# Haemulinae

> Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Haemulinae
> Markdown URL: https://mediated.wiki/source/Haemulinae.md
> Source: https://en.wikipedia.org/wiki/Haemulinae
> Source revision: 1311504502
> License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/)

Subfamily of fishes

Haemulinae Temporal range: Middle Eocene–present PreꞒ Ꞓ O S D C P T J K Pg N Xenocys jessiae, Coiba National Park, Panama Scientific classification Kingdom: Animalia Phylum: Chordata Class: Actinopterygii Order: Acanthuriformes Family: Haemulidae Subfamily: Haemulinae Gill, 1885[1] Genera See text

**Haemulinae** is a subfamily of the [Haemulidae](/source/Haemulidae) and consists of the genera of that family which are regarded as being of New World origin, although they are now widespread.[2] The subfamily is distinguished from the [Plectorhynchinae](/source/Plectorhynchinae) by having a short [dorsal fin](/source/Dorsal_fin) which contains 13-16 soft rays, as opposed to the long dorsal fin with 17-26 soft rays of the subfamily [Plectorhynchinae](/source/Plectorhynchinae).[3]

## Genera

The following genera are included in the Haemulinae by *[Eschmeyer's Catalog of Fishes](/source/Eschmeyer's_Catalog_of_Fishes)*:[4]

- *[Anisotremus](/source/Anisotremus)* Gill 1861

- *[Boridia](/source/Boridia)* [Cuvier](/source/Georges_Cuvier), 1830

- *[Brachydeuterus](/source/Brachydeuterus)* Gill, 1862

- *[Brachygenys](/source/Brachygenys)* [Poey](/source/Felipe_Poey), 1868

- *[Conodon](/source/Conodon)* Cuvier, 1830

- *[Emmelichthyops](/source/Emmelichthyops)* [Schultz](/source/Leonard_Peter_Schultz), 1945

- *[Haemulon](/source/Haemulon)* Cuvier, 1829

- *[Haemulopsis](/source/Haemulopsis)* [Steindachner](/source/Franz_Steindachner), 1869

- *[Isacia](/source/Isacia)* [Jordan](/source/David_Starr_Jordan) & [Fesler](https://en.wikipedia.org/w/index.php?title=Bert_Fesler&action=edit&redlink=1), 1893

- *[Microlepidotus](/source/Microlepidotus)* Gill, 1862

- *[Orthopristis](/source/Orthopristis)* [Girard](/source/Charles_Fr%C3%A9d%C3%A9ric_Girard), 1858

- *[Parakuhlia](/source/Parakuhlia)* [Pellegrin](/source/Jacques_Pellegrin), 1913

- *[Paranisotremus](/source/Paranisotremus)* [Tavera](https://en.wikipedia.org/w/index.php?title=Jose_Tavera&action=edit&redlink=1), [Acero](https://en.wikipedia.org/w/index.php?title=Arturo_Acero_P.&action=edit&redlink=1) & [Wainwright](https://en.wikipedia.org/w/index.php?title=Peter_C._Wainwright&action=edit&redlink=1), 2018

- *[Pomadasys](/source/Pomadasys)* [Lacépède](/source/Bernard_Germain_de_Lac%C3%A9p%C3%A8de), 1802

- *[Rhencus](/source/Rhencus)* Jordan & [Evermann](/source/Barton_Warren_Evermann), 1896

- *[Rhonciscus](/source/Rhonciscus)* Jordan & Evermann, 1896

- *[Xenichthys](/source/Xenichthys)* Gill, 1862

The genus *[Brachygenys](/source/Brachygenys)* is recognised by some authorities as *Haemulon* was determined to be [paraphyletic](/source/Paraphyletic) in molecular studies which showed *[Haemulon chrysargyreum](/source/Haemulon_chrysargyreum)* clustered with *[Xenistius californianus](https://en.wikipedia.org/w/index.php?title=Xenistius_californianus&action=edit&redlink=1)*. The genus also includes the other species in the former genera *[Xenistius](/source/Xenistius)* and *[Xenocys](/source/Xenocys)*.[2] Similarly, these studies also resolved *Pomadasys* as paraphyletic and the genera *[Rhencus](/source/Rhencus)* and *[Rhonciscus](/source/Rhonciscus)* were revived to solve this paraphyly, with the eastern Pacific species *P. macracanthus* and *P. panamensis* being placed in *Rhencus* while *Rhonciscus* contains the eastern Pacific species *P. bayanus* and the western Atlantic species *P. crocro*.[5] These changes are recognised by *[Catalog of Fishes](/source/Catalog_of_Fishes)*[4] but not yet by [Fishbase](/source/Fishbase).[6] In addition, according to these studies, *[Anisotremus moricandi](/source/Anisotremus_moricandi)* makes *[Anisotremus](/source/Anisotremus)* paraphyletic if included and has been placed in the [monotypic](/source/Monotypic) genus *[Paranisotremus](/source/Paranisotremus)*.[7]

Other authorities place the genus *[Genyatremus](/source/Genyatremus)* within the Haemulinae,[5] although both Fishbase and *Catalog of Fishes* put this genus in the other Haemulid subfamily Plectorhinchinae.[6][4]

*[Pomadasys](/source/Pomadasys)* fossils are known from the [Middle Eocene](/source/Middle_Eocene) of Egypt.[8]

## References

1. **[^](#cite_ref-VDLEF_1-0)** Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014). ["Family-group names of Recent fishes"](https://biotaxa.org/Zootaxa/article/view/zootaxa.3882.1.1/10480). *Zootaxa*. **3882** (2): 001–230.

1. ^ [***a***](#cite_ref-BMC_2-0) [***b***](#cite_ref-BMC_2-1) José Julián Tavera; P. Arturo Acero; Eduardo F Balart; Giacomo Bernardi (2012). ["Molecular phylogeny of grunts (Teleostei, Haemulidae), with an emphasis on the ecology, evolution, and speciation history of New World species"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3472276). *BMC Evolutionary Biology*. **12** (57): 57. [doi](/source/Doi_(identifier)):[10.1186/1471-2148-12-57](https://doi.org/10.1186%2F1471-2148-12-57). [PMC](/source/PMC_(identifier)) [3472276](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3472276). [PMID](/source/PMID_(identifier)) [22537107](https://pubmed.ncbi.nlm.nih.gov/22537107).

1. **[^](#cite_ref-Nelson5_3-0)** J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). [*Fishes of the World*](https://web.archive.org/web/20190408194051/https://sites.google.com/site/fotw5th/) (5th ed.). Wiley. p. 457. [ISBN](/source/ISBN_(identifier)) [978-1-118-34233-6](https://en.wikipedia.org/wiki/Special:BookSources/978-1-118-34233-6). Archived from [the original](https://sites.google.com/site/fotw5th/) on 2019-04-08. Retrieved 2021-03-19.

1. ^ [***a***](#cite_ref-CofF_4-0) [***b***](#cite_ref-CofF_4-1) [***c***](#cite_ref-CofF_4-2) Fricke, Ron; [Eschmeyer, William N.](/source/William_N._Eschmeyer) & van der Laan, Richard (eds.). ["Genera in the family Haemulinae"](http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatget.asp?tbl=genus&family=Haemulinae). *[Catalog of Fishes](/source/Catalog_of_Fishes)*. [California Academy of Sciences](/source/California_Academy_of_Sciences). Retrieved 9 September 2025.

1. ^ [***a***](#cite_ref-Parenti_5-0) [***b***](#cite_ref-Parenti_5-1) Paolo Parenti (2019). ["An annotated checklist of the fishes of the family Haemulidae (Teleostei: Perciformes)"](https://www.researchgate.net/publication/332223321). *Iranian Journal of Ichthyology*. **6** (3): 150–196.

1. ^ [***a***](#cite_ref-Fishbase_6-0) [***b***](#cite_ref-Fishbase_6-1) [Froese, Rainer](/source/Rainer_Froese); [Pauly, Daniel](/source/Daniel_Pauly) (eds.). ["Family Haemulidae"](https://www.fishbase.org/summary/FamilySummary.php?family=Haemulidae). *[FishBase](/source/FishBase)*. December 2021 version.

1. **[^](#cite_ref-Tavera2018_7-0)** Jose Tavera; Arturo Acero P.; Peter C. Wainwright (2018). ["Multilocus phylogeny, divergence times, and a major role for the benthic-to- pelagic axis in the diversification of grunts (Haemulidae)"](https://doi.org/10.1016%2Fj.ympev.2017.12.032). *Molecular Phylogenetics and Evolution*. **121**: 212–223. [doi](/source/Doi_(identifier)):[10.1016/j.ympev.2017.12.032](https://doi.org/10.1016%2Fj.ympev.2017.12.032). [PMID](/source/PMID_(identifier)) [29307507](https://pubmed.ncbi.nlm.nih.gov/29307507).

1. **[^](#cite_ref-8)** White, Errol Ivor (1936-07-01). ["V.—On certain Eocene percoid fishes"](https://doi.org/10.1080/00222933608655173). *Annals and Magazine of Natural History*. **18** (103): 43–54. [doi](/source/Doi_(identifier)):[10.1080/00222933608655173](https://doi.org/10.1080%2F00222933608655173). [ISSN](/source/ISSN_(identifier)) [0374-5481](https://search.worldcat.org/issn/0374-5481).

Taxon identifiers Haemulinae Wikidata: Q1550855 Open Tree of Life: 99942 WoRMS: 714648

---
Adapted from the Wikipedia article [Haemulinae](https://en.wikipedia.org/wiki/Haemulinae) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Haemulinae?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.