# Heterosaccus > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Heterosaccus > Markdown URL: https://mediated.wiki/source/Heterosaccus.md > Source: https://en.wikipedia.org/wiki/Heterosaccus > Source revision: 1342269521 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) ***Heterosaccus*** is a genus of [barnacles](/source/Barnacles) in infraclass [Rhizocephala](/source/Rhizocephala). Like other taxa in this group, they [parasitize](/source/Parasitize) crabs. Geoffroy Smith circumscribed the genus in 1906; he initially only included *H. hians*. Smith circumscribed a genus distinct from *[Sacculina](/source/Sacculina)* due to a difference of the [mesentery](/source/Mesentery); in *Heterosaccus*, the mesentery does not stretch down to the mantle opening but rather only is present on the ring of attachment.[1] ## Species As of 2017, [WoRMS](/source/WoRMS) recognizes the following fifteen species.[2] Binominal name (Original combination) Author citation Type host Type locality Ref Heterosaccus californicus Boschma, 1933 "Pugettia producta" "Santa Cruz, California" [3]: 236–237 Heterosaccus distortus Boschma, 1933 "Schizophrys aspera" "vicinity of Jolo," Philippines [3]: 235–236 Heterosaccus dollfusi Boschma, 1960 "Charybdis hoplites" "Gulf of Suez" [4] Heterosaccus gongylus Boschma, 1962 Pleistacantha moseleyi Andaman Sea [5] apud WoRMS Heterosaccus hians (Sacculina hians) (Kossmann, 1872) "Thalamita sp. aff. callianassae" "Java" [6] Heterosaccus indicus Boschma, 1957 "Portunus pelagicus" Mandapam Camp, Tamil Nadu [7] apud WoRMS Heterosaccus lunatus Phillips, 1978 "Charybdis callianassa" "Moreton Bay," Queensland [8] Heterosaccus multilacinensis Phillips, 1978 "Charybdis truncata" "East Moreton Bay," Queensland [8] Heterosaccus occidentalis (Drepanorchis occidentalis) (Boschma, 1928) "Mithrax forceps" (=Mithraculus forceps) "Deadman's Bay, west coast of Florida" [9] Heterosaccus papillosus (Drepanorchis papillosa) (Boschma, 1933) "Charybdis bimaculatus" (=Charybdis bimaculata) "vicinity of Marindugue Island, Philippine Islands" [3]: 234 Heterosaccus pellucidus Shiino, 1943 "Thalamita integra integra" Japan [10] apud WoRMS Heterosaccus ruginosus Boschma, 1931 "Thalamita prymna" "Singapore, coral reef" [11] Heterosaccus setoensis Shiino, 1943 "Thalamita wakensis" (= Thalamita seurati) Seto Inland Sea [10] apud WoRMS Heterosaccus sibogae Boschma, 1931 "Thalamita admeta" (= Thalamita admete) "Siau Reef", Indonesia [12] Heterosaccus tesselatus (Sacculina tessellata) (Boschma, 1925) "Mithrax ruber" (= Teleophrys ruber) "Caracas Bay, Curaçao" [13] ## References 1. Smith, Geoffrey (1906). ["Genus *Heterosaccus*"](https://www.biodiversitylibrary.org/page/37097701). *Rhizocephala*. Vol. 29. Fauna und Flora des Golfes von Neapal. R. Friedländer & Sohn. pp. 113–114. 1. Boschma, H. (1933). ["New species of Sacculinidae in the collection of the United States National Museum"](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=197346). *Tijdschrift der Nederlandsche Dierkundige Vereeniging*. **3**: 234–237. 1. Boschma, H. (1960). "A Rhizocephalan Parasite of the Crab *Charybdis hoplites* (Wood-Mason)". *Crustaceana*. **1** (1): 58–67. [doi:10.1163/156854060X00078](https://doi.org/10.1163/156854060X00078). [JSTOR 20140988](https://www.jstor.org/stable/20140988) 1. Boschma, H. (1962). "A rhizocephalan parasite of a spider crab from the Andaman Sea". *Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen*. **65**: 294–301. Series C. (*fide* [WoRMS](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=197000)). Not seen. 1. Kossmann, R. (1872). ["Beiträge zur Anatomie der schmarotzenden Rankenfüssler"](https://babel.hathitrust.org/cgi/pt?id=mdp.39015076673501;view=1up;seq=337). *Verhandlungen der physikalisch-medicinischen Gesellschaft in Würzburg*. **3**: 329–330. Neue Folge. * Reprinted: Kossmann, R. (1874). ["Beiträge zur Anatomie der schmarotzenden Rankenfüssler"](https://www.biodiversitylibrary.org/page/34290524). *Arbeiten aus dem Zoologisch-Zootomischen Institut in Würzburg*. **1**: 130–131. [Pl. 5](https://www.biodiversitylibrary.org/page/34290913), Fig. 17a, b; [Pl. 6](https://www.biodiversitylibrary.org/page/34290919), Fig. 2. 1. Boschma, H. (1957). "*Heterosaccus indicus*, sp. nov., a rhizocephalan parasite of the crab Portunus pelagicus (L.)". *The Annals of Biology*. **2**: 1–20. (*fide* [WoRMS](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=197107)). Not seen. 1. Philips, W. J. (1978). ["Some parasitic barnacles (Rhizocephala: Sacculinidae) from portunid crabs in Moreton Bay, Queensland"](https://www.biodiversitylibrary.org/page/54219649). *Memoirs of the Queensland Museum*. **18** (2): 255–263. 1. Boschma, H. (1928). ["Two common species of parasitic crustacea (Sacculinidae) of the West Indies"](https://www.biodiversitylibrary.org/page/15693633). *Proceedings of the United States National Museum*. **73** (2726): 4–6. [doi:10.5479/si.00963801.73-2726.1](https://doi.org/10.5479/si.00963801.73-2726.1) 1. Shiino, S. M. (1943). "Rhizocephala of Japan". *Journal of the Sigenkagaku Kenkyusyo*. **1**: 1–36. (*fide* [WoRMS](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=196969)). Not seen. 1. Boschma, H. (1931). ["Papers from Dr. Th. Mortensen's Pacific Expedition 1914–16. LV. Rhizocephala"](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=196950). *Videnskabelige Meddedelser fra den Dansk Naturhistoriske Forening i København*. **89**: 361–365. 1. Boschma, H. (1931). [*Die Rhizocephalen der Siboga Expedition. Supplement*](http://www.marinespecies.org/aphia.php?p=sourcedetails&id=196951). Vol. 31 bis. Siboga-Expeditie. Leiden: E. J. Brill. pp. 54–55. 1. Boschma, H. (1925). ["Rhizocephala of Curaçao"](https://repository.naturalis.nl/pub/504065). *Bijdragen tot de Dierkunde*. **24** (1): 12–13. [doi:10.1163/26660644-02401002](https://doi.org/10.1163/26660644-02401002). [ISSN 0067-8546](https://www.worldcat.org/issn/0067-8546) ## Further reading - Brockerhoff, Annette M.; McLay, Colin L.; Rybakov, Alexi V. (2010). "Occurrence of *Heterosaccus* (Cirripedia: Rhizocephala) in the New Zealand Crab *Metacarcinus novaezelandiae* (Decapoda: Cancridae) and Distribution of Other Rhizocephala in the South Pacific". *Journal of Crustacean Biology*. **30** (3): 377–383. [doi:10.1651/09-3270.1](https://doi.org/10.1651/09-3270.1) - Øksnebjerg, Bo (2000). "The Rhizocephala (Crustacea: Cirripedia) of the Mediterranean and Black Seas: Taxonomy, Biogeography, and Ecology". *Israel Journal of Zoology*. **46**: 72–74. [doi:10.1560/RCLC-NM2U-HV5L-6Q52](https://doi.org/10.1560/RCLC-NM2U-HV5L-6Q52) --- Adapted from the Wikipedia article [Heterosaccus](https://en.wikipedia.org/wiki/Heterosaccus) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Heterosaccus?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.