# Humbertium

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***Humbertium*** is a genus of [land planarians](/source/Land_planarian) of the subfamily [Bipaliinae](/source/Bipaliinae) (hammerhead flatworms).

## Description

Species of *Humbertium* are characterized by the presence of a well developed penis papilla in the [copulatory apparatus](/source/Reproductive_system_of_planarians), similar to the one in *[Bipalium](/source/Bipalium)*, but with the ovovitelloducts entering the female atrium anteriorly and not posteriorly as in *Bipalium*.[1]

## Etymology

The genus *Humbertium* is named after the Swiss naturalist [Aloïs Humbert](/source/Alo%C3%AFs_Humbert) who described several species from [Sri Lanka](/source/Sri_Lanka) that are now classified in the genus.[1]

## Species

The genus *Humbertium* contains the following species:[2]

- *[Humbertium core](/source/Humbertium_core)* (de Beauchamp, 1930)
- *[Humbertium depressum](/source/Humbertium_depressum)* (Ritter-Záhony, 1905)
- *[Humbertium diana](/source/Humbertium_diana)* (Humbert, 1862)
- *[Humbertium dodabettae](/source/Humbertium_dodabettae)* (de Beauchamp, 1930)
- *[Humbertium ferruginoideum](/source/Humbertium_ferruginoideum)* (Sabussowa, 1925)
- *[Humbertium interruptum](/source/Humbertium_interruptum)* (Graff, 1899)[3]
- *[Humbertium ithorense](/source/Humbertium_ithorense)* Solà & Sluys, 2023[3]
- *[Humbertium kelleri](/source/Humbertium_kelleri)* (von Graff, 1899)
- *[Humbertium longicanale](/source/Humbertium_longicanale)* (Sabussowa, 1925)
- *[Humbertium palnisium](/source/Humbertium_palnisium)* (de Beauchamp, 1930)
- *[Humbertium penangense](/source/Humbertium_penangense)* (Kawakatsu, 1986)
- *[Humbertium penrissenense](/source/Humbertium_penrissenense)* (de Beauchamp, 1925)
- *[Humbertium penzigi](/source/Humbertium_penzigi)* (Müller, 1902)[3]
- *[Humbertium phebe](/source/Humbertium_phebe)* (Humbert, 1862)
- *[Humbertium proserpina](/source/Humbertium_proserpina)* (Humbert, 1862)
- *[Humbertium ravenalae](/source/Humbertium_ravenalae)* (von Graff, 1899)
- *[Humbertium subboreale](/source/Humbertium_subboreale)* (Sabussowa, 1925)
- *[Humbertium umbrinum](/source/Humbertium_umbrinum)* (Geba, 1909)
- *[Humbertium voigti](/source/Humbertium_voigti)* (von Graff, 1899)
- *[Humbertium woodworthi](/source/Humbertium_woodworthi)* (von Graff, 1899)

## References

1. Ogren, R. E. & Sluys, R. (2001). "The genus *Humbertium* gen. nov., a new taxon of the land planarian family Bipaliidae (Tricladida, Terricola)". *Belgian Journal of Zoology*. **131** (1): 201–204.

1. ["WoRMS - World Register of Marine Species - Humbertium Ogren & Sluys, 2001"](https://www.marinespecies.org/aphia.php?p=taxdetails&id=479769). *www.marinespecies.org*. Retrieved 2021-10-28.

1. Solà, E.; Sluys, R.; Riutort, M.; Kawakatsu, M. (18 August 2023). "Molecular phylogenetics facilitates the first historical biogeographic analysis of the hammerhead worms (Platyhelminthes: Tricladida: Bipaliinae), with the description of twelve new species and two new genera". *Zootaxa*. **5335** (1): 1–77. [doi:10.11646/zootaxa.5335.1.1](https://doi.org/10.11646/zootaxa.5335.1.1). [ISSN 1175-5334](https://www.worldcat.org/issn/1175-5334). [PMID 38221113](https://pubmed.ncbi.nlm.nih.gov/38221113)

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Adapted from the Wikipedia article [Humbertium](https://en.wikipedia.org/wiki/Humbertium) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Humbertium?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.