# Holyoakia

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> Source: https://en.wikipedia.org/wiki/Holyoakia
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{{Short description|Genus of trilobites}}
{{Automatic taxobox
| fossil_range = late [Botomian](/source/Botomian)<br />~{{fossil range|518|516}}
| image = Holyoakia simpsoni.jpg
| image_caption = ''Holyoakia simpsoni'', © Dave Simpson
| taxon = Holyoakia
| authority = Palmer ''in'' Palmer & Rowell, 1995
| subdivision_ranks = Species
| subdivision = 
* ''Holyoakia granulosa'' <small>Palmer ''in'' Palmer & Rowell, 1995 ([Type](/source/Type_(biology)))</small>
* ''Holyoakia simpsoni'' <small>Paterson & Jago, 2006</small>
}}

'''''Holyoakia''''' is a genus of very small (up to 7½ mm long) [trilobite](/source/trilobite)s of the family [Dorypygidae](/source/Dorypygidae), from the late Lower [Cambrian](/source/Cambrian) (Late Botonian) of South Australia and Antarctica.<ref name="Paterson&Jago"/>

== Etymology ==
''Holyoakia simpsoni'' was named in honor of David Simpson from Adelaide, who collected the first specimens.<ref name="Paterson&Jago"/>

== Distribution ==
* ''Holyoakia granulosa'' has only been collected from the late Lower Cambrian of Antarctica (presumably Botomian, [Shackleton Limestone](/source/Shackleton_Limestone), Central Transantarctic Mountains)
* ''Holyoakia simpsoni'' only occurs in the late Lower Cambrian of South Australia (mid-late Botomian, [Emu Bay Shale](/source/Emu_Bay_Shale), Big Gully, Kangaroo Island).

== Taxonomy ==
The genus was originally with reservation placed in the [Emuellidae](/source/Emuellidae). The genus however lacks the specific characteristics of this family, most notably, the clearly furrowed [glabella](/source/Trilobite) that is rounded at the frontal end, the hook-shaped eye ridges that run parallel to the margin of the head shield (or [cephalon](/source/Trilobite)), clearly distinguishable prothorax of six segments, the most backward one with much larger spines than all others, and opistothorax of many segments, while the pygidium is tiny. ''Holyoakia'' however, has a rectangular glabella, difficult to discern eye ridges, a thorax of eight segments which is not differentiated and has no macropleural segment, and the pygidium is not much smaller than the cephalon.<ref name="Paterson&Jago">{{cite journal|last1= Paterson|first1= R.J.|last2= Jago|first2=J.B.|year= 2006|title= New trilobites from the Lower Cambrian Emu Bay Shale Lagerstätte at Big Gully, Kangaroo Island, South Australia.|journal= Memoirs of the Association of Australasian Palaeontologists|volume= 32|pages= 43–57|hdl= 1959.14/10651|issn= 0810-8889}}</ref>

== Ecology ==
''Holyoakia simpsoni'' occurs together with ''[Redlichia takooensis](/source/Redlichia_takooensis)'', ''[Estaingia bilobata](/source/Estaingia_bilobata)'', ''[Balcoracania dailyi](/source/Balcoracania)'', and ''[Megapharanaspis nedini](/source/Megapharanaspis)''.

== Key to the species ==
{|
|- valign="top"  align="left"
! scope="row" width="3%" | '''1''' 
|Pygidium subtriangular, about 1½× as wide as long. Pygidial axis with 8 discernible rings. 6 pairs of pygidial pleura terminate in well developed spines.<br />→ '''''Holyoakia granulosa'''''

|- valign="top" align="left"
! scope="row" width="3%" | '''-''' 
|Pygidium transversely elliptical, more than 2× as wide as long. Pygidial axis with 5 discernible rings. 4 pairs of pygidial pleural furrows end in nodes.<br />→ '''''Holyoakia simpsoni'''''<ref name="Paterson&Jago"/>
|}

== References ==
{{Portal|Paleontology}}
{{Reflist}}

== Further reading ==
<!--- Note: to edit Australian Trilobite References go to the Template:Australian Trilobite References template --->
{{refbegin}}
* {{cite book
 | chapter = Burgess Shale-type Localities: The global picture
 | author = Hagadorn, J.W.
 | editor = Bottjer, D.J. |editor2=W. Etter |editor3=J.W. Hagadorn |editor4=C.M. Tang
 | title=Exceptional Fossil Preservation -- A Unique View on the Evolution of Marine Life
 | publisher=Columbia University Press 
 | year=2002
 | isbn =0231102542
 }}
* {{cite web
 |author1=Greg Edgecombe |website=The Australian Museum 
 | title=Australian Trilobites: A Species List and Bibliography
 | url=http://www.austmus.gov.au/palaeontology/research/trilobites03.htm?
 | accessdate=August 23, 2005
 }}
* {{cite web
 | author=Sam Gon III
 | title=A guide to the Orders of Trilobites
 | url=http://www.trilobites.info/index.htm
 | accessdate=August 23, 2005
 }}
* {{cite journal
 | author=Nedin, C.
 | title=The Emu Bay Shale, a Lower Cambrian fossil Lagerstätte, Kangaroo Island, South Australia 
 | journal=Memoirs of the Association of Australasian Palaeontologists
 | year=1995 | volume=18 | pages=31–40
 }}
* {{cite web
 |author      = Simpson, Dave
 |title       = Trilobites of South Australia
 |url         = http://info.esc.net.au/~dasimpson/index.htm
 |accessdate  = August 23, 2005
 |url-status     = dead
 |archiveurl  = https://web.archive.org/web/20060219192556/http://info.esc.net.au/~dasimpson/index.htm
 |archivedate = February 19, 2006
}}
{{refend}}

{{Taxonbar|from=Q5887050}}

Category:Corynexochida genera
Category:Dorypygidae
Category:Cambrian trilobites
Category:Trilobites of Antarctica
Category:Trilobites of Australia
Category:Fossil taxa described in 1995

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Adapted from the Wikipedia article [Holyoakia](https://en.wikipedia.org/wiki/Holyoakia) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Holyoakia?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.
