# Heiomasia

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Genus of fungi

Heiomasia Heiomasia seaveyorum Scientific classification Kingdom: Fungi Division: Ascomycota Class: Lecanoromycetes Order: Graphidales Family: Graphidaceae Genus: Heiomasia Nelsen, Lücking & Rivas Plata (2010) Type species Heiomasia sipmanii (Aptroot, Lücking & Rivas Plata) Nelsen, Lücking & Rivas Plata (2010) Species H. annamariae H. pallescens H. seaveyorum H. siamensis H. sipmanii

***Heiomasia*** is a [genus](/source/Genus) of [corticolous](/source/Corticolous_lichen) (bark-dwelling), [crustose lichens](/source/Crustose_lichen) in the family [Graphidaceae](/source/Graphidaceae). It has five species.

## Taxonomy

The genus was [circumscribed](/source/Circumscription_(taxonomy)) in 2010 by Matthew Nelsen, [Robert Lücking](/source/Robert_L%C3%BCcking) and Eimy Rivas Plata, with *[Heiomasia sipmanii](https://en.wikipedia.org/w/index.php?title=Heiomasia_sipmanii&action=edit&redlink=1)* assigned as the [type species](/source/Type_species).[1] This species, found in Thailand and the Philippines, was first described as *Herpothallon sipmanii*.[2] [Molecular phylogenetic](/source/Molecular_phylogenetic) analysis showed that it, along with another then-undescribed species, *[H. seaveyorum](https://en.wikipedia.org/w/index.php?title=Heiomasia_seaveyorum&action=edit&redlink=1)*, formed a [clade](/source/Clade) with a distinct lineage in the Graphidaceae, and so *Heiomasia* was created to contain them. The genus name honours [Harrie Sipman](/source/Harrie_Sipman), "recognizing his substantial contributions to tropical lichenology". The name is constructed from letters of his full name: **He**nricus **Jo**hannes **Ma**ria **Si**pman (with the "J" changed to "I").[1]

## Description

*Heiomasia* species have pale green to whitish [thalli](/source/Thallus) that are effuse (spread out) and byssoid (wispy, like cotton), and often have a white [prothallus](/source/Prothallus#In_lichens). A major characteristic of the genus is the presence of large sausage-shaped or disc-shaped [isidia](/source/Isidia)-like structures. [Lichen products](/source/Lichen_product) found in *Heiomasia* are [protocetraric acid](/source/Protocetraric_acid), [hypoprotocetraric acid](https://en.wikipedia.org/w/index.php?title=Hypoprotocetraric_acid&action=edit&redlink=1), and metabolically related satellite compounds.[1]

[Morphologically](/source/Morphology_(biology)), *Heiomasia* resembles species in *[Cryptothecia](/source/Cryptothecia)*, *[Herpothallon](/source/Herpothallon)*, and *[Dichosporidium](/source/Dichosporidium)*, but those genera are in the order [Arthoniales](/source/Arthoniales) and are not closely related.[1]

## Species

[Klaus Kalb](/source/Klaus_Kalb) published a [key](/source/Identification_key) to the five known *Heiomasia* species in 2020.[3]

- *[Heiomasia annamariae](https://en.wikipedia.org/w/index.php?title=Heiomasia_annamariae&action=edit&redlink=1)* Kalb (2020)[3]

- *[Heiomasia pallescens](https://en.wikipedia.org/w/index.php?title=Heiomasia_pallescens&action=edit&redlink=1)* Jagad.Ram (2013)[4] – [Andaman Islands](/source/Andaman_Islands)

- *[Heiomasia seaveyorum](https://en.wikipedia.org/w/index.php?title=Heiomasia_seaveyorum&action=edit&redlink=1)* Nelsen & Lücking (2010)[1] – [Florida](/source/Florida), USA

- *[Heiomasia siamensis](https://en.wikipedia.org/w/index.php?title=Heiomasia_siamensis&action=edit&redlink=1)* Kalb (2020)[3] – Malaysia

- *[Heiomasia sipmanii](https://en.wikipedia.org/w/index.php?title=Heiomasia_sipmanii&action=edit&redlink=1)* (Aptroot, Lücking & Rivas Plata) Nelsen, Lücking & Rivas Plata (2010)[1] – Philippines; Thailand

## References

1. ^ [***a***](#cite_ref-Nelsen_et_al._2010_1-0) [***b***](#cite_ref-Nelsen_et_al._2010_1-1) [***c***](#cite_ref-Nelsen_et_al._2010_1-2) [***d***](#cite_ref-Nelsen_et_al._2010_1-3) [***e***](#cite_ref-Nelsen_et_al._2010_1-4) [***f***](#cite_ref-Nelsen_et_al._2010_1-5) Nelsen, Matthew P.; Lücking, Robert; Plata, Eimy Rivas; Mbatchou, Joelle S. (2010). "*Heiomasia*, a new genus in the lichen-forming family Graphidaceae (Ascomycota: Lecanoromycetes: Ostropales) with disjunct distribution in Southeastern North America and Southeast Asia". *The Bryologist*. **113** (4): 742–751. [doi](/source/Doi_(identifier)):[10.1639/0007-2745-113.4.742](https://doi.org/10.1639%2F0007-2745-113.4.742). [S2CID](/source/S2CID_(identifier)) [84013904](https://api.semanticscholar.org/CorpusID:84013904).

1. **[^](#cite_ref-Aptroot_et_al._2009_2-0)** Aptroot, A.; Thor, G.; Lücking, R.; Elix, J.A.; Chaves, J.L. (2009). "The lichen genus *Herpothallon* reinstated". *Bibliotheca Lichenologica*. **99**: 19–67.

1. ^ [***a***](#cite_ref-Kalb_2020_3-0) [***b***](#cite_ref-Kalb_2020_3-1) [***c***](#cite_ref-Kalb_2020_3-2) Kalb, K. (2020). ["New or otherwise interesting lichens. VII, including a world key to the lichen genus *Heiomasia*"](http://fschumm.de/Archive/Vol15_Kalb_New_lichens_VII.pdf) (PDF). *Archive for Lichenology*. **15**: 1–18.

1. **[^](#cite_ref-Jagadeesh_Ram_2014_4-0)** Jagadeesh Ram, T.A.M. (2014). "The genus *Herpothallon* (Arthoniaceae) in the Andaman Islands, India". *The Lichenologist*. **46** (1): 39–49. [doi](/source/Doi_(identifier)):[10.1017/s0024282913000571](https://doi.org/10.1017%2Fs0024282913000571). [S2CID](/source/S2CID_(identifier)) [232393678](https://api.semanticscholar.org/CorpusID:232393678).

Taxon identifiers Heiomasia Wikidata: Q21231667 Wikispecies: Heiomasia CoL: 62THM EoL: 21305381 GBIF: 8310991 iNaturalist: 474424 IndexFungorum: 516050 IRMNG: 1465898 MycoBank: 516050 NCBI: 941792 Open Tree of Life: 952201 SpeciesFungorum: 516050

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