'' <small>Ahl, 1931</small>
* ''[Hyperolius argus](/source/Hyperolius_argus)'' <small>Peters, 1854</small>
* ''[Hyperolius atrigularis](/source/Hyperolius_atrigularis)'' <small>Laurent, 1941</small>
* ''[Hyperolius balfouri](/source/Hyperolius_balfouri)'' <small>(Werner, 1908)</small>
* ''[Hyperolius baumanni](/source/Hyperolius_baumanni)'' <small>Ahl, 1931</small>
* ''[Hyperolius benguellensis](/source/Hyperolius_benguellensis)'' <small>(Bocage, 1893)</small>
* ''[Hyperolius bicolor](/source/Hyperolius_bicolor)'' <small>Ahl, 1931</small>
* ''[Hyperolius bobirensis](/source/Hyperolius_bobirensis)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius bocagei](/source/Hyperolius_bocagei)'' <small>Steindachner, 1867</small>
* ''[Hyperolius bolifambae](/source/Hyperolius_bolifambae)'' <small>Mertens, 1938</small>
* ''[Hyperolius bopeleti](/source/Hyperolius_bopeleti)'' <small>Amiet, 1980</small>
* ''[Hyperolius brachiofasciatus](/source/Hyperolius_brachiofasciatus)'' <small>Ahl, 1931</small>
* ''[Hyperolius burgessi](/source/Hyperolius_burgessi)'' <small>Loader, Lawson, Portik, and Menegon, 2015</small>
* ''[Hyperolius camerunensis](/source/Hyperolius_camerunensis)'' <small>Amiet, 2004</small>
* ''[Hyperolius castaneus](/source/Hyperolius_castaneus)'' <small>Ahl, 1931</small>
* ''[Hyperolius chelaensis](/source/Hyperolius_chelaensis)'' <small>Conradie, Branch, Measey, and Tolley, 2012</small>
* ''[Hyperolius chlorosteus](/source/Hyperolius_chlorosteus)'' <small>(Boulenger, 1915)</small>
* ''[Hyperolius chrysogaster](/source/Hyperolius_chrysogaster)'' <small>Laurent, 1950</small>
* ''[Hyperolius cinereus](/source/Hyperolius_cinereus)'' <small>Monard, 1937</small>
* ''[Hyperolius cinnamomeoventris](/source/Hyperolius_cinnamomeoventris)'' <small>Bocage, 1866</small>
* ''[Hyperolius concolor](/source/Hyperolius_concolor)'' <small>(Hallowell, 1844)</small>
* ''[Hyperolius constellatus](/source/Hyperolius_constellatus)'' <small>Laurent, 1951</small>
* ''[Hyperolius cystocandicans](/source/Hyperolius_cystocandicans)'' <small>Richards and Schiøtz, 1977</small>
* ''[Hyperolius dartevellei](/source/Hyperolius_dartevellei)'' <small>Laurent, 1943</small>
* ''[Hyperolius davenporti](/source/Hyperolius_davenporti)'' <small>Loader, Lawson, Portik, and Menegon, 2015</small>
* ''[Hyperolius diaphanus](/source/Hyperolius_diaphanus)'' <small>Laurent, 1972</small>
* ''[Hyperolius dintelmanni](/source/Hyperolius_dintelmanni)'' <small>Lötters and Schmitz, 2004</small>
* ''[Hyperolius discodactylus](/source/Hyperolius_discodactylus)'' <small>Ahl, 1931</small>
* ''[Hyperolius drewesi](/source/Hyperolius_drewesi)'' <small>Bell, 2016</small>
* ''[Hyperolius endjami](/source/Hyperolius_endjami)'' <small>Amiet, 1980</small>
* ''[Hyperolius ferrugineus](/source/Hyperolius_ferrugineus)'' <small>Laurent, 1943</small>
* ''[Hyperolius friedemanni](/source/Hyperolius_friedemanni)'' <small>Mercurio and Rödel ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius frontalis](/source/Hyperolius_frontalis)'' <small>Laurent, 1950</small>
* ''[Hyperolius fuscigula](/source/Hyperolius_fuscigula)'' <small>Bocage, 1866</small>
* ''[Hyperolius fusciventris](/source/Hyperolius_fusciventris)'' <small>Peters, 1876</small>
* ''[Hyperolius ghesquieri](/source/Hyperolius_ghesquieri)'' <small>Laurent, 1943</small>
* ''[Hyperolius glandicolor](/source/Hyperolius_glandicolor)'' <small>Peters, 1878</small>
* ''[Hyperolius gularis](/source/Hyperolius_gularis)'' <small>Ahl, 1931</small>
* ''[Hyperolius guttulatus](/source/Hyperolius_guttulatus)'' <small>Günther, 1858</small>
* ''[Hyperolius horstockii](/source/Hyperolius_horstockii)'' <small>(Schlegel, 1837)</small>
* ''[Hyperolius houyi](/source/Hyperolius_houyi)'' <small>Ahl, 1931</small>
* ''[Hyperolius howelli](/source/Hyperolius_howelli)'' <small>Du Preez and Channing ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius hutsebauti](/source/Hyperolius_hutsebauti)'' <small>Laurent, 1956</small>
* ''[Hyperolius hypsiphonus](/source/Hyperolius_hypsiphonus)'' <small>(Amiet, 2000)</small>
* ''[Hyperolius igbettensis](/source/Hyperolius_igbettensis)'' <small>Schiøtz, 1963</small>
* ''[Hyperolius inornatus](/source/Hyperolius_inornatus)'' <small>Laurent, 1943</small>
* ''[Hyperolius inyangae](/source/Hyperolius_inyangae)'' <small>Channing ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius jackie](/source/Hyperolius_jackie)'' <small>Dehling, 2012</small>
* ''[Hyperolius jacobseni](/source/Hyperolius_jacobseni)'' <small>Channing ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius jynx](/source/Hyperolius_jynx)'' <small>(Amiet, 2000)</small>
* ''[Hyperolius kachalolae](/source/Hyperolius_kachalolae)'' <small>Schiøtz, 1975</small>
* ''[Hyperolius kibarae](/source/Hyperolius_kibarae)'' <small>Laurent, 1957</small>
* ''[Hyperolius kihangensis](/source/Hyperolius_kihangensis)'' <small>Schiøtz and Westergaard, 1999</small>
* ''[Hyperolius kivuensis](/source/Hyperolius_kivuensis)'' <small>Ahl, 1931</small>
* ''[Hyperolius koehleri](/source/Hyperolius_koehleri)'' <small>Mertens, 1940</small>
* ''[Hyperolius kuligae](/source/Hyperolius_kuligae)'' <small>Mertens, 1940</small>
* ''[Hyperolius lamottei](/source/Hyperolius_lamottei)'' <small>Laurent, 1958</small>
* ''[Hyperolius langi](/source/Hyperolius_langi)'' <small>Noble, 1924</small>
* ''[Hyperolius lateralis](/source/Hyperolius_lateralis)'' <small>Laurent, 1940</small>
* ''[Hyperolius laurenti](/source/Hyperolius_laurenti)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius leleupi](/source/Hyperolius_leleupi)'' <small>Laurent, 1951</small>
* ''[Hyperolius leucotaenius](/source/Hyperolius_leucotaenius)'' <small>Laurent, 1950</small>
* ''[Hyperolius lucani](/source/Hyperolius_lucani)'' <small>Rochebrune, 1885</small>
* ''[Hyperolius lupiroensis](/source/Hyperolius_lupiroensis)'' <small>Channing ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius maestus](/source/Hyperolius_maestus)'' <small>Rochebrune, 1885</small>
* ''[Hyperolius major](/source/Hyperolius_major)'' <small>Laurent, 1957</small>
* ''[Hyperolius marginatus](/source/Hyperolius_marginatus)'' <small>Peters, 1854</small>
* ''[Hyperolius mariae](/source/Hyperolius_mariae)'' <small>Barbour and Loveridge, 1928</small>
* ''[Hyperolius marmoratus](/source/Hyperolius_marmoratus)'' <small>Rapp, 1842</small>
* ''[Hyperolius microps](/source/Hyperolius_microps)'' <small>Günther, 1864</small>
* ''[Hyperolius minutissimus](/source/Hyperolius_minutissimus)'' <small>Schiøtz, 1975</small>
* ''[Hyperolius mitchelli](/source/Hyperolius_mitchelli)'' <small>Loveridge, 1953</small>
* ''[Hyperolius molleri](/source/Hyperolius_molleri)'' <small>(Bedriaga, 1892)</small>
* ''[Hyperolius montanus](/source/Hyperolius_montanus)'' <small>(Angel, 1924)</small>
* ''[Hyperolius mosaicus](/source/Hyperolius_mosaicus)'' <small>Perret, 1959</small>
* ''[Hyperolius nasicus](/source/Hyperolius_nasicus)'' <small>Laurent, 1943</small>
* ''[Hyperolius nasutus](/source/Hyperolius_nasutus)'' <small>Günther, 1865</small>
* ''[Hyperolius nienokouensis](/source/Hyperolius_nienokouensis)'' <small>Rödel, 1998</small>
* ''[Hyperolius nimbae](/source/Hyperolius_nimbae)'' <small>Laurent, 1958</small>
* ''[Hyperolius nitidulus](/source/Hyperolius_nitidulus)'' <small>Peters, 1875</small>
* ''[Hyperolius obscurus](/source/Hyperolius_obscurus)'' <small>Laurent, 1943</small>
* ''[Hyperolius obstetricans](/source/Hyperolius_obstetricans)'' <small>(Ahl, 1931)</small>
* ''[Hyperolius occidentalis](/source/Hyperolius_occidentalis)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius ocellatus](/source/Hyperolius_ocellatus)'' <small>Günther, 1858</small>
* ''[Hyperolius olivaceus](/source/Hyperolius_olivaceus)'' <small>Buchholz and Peters, 1876</small>
* ''[Hyperolius papyri](/source/Hyperolius_papyri)'' <small>(Werner, 1908 "1907")</small>
* ''[Hyperolius parallelus](/source/Hyperolius_parallelus)'' <small>Günther, 1858</small>
* ''[Hyperolius pardalis](/source/Hyperolius_pardalis)'' <small>Laurent, 1948</small>
* ''[Hyperolius parkeri](/source/Hyperolius_parkeri)'' <small>Loveridge, 1933</small>
* ''[Hyperolius phantasticus](/source/Hyperolius_phantasticus)'' <small>(Boulenger, 1899)</small>
* ''[Hyperolius pickersgilli](/source/Hyperolius_pickersgilli)'' <small>Raw, 1982</small>
* ''[Hyperolius picturatus](/source/Hyperolius_picturatus)'' <small>Peters, 1875</small>
* ''[Hyperolius pictus](/source/Hyperolius_pictus)'' <small>Ahl, 1931</small>
* ''[Hyperolius platyceps](/source/Hyperolius_platyceps)'' <small>(Boulenger, 1900)</small>
* ''[Hyperolius polli](/source/Hyperolius_polli)'' <small>Laurent, 1943</small>
* ''[Hyperolius polystictus](/source/Hyperolius_polystictus)'' <small>Laurent, 1943</small>
* ''[Hyperolius poweri](/source/Hyperolius_poweri)'' <small>Loveridge, 1938</small>
* ''[Hyperolius protchei](/source/Hyperolius_protchei)'' <small>Rochebrune, 1885</small>
* ''[Hyperolius pseudargus](/source/Hyperolius_pseudargus)'' <small>Schiøtz and Westergaard, 1999</small>
* ''[Hyperolius puncticulatus](/source/Hyperolius_puncticulatus)'' <small>(Pfeffer, 1893)</small>
* ''[Hyperolius pusillus](/source/Hyperolius_pusillus)'' <small>(Cope, 1862)</small>
* ''[Hyperolius pustulifer](/source/Hyperolius_pustulifer)'' <small>Laurent, 1940</small>
* ''[Hyperolius pyrrhodictyon](/source/Hyperolius_pyrrhodictyon)'' <small>Laurent, 1965</small>
* ''[Hyperolius quadratomaculatus](/source/Hyperolius_quadratomaculatus)'' <small>Ahl, 1931</small>
* ''[Hyperolius quinquevittatus](/source/Hyperolius_quinquevittatus)'' <small>Bocage, 1866</small>
* ''[Hyperolius raymondi](/source/Hyperolius_raymondi)'' <small>Conradie, Branch, and Tolley, 2013</small>
* ''[Hyperolius rhizophilus](/source/Hyperolius_rhizophilus)'' <small>Rochebrune, 1885</small>
* ''[Hyperolius rhodesianus](/source/Hyperolius_rhodesianus)'' <small>Laurent, 1948</small>
* ''[Hyperolius riggenbachi](/source/Hyperolius_riggenbachi)'' <small>(Nieden, 1910)</small>
* ''[Hyperolius rubrovermiculatus](/source/Hyperolius_rubrovermiculatus)'' <small>Schiøtz, 1975</small>
* ''[Hyperolius ruvuensis](/source/Hyperolius_ruvuensis)'' <small>Barratt, Lawson and Loader, 2017</small>
* ''[Hyperolius rwandae](/source/Hyperolius_rwandae)'' <small>Dehling, Sinsch, Rodel, and Channing ''in'' Channing, Hillers, Lötters, Rödel, Schick, Conradie, Rödder, Mercurio, Wagner, Dehling, Du Preez, Kielgast, and Burger, 2013</small>
* ''[Hyperolius sankuruensis](/source/Hyperolius_sankuruensis)'' <small>Laurent, 1979</small>
* ''[Hyperolius schoutedeni](/source/Hyperolius_schoutedeni)'' <small>Laurent, 1943</small>
* ''[Hyperolius semidiscus](/source/Hyperolius_semidiscus)'' <small>Hewitt, 1927</small>
* ''[Hyperolius sheldricki](/source/Hyperolius_sheldricki)'' <small>Duff-MacKay and Schiøtz, 1971</small>
* ''[Hyperolius soror](/source/Hyperolius_soror)'' <small>(Chabanaud, 1921)</small>
* ''[Hyperolius spatzi](/source/Hyperolius_spatzi)'' <small>Ahl, 1931</small>
* ''[Hyperolius spinigularis](/source/Hyperolius_spinigularis)'' <small>Stevens, 1971</small>
* ''[Hyperolius steindachneri](/source/Hyperolius_steindachneri)'' <small>Bocage, 1866</small>
* ''[Hyperolius stenodactylus](/source/Hyperolius_stenodactylus)'' <small>Ahl, 1931</small>
* ''[Hyperolius stictus](/source/Hyperolius_stictus)'' <small>Conradie, Verburgt, Portik, Ohler, Bwong, and Lawson, 2018</small>
* ''[Hyperolius substriatus](/source/Hyperolius_substriatus)'' <small>Ahl, 1931</small>
* ''[Hyperolius swynnertoni](/source/Hyperolius_swynnertoni)'' <small>FitzSimons, 1941</small>
* ''[Hyperolius sylvaticus](/source/Hyperolius_sylvaticus)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius tanneri](/source/Hyperolius_tanneri)'' <small>Schiøtz, 1982</small>
* ''[Hyperolius thomensis](/source/Hyperolius_thomensis)'' <small>Bocage, 1886</small>
* ''[Hyperolius tornieri](/source/Hyperolius_tornieri)'' <small>Ahl, 1931</small>
* ''[Hyperolius torrentis](/source/Hyperolius_torrentis)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius tuberculatus](/source/Hyperolius_tuberculatus)'' <small>(Mocquard, 1897)</small>
* ''[Hyperolius tuberilinguis](/source/Hyperolius_tuberilinguis)'' <small>Smith, 1849</small>
* ''[Hyperolius ukwiva](/source/Hyperolius_ukwiva)'' <small>Loader, Lawson, Portik, and Menegon, 2015</small>
* ''[Hyperolius veithi](/source/Hyperolius_veithi)'' <small>Schick, Kielgast, Rödder, Muchai, Burger, and Lötters, 2010</small>
* ''[Hyperolius vilhenai](/source/Hyperolius_vilhenai)'' <small>Laurent, 1964</small>
* ''[Hyperolius viridiflavus](/source/Hyperolius_viridiflavus)'' <small>(Duméril and Bibron, 1841)</small>
* ''[Hyperolius viridigulosus](/source/Hyperolius_viridigulosus)'' <small>Schiøtz, 1967</small>
* ''[Hyperolius viridis](/source/Hyperolius_viridis)'' <small>Schiøtz, 1975</small>
* ''[Hyperolius watsonae](/source/Hyperolius_watsonae)'' <small>Pickersgill, 2007</small>
* ''[Hyperolius wermuthi](/source/Hyperolius_wermuthi)'' <small>Laurent, 1961</small>
* ''[Hyperolius xenorhinus](/source/Hyperolius_xenorhinus)'' <small>Laurent, 1972</small>
* ''[Hyperolius zonatus](/source/Hyperolius_zonatus)'' <small>Laurent, 1958</small>
}}

==Nomina inquirenda==
The following species are considered ''[nomina inquirenda](/source/species_inquirenda)'' (species of doubtful identity):<ref name="Frost-2022"/>
* ''Rappia granulata'' <small>Tornier, 1896</small>
* ''Rappia fimbriata'' <small>Tornier, 1896</small>
* ''[Hyperolius laticeps](/source/Hyperolius_laticeps)'' <small>Ahl, 1931</small>
* ''[Hyperolius thoracotuberculatus](/source/Hyperolius_thoracotuberculatus)'' <small>Ahl, 1931</small>

==References==
{{Reflist|30em|refs=
<ref name="Channing and Rödel 2019">{{cite book |title=Field Guide to the Frogs & other Amphibians of Africa |last1=Channing |first1=Allan |last2=Rödel |first2=Mark-Oliver |name-list-style=amp |year=2019 |publisher=Struik Nature |location=Cape Town |isbn=978-1-77584-512-6 |pages=170–209 }}</ref>

<ref name="Frost-2022">{{cite web |url=https://amphibiansoftheworld.amnh.org/Amphibia/Anura/Hyperoliidae/Hyperoliinae/Hyperolius |title=''Hyperolius'' Rapp, 1842 |author=Frost, Darrel R. |year=2022 |work=Amphibian Species of the World: An Online Reference. Version 6.1 |publisher=American Museum of Natural History |doi=10.5531/db.vz.0001 |access-date=2026-03-01}}</ref>

}}

{{Taxonbar|from=Q2405655|from2=Q622396}}

Category:Hyperolius
Category:Hyperoliidae
Category:Amphibians of Sub-Saharan Africa
Category:Amphibian genera
Category:Taxonomy articles created by Polbot
Category:Long stubs with short prose

{{Hyperoliidae-stub}}

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Adapted from the Wikipedia article [Hyperolius](https://en.wikipedia.org/wiki/Hyperolius) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Hyperolius?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.