# Gyalectidium

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Genus of lichen-forming fungi

Gyalectidium Gyalectidium puntilloi Scientific classification Kingdom: Fungi Division: Ascomycota Class: Lecanoromycetes Order: Graphidales Family: Gomphillaceae Genus: Gyalectidium Müll.Arg. (1881) Type species Gyalectidium filicinum Müll.Arg. (1881) Species See text Synonyms[1] List Gonothecis Clem. (1909) Gonothecium (Vain.) Clem. & Shear (1931) Lecidea sect. Gonothecium Vain. (1890) Lopadiopsidomyces Cif. & Tomas. (1953) Lopadiopsis Vain. (1896) Tauromyces Cavalc. & A.A.Silva (1972)

***Gyalectidium*** is a [genus](/source/Genus) of [lichen](/source/Lichen)-forming [fungi](/source/Fungi) in the family [Gomphillaceae](/source/Gomphillaceae). Established in 1881 by Swiss lichenologist [Johannes Müller Argoviensis](/source/Johannes_M%C3%BCller_Argoviensis), the genus comprises about 50 species of predominantly [leaf-dwelling lichens](/source/Foliicolous_lichen) that form small, pale greenish to whitish-grey patches typically only a few millimetres across. Members are characterised by specialised [asexual](/source/Asexual_reproduction) reproductive structures called [hyphophores](/source/Glossary_of_lichen_terms#hyphophores), which are small scale-like outgrowths that produce [propagules](/source/Propagule) containing both fungal and algal partners, and by [fruiting bodies](/source/Fruiting_bodies) that contain single, large, multi-chambered [ascospores](/source/Ascospore). The genus has a mainly [pantropical](/source/Pantropical) to [subtropical](/source/Subtropical) distribution with highest diversity in the [Neotropics](/source/Neotropics), occurring primarily on living leaves in humid tropical forests but also documented from [temperate](/source/Temperate) regions including Europe and parts of Australasia.

## Taxonomy

The genus was [circumscribed](/source/Circumscription_(taxonomy)) by the Swiss lichenologist [Johannes Müller Argoviensis](/source/Johannes_M%C3%BCller_Argoviensis) in 1881. In the protologue, he characterised *Gyalectidium* as a [crustose lichen](/source/Crustose_lichen) with globose, green [gonidia](/source/Glossary_of_lichen_terms#gonidia) (green algae), [lecanorine](/source/Glossary_of_lichen_terms#lecanorine) apothecia with a simple margin that is thalline externally, and [paraphyses](/source/Paraphyses) that are interconnected in a lattice. The spores were described as [hyaline](/source/Hyaline) and "parenchymatous" (divided into many chambers). He distinguished the genus from *[Gyalecta](/source/Gyalecta)* by the structure of the paraphyses, and remarked that the species then known were [foliicolous](/source/Foliicolous_lichen) and had one-spored [asci](/source/Ascus). He included 3 species: *G. xantholeucum*, *G. dispersum*, and *G. filicinum*;[2] the last of these is now the [type species](/source/Type_species) of the genus.[3]

In their 2001 monograph, Ferraro and colleagues proposed an infrageneric classification that organised the species treated there into [sections](/source/Section_(botany)) and [series](/source/Series_(botany)), largely reflecting thallus texture (especially the distribution of calcium oxalate crystals) and the form and placement of the hyphophores. They recognised three main groupings as sections, each with a single series: sect. *Placolectidium* (ser. *Caucasicae*), comprising species with [placoid](/source/Glossary_of_lichen_terms#placoid) to [bullate](/source/Glossary_of_lichen_terms#bullate) thalli that are evenly encrusted with crystals and often form their hyphophores at the thallus margin; sect. *Areolectidium* (ser. *Areolatae*), including species with smoother thalli lacking crystals or with a distinctly areolate pattern, typically with well-developed hyphophore scales; and the autonymous sect. *Gyalectidium* (ser. *Gyalectidium*), centred on the *G. filicinum* group, characterised by a finely [verrucose](/source/Glossary_of_lichen_terms#verrucose) thallus in which crystals are aggregated into small wart-like patches and by a wide range of hyphophore forms. Two species that appeared isolated in their analysis were placed in [monotypic](/source/Monotypic) sections and series: sect. *Setolectidium* (ser. *Microcarpae*) for *G. microcarpum*, which has abundant sterile setae and lacks hyphophores, and sect. *Goniolectidium* (ser. *Yahriae*) for *G. yahriae*, in which the hyphophores are modified into multi-part structures with a goniocystangium-like appearance.[4]

## Description

*Gyalectidium* is a genus of mostly leaf-dwelling ([foliicolous](/source/Foliicolous_lichen)) [crustose lichens](/source/Crustose_lichen) that form small, pale greenish to whitish-grey patches, often only a few millimetres across (rarely approaching about 10 mm). The [thallus](/source/Thallus) is always [corticate](/source/Glossary_of_lichen_terms#corticate) (i.e., it has a surface layer of fungal tissue), and it commonly carries colourless [calcium oxalate](/source/Calcium_oxalate) crystals that give the surface a finely warty, coarsely [verrucose](/source/Glossary_of_lichen_terms#verrucose), or sometimes distinctly [areolate](/source/Glossary_of_lichen_terms#areolate) look; in a few species the thallus is smoother because crystals are sparse or absent. The [photosynthetic](/source/Photosynthesis) partner is a [green alga](/source/Green_alga) of the genus *[Trebouxia](/source/Trebouxia)*.[4]

The genus is recognised by its specialised asexual structures called [hyphophores](/source/Glossary_of_lichen_terms#hyphophores): small, scale-like outgrowths that may stand upright, slant, or lie close to the thallus surface, and in some species are much reduced or missing. At the base (or centre) of each hyphophore a mass of short, branched hyphae (the diahyphal or "conidial" mass) develops; it is typically intermixed with algal cells and often breaks into smaller rounded [propagules](/source/Propagule) that can disperse both partners together. Sexual [fruiting bodies](/source/Fruiting_bodies) ([apothecia](/source/Apothecia)) are immersed in the thallus and have a [thalline margin](/source/Glossary_of_lichen_terms#thalline_margin). Internally they have a colourless, non-[amyloid](/source/Amyloid_(mycology)) hymenium with richly branched and interconnected [paraphyses](/source/Paraphyses), and [asci](/source/Ascus) that usually contain a single, colourless [ascospore](/source/Ascospore). The ascospore is [muriform](/source/Glossary_of_lichen_terms#muriform) (divided by both transverse and longitudinal [septa](/source/Septum) into many chambers). [Pycnidia](/source/Pycnidia) are uncommon in the genus.[4]

## Habitat and distribution

Most *Gyalectidium* species are best known as [foliicolous lichens](/source/Foliicolous_lichen), forming tiny crusts on living leaves in humid climates. In the 2001 world monograph, the genus was reported most often from [tropical](/source/Tropical) forests, where at least some species can occur across a wide range of light conditions—from shaded [understory](/source/Understory) leaves to more exposed leaves in the [canopy](/source/Canopy_(botany)). Records discussed there also suggest that some *Gyalectidium* lichens can [colonise](/source/Colonisation_(biology)) very young leaves early in succession (behaving as [pioneer](/source/Pioneer_species) colonists), yet remain present on older leaves later in the life of the leaf. Several species may be found together on the same leaf, which implies broadly similar [microhabitat](/source/Microhabitat) requirements within a site.[4]

As treated in that monograph, *Gyalectidium* had a mainly [pantropical](/source/Pantropical) to [subtropical](/source/Subtropical) distribution overall, with the highest recorded diversity in the [Neotropics](/source/Neotropics) (especially areas that had been intensively collected), and additional species documented from tropical Africa, Europe (including the [Caucasus](/source/Caucasus) and [Macaronesia](/source/Macaronesia)), and [Australasia](/source/Australasia). The authors stressed that broad patterns were still hard to generalise, because many species were known from few collections and some regions (especially tropical Asia) appeared undercollected at the time. Even within better-sampled areas, the monograph suggests some ecological separation: certain species were reported more often from open or canopy situations, while others were more frequently detected in secondary vegetation. Although the genus was treated as overwhelmingly leaf-dwelling, the monograph also recorded a small number of exceptions, including a species occurring on twigs/decorticated wood, and an occasional streamside-rock record.[4]

## Species

*Gyalectidium minus*

As of February 2026[\[update\]](https://en.wikipedia.org/w/index.php?title=Gyalectidium&action=edit), [Species Fungorum](/source/Species_Fungorum) (in the [Catalogue of Life](/source/Catalogue_of_Life)) accept 46 species of *Gyalectidium*.[5]

- *[Gyalectidium appendiculatum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_appendiculatum&action=edit&redlink=1)* Lücking, Lendemer & E.A.Tripp (2008)[6]

- *[Gyalectidium areolatum](/source/Gyalectidium_areolatum)* L.I.Ferraro & Lücking (2001)[4]

- *[Gyalectidium atrosquamulatum](/source/Gyalectidium_atrosquamulatum)* Lücking & Kalb (2001)[4]

- *[Gyalectidium aurelii](https://en.wikipedia.org/w/index.php?title=Gyalectidium_aurelii&action=edit&redlink=1)* L.I.Ferraro & Lücking (2003)[7]

- *[Gyalectidium australe](/source/Gyalectidium_australe)* Lücking (2001)[4]

- *[Gyalectidium barbatum](/source/Gyalectidium_barbatum)* Herrera-Camp. & Lücking (2002)[8] – Mexico

- *[Gyalectidium catenulatum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_catenulatum&action=edit&redlink=1)* (Cavalc. & A.A.Silva) L.I.Ferraro, Lücking & Sérus. (2000)

- *[Gyalectidium caucasicum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_caucasicum&action=edit&redlink=1)* (Elenkin & Woron.) Vězda (1983)

- *[Gyalectidium chilense](https://en.wikipedia.org/w/index.php?title=Gyalectidium_chilense&action=edit&redlink=1)* M.Cáceres & Lücking (2003)[9]

- *[Gyalectidium ciliatum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_ciliatum&action=edit&redlink=1)* Lücking, G.Thor & Tat. Matsumoto (2000)[10]

- *[Gyalectidium cinereodiscus](https://en.wikipedia.org/w/index.php?title=Gyalectidium_cinereodiscus&action=edit&redlink=1)* Herrera-Camp. & Lücking (2004)[11]

- *[Gyalectidium conchiferum](/source/Gyalectidium_conchiferum)* Lücking & V.Wirth (2001)[4]

- *[Gyalectidium denticulatum](/source/Gyalectidium_denticulatum)* Lücking (2001)[4]

- *[Gyalectidium fantasticum](/source/Gyalectidium_fantasticum)* L.I.Ferraro & Lücking (2001)[4]

- *[Gyalectidium filicinum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_filicinum&action=edit&redlink=1)* Müll.Arg. (1881)

- *[Gyalectidium flabellatum](/source/Gyalectidium_flabellatum)* Sérus. (2001)[4]

- *[Gyalectidium floridense](/source/Gyalectidium_floridense)* Safranek & Lücking (2005)[12] – [Florida](/source/Florida), USA

- *[Gyalectidium fuscum](/source/Gyalectidium_fuscum)* Lücking & Sérus. (2001)[4]

- *[Gyalectidium gahavisukanam](https://en.wikipedia.org/w/index.php?title=Gyalectidium_gahavisukanam&action=edit&redlink=1)* Sérus. (2001)[4]

- *[Gyalectidium imperfectum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_imperfectum&action=edit&redlink=1)* Vězda (1994)[13]

- *[Gyalectidium kenyanum](/source/Gyalectidium_kenyanum)* Lücking & Kalb (2001)[4]

- *[Gyalectidium laciniatum](/source/Gyalectidium_laciniatum)* Lücking (2001)[4]

- *[Gyalectidium macaronesicum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_macaronesicum&action=edit&redlink=1)* Sérus. (2000)[14]

- *[Gyalectidium maracae](https://en.wikipedia.org/w/index.php?title=Gyalectidium_maracae&action=edit&redlink=1)* Lücking (2001)[4]

- *[Gyalectidium membranaceum](/source/Gyalectidium_membranaceum)* Sérus. & Lücking (2001)[4]

- *[Gyalectidium microcarpum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_microcarpum&action=edit&redlink=1)* (Vězda) Lücking, Sérus. & Vězda (2001)[4]

- *[Gyalectidium minus](/source/Gyalectidium_minus)* Sérus. (2001)[4]

- *[Gyalectidium nashii](https://en.wikipedia.org/w/index.php?title=Gyalectidium_nashii&action=edit&redlink=1)* Herrera-Camp. & Lücking (2003)[15]

- *[Gyalectidium novoguineense](/source/Gyalectidium_novoguineense)* Sérus. (2001)[4]

- *[Gyalectidium pallidum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_pallidum&action=edit&redlink=1)* Herrera-Camp. & Lücking (2002)[8]

- *[Gyalectidium paolae](https://en.wikipedia.org/w/index.php?title=Gyalectidium_paolae&action=edit&redlink=1)* Herrera-Camp. & Lücking (2003)[15]

- *[Gyalectidium plicatum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_plicatum&action=edit&redlink=1)* L.I.Ferraro & Lücking (2003)[9]

- *[Gyalectidium puntilloi](/source/Gyalectidium_puntilloi)* Sérus. (2001)[4]

- *[Gyalectidium radiatum](https://en.wikipedia.org/w/index.php?title=Gyalectidium_radiatum&action=edit&redlink=1)* Lücking, G.Thor & Tat.Matsumoto (2000)[10]

- *[Gyalectidium rosae-emiliae](https://en.wikipedia.org/w/index.php?title=Gyalectidium_rosae-emiliae&action=edit&redlink=1)* Herrera-Camp. & Lücking (2003)[15]

- *[Gyalectidium sanmartinense](https://en.wikipedia.org/w/index.php?title=Gyalectidium_sanmartinense&action=edit&redlink=1)* Herrera-Camp. & Lücking (2004)[11]

- *[Gyalectidium shimanense](https://en.wikipedia.org/w/index.php?title=Gyalectidium_shimanense&action=edit&redlink=1)* Y.Suto (2018)[16] – Japan

- *[Gyalectidium tuckerae](https://en.wikipedia.org/w/index.php?title=Gyalectidium_tuckerae&action=edit&redlink=1)* Lücking & Lendemer (2007)[17] – [Louisiana](/source/Louisiana), USA

- *[Gyalectidium ulloae](https://en.wikipedia.org/w/index.php?title=Gyalectidium_ulloae&action=edit&redlink=1)* Herrera-Camp. & Lücking (2002)[8] – Mexico

- *[Gyalectidium verruculosum](/source/Gyalectidium_verruculosum)* Sérus. (2001)[4]

- *[Gyalectidium viride](https://en.wikipedia.org/w/index.php?title=Gyalectidium_viride&action=edit&redlink=1)* Lücking, W.R.Buck & Rivas Plata (2007)[17] – [Alabama](/source/Alabama), USA

- *[Gyalectidium yahriae](/source/Gyalectidium_yahriae)* W.R.Buck & Sérus. (2000)[18] – Florida, USA

## References

1. **[^](#cite_ref-Species_Fungorum_synonymy_1-0)** ["Synonymy: *Gyalectidium* Müll. Arg., Flora, Regensburg 64(7): 100 (1881)"](https://www.speciesfungorum.org/Names/SynSpecies.asp?RecordID=2158). [Species Fungorum](/source/Species_Fungorum). Retrieved 5 March 2022.

1. **[^](#cite_ref-Müller_1881_2-0)** Müller, J. (1881). ["Lichenologische Beiträge XII"](https://www.biodiversitylibrary.org/page/66158). *Flora (Regensburg)* (in Latin). **64** (7): 100–112.

1. **[^](#cite_ref-IF:_Gyalectidium_3-0)** ["Record Details: *Gyalectidium* Müll. Arg., Flora, Regensburg 64(7): 100 (1881)"](https://www.indexfungorum.org/Names/NamesRecord.asp?RecordID=2158). [Index Fungorum](/source/Index_Fungorum). Retrieved 1 February 2026.

1. ^ [***a***](#cite_ref-Ferraro_et_al._2001_4-0) [***b***](#cite_ref-Ferraro_et_al._2001_4-1) [***c***](#cite_ref-Ferraro_et_al._2001_4-2) [***d***](#cite_ref-Ferraro_et_al._2001_4-3) [***e***](#cite_ref-Ferraro_et_al._2001_4-4) [***f***](#cite_ref-Ferraro_et_al._2001_4-5) [***g***](#cite_ref-Ferraro_et_al._2001_4-6) [***h***](#cite_ref-Ferraro_et_al._2001_4-7) [***i***](#cite_ref-Ferraro_et_al._2001_4-8) [***j***](#cite_ref-Ferraro_et_al._2001_4-9) [***k***](#cite_ref-Ferraro_et_al._2001_4-10) [***l***](#cite_ref-Ferraro_et_al._2001_4-11) [***m***](#cite_ref-Ferraro_et_al._2001_4-12) [***n***](#cite_ref-Ferraro_et_al._2001_4-13) [***o***](#cite_ref-Ferraro_et_al._2001_4-14) [***p***](#cite_ref-Ferraro_et_al._2001_4-15) [***q***](#cite_ref-Ferraro_et_al._2001_4-16) [***r***](#cite_ref-Ferraro_et_al._2001_4-17) [***s***](#cite_ref-Ferraro_et_al._2001_4-18) [***t***](#cite_ref-Ferraro_et_al._2001_4-19) [***u***](#cite_ref-Ferraro_et_al._2001_4-20) [***v***](#cite_ref-Ferraro_et_al._2001_4-21) [***w***](#cite_ref-Ferraro_et_al._2001_4-22) Ferraro, L.I.; Lücking, R.; Sérusiaux, E. (2001). ["A world monograph of the lichen genus *Gyalectidium* (Gomphillaceae)"](https://doi.org/10.1111%2Fj.1095-8339.2001.tb01126.x). *Botanical Journal of the Linnean Society*. **137** (3): 311–345. [doi](/source/Doi_(identifier)):[10.1111/j.1095-8339.2001.tb01126.x](https://doi.org/10.1111%2Fj.1095-8339.2001.tb01126.x). [hdl](/source/Hdl_(identifier)):[11336/31010](https://hdl.handle.net/11336%2F31010).

1. **[^](#cite_ref-CoL_4RP4_5-0)** ["*Gyalectidium* Müll.Arg"](https://www.catalogueoflife.org/data/taxon/4RP4). *[Catalogue of Life](/source/Catalogue_of_Life)*. [Species 2000](/source/Species_2000): Leiden, the Netherlands. Retrieved 1 February 2026.

1. **[^](#cite_ref-Lendemer_&_Tripp_2008_6-0)** Lendemer, James C.; Tripp, Erin A. (2008). "Contributions to the lichen flora of North Carolina: a preliminary checklist of the lichens of Gorges State Park". *The Bryologist*. **111** (1): 57–67. [doi](/source/Doi_(identifier)):[10.1639/0007-2745(2008)111\[57:CTTLFO\]2.0.CO;2](https://doi.org/10.1639%2F0007-2745%282008%29111%5B57%3ACTTLFO%5D2.0.CO%3B2).

1. **[^](#cite_ref-Ferraro_&_Lücking_2003_7-0)** Ferraro, L.I.; Lücking, R. (2003). "*Gyalectidium aurellii* (Ostropales: Gomphillaceae), a new foliicolous lichen from the state of Mato Grosso, Brazil". *Acta Botanica Brasilica*. **17** (4): 619–622.

1. ^ [***a***](#cite_ref-Herrera-Campos_&_Lücking_2002_8-0) [***b***](#cite_ref-Herrera-Campos_&_Lücking_2002_8-1) [***c***](#cite_ref-Herrera-Campos_&_Lücking_2002_8-2) Herrera-Campos, M.A.; Lücking, R. (2002). "The foliicolous lichen flora of Mexico. I. New species from Los Tuxtlas Tropical Biology Station, Veracruz". *The Lichenologist*. **34** (3): 211–222. [doi](/source/Doi_(identifier)):[10.1006/lich.2002.0397](https://doi.org/10.1006%2Flich.2002.0397).

1. ^ [***a***](#cite_ref-Lücking_et_al._2003_9-0) [***b***](#cite_ref-Lücking_et_al._2003_9-1) Lücking, R.; Wirth, V.; Ferraro, L.I.; Cáceres, M.E.S. (2003). "Foliicolous lichens from Valdivian temperate rainforest of Chile and Argentina: evidence of an austral element, with the description of seven new taxa". *Global Ecology and Biogeography*. **12**: 21–36.

1. ^ [***a***](#cite_ref-Thor_et_al._2000_10-0) [***b***](#cite_ref-Thor_et_al._2000_10-1) Thor, G.; Lücking, R.; Matsumoto, T. (2000). "The foliicolous lichen flora of Japan". *Symbolae Botanicae Upsalienses*. **32** (3): 1–72.

1. ^ [***a***](#cite_ref-Herrera-Campos_et_al._2004_11-0) [***b***](#cite_ref-Herrera-Campos_et_al._2004_11-1) Herrera-Campos, M.A.; Colín, P.M.; Peña, A.B.; Lücking, R. (2004). "The foliicolous lichen flora of Mexico. III. New species from Volcán San Martín Tuxtla (Sierra de Los Tuxtlas), Veracruz, with notes on Fellhanera santessonii". *Phyton*. **44** (2): 167–184.

1. **[^](#cite_ref-Safranek_&_Lücking_2005_12-0)** Safranek, William W.; Lücking, Robert (2005). "*Gyalectidium floridense*, a new foliicolous lichen from the Southeastern United States". *The Bryologist*. **108** (2): 295–297. [doi](/source/Doi_(identifier)):[10.1639/0007-2745(2005)108\[0295:GFANFL\]2.0.CO;2](https://doi.org/10.1639%2F0007-2745%282005%29108%5B0295%3AGFANFL%5D2.0.CO%3B2).

1. **[^](#cite_ref-Vězda_1994_13-0)** Vězda, A. (1994). "Neue foliicole Flechten II" [New foliicolous lichens II]. *Nova Hedwigia* (in German). **58**: 123–143.

1. **[^](#cite_ref-Lücking_&_Kalb_2000_14-0)** Lücking, R.; Kalb, K. (2000). "Foliikole Flechten aus Brasilien (vornehmlich Amazonien), inklusive einer Checkliste und Bemerkungen zu *Coenogonium* und *Dimerella* (Gyalectaceae)" [Foliicolous lichens from Brazil (primarily Amazonia), including a checklist and remarks on *Coenogonium* and *Dimerella* (Gyalectaceae)]. *Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie* (in German). **122** (1): 1–61.

1. ^ [***a***](#cite_ref-Herrera-Campos_&_Lücking_2003_15-0) [***b***](#cite_ref-Herrera-Campos_&_Lücking_2003_15-1) [***c***](#cite_ref-Herrera-Campos_&_Lücking_2003_15-2) Herrera-Campos, María De Los Angeles; Lücking, Robert (2003). "The foliicolous lichen flora of Mexico II. New species from the montane forest in Oaxaca and Puebla". *The Bryologist*. **106** (1): 1–8. [doi](/source/Doi_(identifier)):[10.1639/0007-2745(2003)106\[0001:TFLFOM\]2.0.CO;2](https://doi.org/10.1639%2F0007-2745%282003%29106%5B0001%3ATFLFOM%5D2.0.CO%3B2).

1. **[^](#cite_ref-Suto_&_Ohtani_2018_16-0)** Suto, Y.; Ohtani, S. (2018). "Two new species of foliicolous lichenized Ascomycota, *Asterothyrium sasae* (Asterothyriaceae) and *Gyalectidium shimanense* (Gomphillaceae) from Shimane-ken, Western Japan". *Lichenology*. **17** (2): 35–44.

1. ^ [***a***](#cite_ref-Lücking_et_al._2007_17-0) [***b***](#cite_ref-Lücking_et_al._2007_17-1) Lücking, Robert; Buck, William R.; Plata, Eimy Rivas (2007). "The lichen family Gomphillaceae (Ostropales) in eastern North America, with notes on hyphophore development in *Gomphillus* and *Gyalideopsis*". *The Bryologist*. **110** (4): 622–672. [doi](/source/Doi_(identifier)):[10.1639/0007-2745(2007)110\[622:TLFGOI\]2.0.CO;2](https://doi.org/10.1639%2F0007-2745%282007%29110%5B622%3ATLFGOI%5D2.0.CO%3B2).

1. **[^](#cite_ref-Buck_&_Sérusiaux_2000_18-0)** Buck, William R.; Sérusiaux, Emmanuël (2000). "*Gyalectidium yahriae*, sp. nov. (Lichenized Ascomycetes, Gomphillaceae) from Florida and Papua New Guinea". *The Bryologist*. **103** (1): 134–138. [doi](/source/Doi_(identifier)):[10.1639/0007-2745(2000)103\[0134:GYSNLA\]2.0.CO;2](https://doi.org/10.1639%2F0007-2745%282000%29103%5B0134%3AGYSNLA%5D2.0.CO%3B2). [hdl](/source/Hdl_(identifier)):[2268/175233](https://hdl.handle.net/2268%2F175233).

Taxon identifiers Gyalectidium Wikidata: Q5623943 Wikispecies: Gyalectidium AusLichen: 30017753 CoL: 4RP4 EoL: 16703 GBIF: 2601547 iNaturalist: 175775 IndexFungorum: 2158 IRMNG: 1065030 MycoBank: 2158 NCBI: 248113 NZOR: b4885619-83f2-45ee-bb74-63f943185b0e Open Tree of Life: 730337 PLANTS: GYALE3 SpeciesFungorum: 2158

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Adapted from the Wikipedia article [Gyalectidium](https://en.wikipedia.org/wiki/Gyalectidium) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Gyalectidium?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.