# Handkea

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{{Short description|Genus of fungi}}
{{Automatic_taxobox
| image = Hand.utri.jpg
| image_caption = ''Handkea utriformis''
| taxon = Handkea
| authority = Kreisel
| type_species = ''[Handkea utriformis](/source/Handkea_utriformis)''
| type_species_authority = (Bull.) Kreisel
}}

'''''Handkea''''' is a genus of [puffball](/source/puffball) mushrooms in the family [Agaricaceae](/source/Agaricaceae). In 1989, German mycologist Hanns Kreisel described the genus ''Handkea'' to include species of ''[Calvatia](/source/Calvatia)'' with distinct microscopic features, including a unique type of [capillitium](/source/capillitium) (coarse thick-walled [hyphae](/source/hyphae) in the [gleba](/source/gleba)), with curvy slits instead of the usual pores.<ref>{{cite journal|last=Kreisel H.|year=1989|title=Studies in the ''Calvatia'' complex (basidiomycetes)|journal=Nova Hedwigia|volume=48|issue=3–4|pages=281–96}}</ref> Although accepted by some authors,<ref name="isbn0-947643-81-8">{{cite book|vauthors=Laessøe T, Pegler DN, Spooner B |title=British puffballs, earthstars and stinkhorns: an account of the British gasteroid fungi|publisher=Royal Botanic Gardens|location=Kew|year=1995|isbn=0-947643-81-8}}</ref> the genus concept has been rejected by others.<ref name=Demoulin1990>{{cite journal | last1 = Demoulin | first1 = V | last2 = Lange | first2 = M | year = 1990 | title = ''Calvatia turneri'' (Ellis et Everh.) Demoulin et M. Lange, comb. nov., the correct name for ''C. tatrensis'' Hollós | journal = Mycotaxon | volume = 38 | pages = 221–26 }}</ref><ref>Demoulin V. (1971). "Le genre ''Lycoperdon'' en Europe et en Amérique du Nord. Etude taxonomique et phytogéographique". PhD. thesis, Université de Liège.</ref>

The genus name of ''Handkea'' is in honour of Horst-Herbert Handke (1913-2005), who was a German botanist and Professor at the [University of Halle](/source/University_of_Halle).<ref>{{cite book | last=Burkhardt | first=Lotte | title=Eine Enzyklopädie zu eponymischen Pflanzennamen |trans-title=Encyclopedia of eponymic plant names | publisher=Botanic Garden and Botanical Museum, Freie Universität Berlin | year=2022 | isbn=978-3-946292-41-8 | url=https://doi.org/10.3372/epolist2022|format=pdf |language=German |location=Berlin | doi=10.3372/epolist2022 |access-date=January 27, 2022}}</ref>

[Phylogenetic](/source/Phylogenetic) analyses published in 2008 shows that ''Handkea'' may be grouped in a [clade](/source/clade) along with species from several other genera, including ''[Lycoperdon](/source/Lycoperdon)'', ''[Vascellum](/source/Vascellum)'', ''[Morganella](/source/Morganella_(fungus))'', ''[Bovistella](/source/Bovistella)'', and ''[Calvatia](/source/Calvatia)''.<ref name="pmid18207380">{{cite journal|vauthors=Larsson E, Jeppson M |title=Phylogenetic relationships among species and genera of Lycoperdaceae based on ITS and LSU sequence data from north European taxa|journal=Mycological Research|volume=112|issue=Pt 1|pages=4–22|year=2008|pmid=18207380|doi=10.1016/j.mycres.2007.10.018}}</ref>

==Species==
{{Columns-list|colwidth=30em|
*''[H. canadensis](/source/Handkea_canadensis)''
*''[H. capensis](/source/Handkea_capensis)''
*''[H. fumosa](/source/Handkea_fumosa)''
*''[H. hesperia](/source/Handkea_hesperia)''
*''[H. lloydii](/source/Handkea_lloydii)''
*''[H. lycoperdoides](/source/Handkea_lycoperdoides)''
*''[H. subcretacea](/source/Handkea_subcretacea)''
*''[H. wandae](/source/Handkea_wandae)''
}}

==References==
{{Reflist}}

==External links==
*{{cite web |title=''Handkea'' Kreisel |work=Atlas of Living Australia |url=https://bie.ala.org.au/species/urn:lsid:indexfungorum.org:names:25306}}

{{Taxonbar |from=Q5647449}}

Category:Lycoperdaceae
Category:Agaricales genera
Category:Puffballs

{{Agaricales-stub}}

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Adapted from the Wikipedia article [Handkea](https://en.wikipedia.org/wiki/Handkea) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Handkea?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.