# Etmopterus

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Genus of sharks

Etmopterus Temporal range: 48–0 Ma[1] PreꞒ Ꞓ O S D C P T J K Pg N Middle Eocene to Present Velvet belly lanternshark (Etmopterus spinax) Green lanternshark (Etmopterus virens) Scientific classification Kingdom: Animalia Phylum: Chordata Class: Chondrichthyes Subclass: Elasmobranchii Division: Selachii Order: Squaliformes Family: Etmopteridae Genus: Etmopterus Rafinesque, 1810 Type species Etmopterus aculeatus Rafinesque, 1810

***Etmopterus*** is a [genus](/source/Genus) of [lantern sharks](/source/Etmopteridae) in the [squaliform](/source/Squaliformes) [family](/source/Family_(taxonomy)) Etmopteridae. They are found in [deep sea](/source/Deep_sea) ecosystems of the [Atlantic](/source/Atlantic_Ocean), [Indian](/source/Indian_Ocean) and [Pacific Oceans](/source/Pacific_Ocean).[2]

## Ecology

A number of species in this genus function as [host](/source/Host_(biology)) to the specialized [parasitic](/source/Parasitic) barnacle *[Anelasma squalicola](/source/Anelasma_squalicola)*, which embeds itself into the skin of the shark and extracts [nutrients](/source/Nutrient) from its bloodstream.[3]

## Species

*E. bigelowi*

*E. bullisi*

*E. carteri*

*E. gracilispinis*

*E. granulosus*

*E. hillianus*

*E. lucifer*

*E. perryi*

*E. polli*

*E. princeps*

*E. pusillus*

*E. robinsi*

There are currently 45 recognized species in this genus:

- *[Etmopterus alphus](/source/Etmopterus_alphus)* [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), [Straube](https://en.wikipedia.org/w/index.php?title=Nicolas_Straube&action=edit&redlink=1), [Leslie](https://en.wikipedia.org/w/index.php?title=Robin_W._Leslie&action=edit&redlink=1) & [Weigmann](https://en.wikipedia.org/w/index.php?title=Simon_Weigmann&action=edit&redlink=1), 2016 (whitecheek lanternshark)[4]

- *[Etmopterus baxteri](/source/Etmopterus_baxteri)* [Garrick](/source/Jack_Garrick), 1957 (New Zealand lanternshark)[5]

- *[Etmopterus benchleyi](/source/Etmopterus_benchleyi)* [Vásquez](https://en.wikipedia.org/w/index.php?title=Victoria_Elena_V%C3%A1squez&action=edit&redlink=1), [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1) & [D. J. Long](https://en.wikipedia.org/w/index.php?title=Douglas_J._Long&action=edit&redlink=1), 2015 (ninja lanternshark)[6]

- *[Etmopterus bigelowi](/source/Etmopterus_bigelowi)* [Shirai](/source/Shigeru_Shirai) & [Tachikawa](https://en.wikipedia.org/w/index.php?title=Hiroyuki_Tachikawa&action=edit&redlink=1), 1993 (blurred lanternshark)

- *[Etmopterus brachyurus](/source/Etmopterus_brachyurus)* [H. M. Smith](/source/Hugh_McCormick_Smith) & [Radcliffe](/source/Lewis_Radcliffe), 1912 (short-tail lanternshark)

- *[Etmopterus brosei](/source/Etmopterus_brosei)* [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), [Leslie](https://en.wikipedia.org/w/index.php?title=Robin_W._Leslie&action=edit&redlink=1) & [Weigmann](https://en.wikipedia.org/w/index.php?title=Simon_Weigmann&action=edit&redlink=1), 2021 (Barrie's lanternshark)[7]

- *[Etmopterus bullisi](/source/Etmopterus_bullisi)* [Bigelow](/source/Henry_Bryant_Bigelow) & [Schroeder](/source/William_Charles_Schroeder), 1957

- *[Etmopterus burgessi](/source/Etmopterus_burgessi)* [Schaaf-Da Silva](https://en.wikipedia.org/w/index.php?title=Jayna_Ann_Schaaf-Da_Silva&action=edit&redlink=1) & [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), 2006[8]

- *[Etmopterus carteri](/source/Etmopterus_carteri)* [S. Springer](/source/Stewart_Springer) & [G. H. Burgess](/source/George_H._Burgess), 1985 (cylindrical lanternshark)

- *[Etmopterus caudistigmus](/source/Etmopterus_caudistigmus)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002 (tail-spot lanternshark)

- *[Etmopterus compagnoi](/source/Etmopterus_compagnoi)* [R. Fricke](/source/Ronald_Fricke) & [Koch](https://en.wikipedia.org/w/index.php?title=Isabel_Koch&action=edit&redlink=1), 1990[9]

- *[Etmopterus decacuspidatus](/source/Etmopterus_decacuspidatus)* [W. L. Y. Chan](https://en.wikipedia.org/w/index.php?title=Chan_William_Lai-Yee&action=edit&redlink=1), 1966 (comb-tooth lanternshark)

- *[Etmopterus dianthus](/source/Etmopterus_dianthus)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002 (pink lanternshark)

- *[Etmopterus dislineatus](/source/Etmopterus_dislineatus)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002

- *[Etmopterus evansi](/source/Etmopterus_evansi)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002 (black-mouth lanternshark)

- *[Etmopterus fusus](/source/Etmopterus_fusus)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002 (pygmy lanternshark)

- *[Etmopterus gracilispinis](/source/Etmopterus_gracilispinis)* [G. Krefft](/source/Gerhard_Krefft), 1968 (broad-banded lanternshark)

- *[Etmopterus granulosus](/source/Etmopterus_granulosus)* ([Günther](/source/Albert_Karl_Ludwig_Gotthilf_G%C3%BCnther), 1880) (southern lanternshark)

- *[Etmopterus hillianus](/source/Etmopterus_hillianus)* ([Poey](/source/Felipe_Poey_y_Aloy), 1861) (Caribbean lanternshark)

- *[Etmopterus joungi](/source/Etmopterus_joungi)* [Knuckey](https://en.wikipedia.org/w/index.php?title=James_D._S._Knuckey&action=edit&redlink=1), [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1) & [G. H. Burgess](/source/George_H._Burgess), 2011 (short-fin smooth lanternshark)[10]

- *[Etmopterus lailae](/source/Etmopterus_lailae)* [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), [Papastamatiou](https://en.wikipedia.org/w/index.php?title=Yannis_Peter_Papastamatiou&action=edit&redlink=1), [Kajiura](https://en.wikipedia.org/w/index.php?title=Stephen_Michael_Kajiura&action=edit&redlink=1) & [Wetherbee](https://en.wikipedia.org/w/index.php?title=Bradley_M._Wetherbee&action=edit&redlink=1), 2017 (Laila's lanternshark)[11]

- *[Etmopterus litvinovi](/source/Etmopterus_litvinovi)* [Parin](/source/Nikolai_Vasilyevich_Parin) & [Kotlyar](/source/Aleksandr_Nicholaevich_Kotlyar), 1990 (small-eye lanternshark)

- *[Etmopterus lucifer](/source/Etmopterus_lucifer)* [D. S. Jordan](/source/David_Starr_Jordan) & [Snyder](/source/John_Otterbein_Snyder), 1902 (black-belly lanternshark)

- *[Etmopterus marshae](/source/Etmopterus_marshae)* [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1) & [Van Hees](https://en.wikipedia.org/w/index.php?title=Kelley_E._Van_Hees&action=edit&redlink=1) (Marsha's lanternshark)[12]

- *[Etmopterus molleri](/source/Etmopterus_molleri)* ([Whitley](/source/Gilbert_Percy_Whitley), 1939) (Moller's lanternshark or also Slendertail lanternshark)

- *[Etmopterus perryi](/source/Etmopterus_perryi)* [S. Springer](/source/Stewart_Springer) & [G. H. Burgess](/source/George_H._Burgess), 1985 (dwarf lanternshark)

- *[Etmopterus polli](/source/Etmopterus_polli)* [Bigelow](/source/Henry_Bryant_Bigelow), [Schroeder](/source/William_Charles_Schroeder) & [S. Springer](/source/Stewart_Springer), 1953 (African lanternshark)

- *[Etmopterus princeps](/source/Etmopterus_princeps)* [Collett](/source/Robert_Collett), 1904 (great lanternshark)

- *[Etmopterus pseudosqualiolus](/source/Etmopterus_pseudosqualiolus)* [Last](/source/Peter_Robert_Last), [G. H. Burgess](/source/George_H._Burgess) & [Séret](https://en.wikipedia.org/w/index.php?title=Bernard_S%C3%A9ret&action=edit&redlink=1), 2002 (false lanternshark)

- *[Etmopterus pusillus](/source/Etmopterus_pusillus)* ([R. T. Lowe](/source/Richard_Thomas_Lowe), 1839) (smooth lanternshark)

- *[Etmopterus pycnolepis](/source/Etmopterus_pycnolepis)* [Kotlyar](/source/Aleksandr_Nicholaevich_Kotlyar), 1990 (dense-scale lanternshark)

- *[Etmopterus robinsi](/source/Etmopterus_robinsi)* [Schofield](https://en.wikipedia.org/w/index.php?title=Pamela_J._Schofield&action=edit&redlink=1) & [G. H. Burgess](/source/George_H._Burgess), 1997 (West Indian lanternshark)

- *[Etmopterus samadiae](/source/Etmopterus_samadiae)* [W. T. White](/source/William_Toby_White), [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), [Mana](https://en.wikipedia.org/w/index.php?title=Ralph_Reeves_Mana&action=edit&redlink=1) & [Corrigan](https://en.wikipedia.org/w/index.php?title=Shannon_Leah_Corrigan&action=edit&redlink=1), 2017 (Papuan lanternshark)[13]

- *[Etmopterus schultzi](/source/Etmopterus_schultzi)* [Bigelow](/source/Henry_Bryant_Bigelow), [Schroeder](/source/William_Charles_Schroeder) & [S. Springer](/source/Stewart_Springer), 1953 (fringe-fin lanternshark)

- *[Etmopterus sculptus](/source/Etmopterus_sculptus)* [Ebert](https://en.wikipedia.org/w/index.php?title=David_A._Ebert&action=edit&redlink=1), [Compagno](/source/Leonard_Joseph_Victor_Compagno) & [De Vries](https://en.wikipedia.org/w/index.php?title=Marlee_J._De_Vries&action=edit&redlink=1), 2011 (sculpted lanternshark)[14]

- *[Etmopterus sentosus](/source/Etmopterus_sentosus)* [Bass](https://en.wikipedia.org/w/index.php?title=A._John_Bass&action=edit&redlink=1), [D'Aubrey](https://en.wikipedia.org/w/index.php?title=Jeannette_Dymock_D%27Aubrey&action=edit&redlink=1) & [Kistnasamy](https://en.wikipedia.org/w/index.php?title=Nadaraj_Kistnasamy&action=edit&redlink=1), 1976 (thorny lanternshark)

- *[Etmopterus sheikoi](/source/Etmopterus_sheikoi)* ([Dolganov](https://en.wikipedia.org/w/index.php?title=Vladimir_Nikolaevich_Dolganov&action=edit&redlink=1), 1986) (rasp-tooth dogfish)[15]

- *[Etmopterus spinax](/source/Etmopterus_spinax)* ([Linnaeus](/source/Carl_Linnaeus), [1758](/source/10th_edition_of_Systema_Naturae)) (velvet-belly lanternshark)

- *[Etmopterus splendidus](/source/Etmopterus_splendidus)* [Ka. Yano](https://en.wikipedia.org/w/index.php?title=Kazunari_Yano&action=edit&redlink=1), 1988 (splendid lanternshark)

- *[Etmopterus unicolor](/source/Etmopterus_unicolor)* ([Engelhardt](https://en.wikipedia.org/w/index.php?title=Robert_Engelhardt&action=edit&redlink=1), 1912) (bristled lanternshark)

- *[Etmopterus viator](/source/Etmopterus_viator)* [Straube](https://en.wikipedia.org/w/index.php?title=Nicolas_Straube&action=edit&redlink=1), 2011 (traveller lanternshark)[16]

- *[Etmopterus villosus](/source/Etmopterus_villosus)* [C. H. Gilbert](/source/Charles_Henry_Gilbert), 1905 (Hawaiian lanternshark)

- *[Etmopterus virens](/source/Etmopterus_virens)* [Bigelow](/source/Henry_Bryant_Bigelow), [Schroeder](/source/William_Charles_Schroeder) & [S. Springer](/source/Stewart_Springer), 1953 (green lanternshark)

- [*Etmopterus westraliensis*](/source/Etmopterus_westraliensis) Ng, White, Liu & Joung 2025

### Extinct species

There are at least two extinct species found in *Etmopterus*.[17]

- †*[Etmopterus acutidens](https://en.wikipedia.org/w/index.php?title=Etmopterus_acutidens&action=edit&redlink=1)* Casier 1966

- †*[Etmopterus cahuzaci](https://en.wikipedia.org/w/index.php?title=Etmopterus_cahuzaci&action=edit&redlink=1)* Adnet 2006

## See also

- [List of prehistoric cartilaginous fish](/source/List_of_prehistoric_cartilaginous_fish)

## References

1. **[^](#cite_ref-Adnet2001_1-0)** Adnet S., Cappetta H. (2001). "A palaeontological and phylogenetical analysis of squaliform sharks (Chondrichthyes: Squaliformes) based on dental characters". *Lethaia*. **34** (3): 234–248. [doi](/source/Doi_(identifier)):[10.1111/j.1502-3931.2001.tb00052.x](https://doi.org/10.1111%2Fj.1502-3931.2001.tb00052.x).

1. **[^](#cite_ref-FB_2-0)** [Froese, Rainer](/source/Rainer_Froese); [Pauly, Daniel](/source/Daniel_Pauly) (eds.). ["Species in genus *Etmopterus*"](https://www.fishbase.org/identification/SpeciesList.php?genus=Etmopterus). *[FishBase](/source/FishBase)*. June 2017 version.

1. **[^](#cite_ref-Yano2000_3-0)** Yano K., Musick J.A. (2000). "The effect of the mesoparasitic barnacle *Anelasma* on the development of reproductive organs of deep-sea squaloid sharks, *Centroscyllium* and *Etmopterus*". *Environmental Biology of Fishes*. **59** (3): 329–339. [doi](/source/Doi_(identifier)):[10.1023/A:1007649227422](https://doi.org/10.1023%2FA%3A1007649227422). [S2CID](/source/S2CID_(identifier)) [6049145](https://api.semanticscholar.org/CorpusID:6049145).

1. **[^](#cite_ref-Ebert2016_4-0)** Ebert D.A.; Straube N.; Leslie R.W.; Weigmann S. (2016). "*Etmopterus alphus* n. sp.: a new lanternshark (Squaliformes: Etmopteridae) from the south-western Indian Ocean". *African Journal of Marine Science*. **38** (3): 329–340. [doi](/source/Doi_(identifier)):[10.2989/1814232X.2016.1198275](https://doi.org/10.2989%2F1814232X.2016.1198275). [S2CID](/source/S2CID_(identifier)) [89547111](https://api.semanticscholar.org/CorpusID:89547111).

1. **[^](#cite_ref-5)** N. Bailly (2008). Bailly N (ed.). ["Etmopterus baxteri Garrick, 1957"](https://www.marinespecies.org/aphia.php?p=taxdetails&id=271627). *[FishBase](/source/FishBase)*. [World Register of Marine Species](/source/World_Register_of_Marine_Species). Retrieved October 11, 2017.

1. **[^](#cite_ref-Vasquez2015_6-0)** Vásquez V.E.; Ebert D.A.; Long D.J. (2015). ["*Etmopterus benchleyi* n. sp., a new lanternshark (Squaliformes: Etmopteridae) from the central eastern Pacific Ocean"](http://www.oceansciencefoundation.org/josf/josf17d.pdf) (PDF). *Journal of the Ocean Science Foundation*. **17**: 43–55.

1. **[^](#cite_ref-7)** Ebert, D.A.; Leslie, R.W.; Weigmann, S. (31 May 2021). "*Etmopterus brosei* sp. nov.: a new lanternshark (Squaliformes: Etmopteridae) from the southeastern Atlantic and southwestern Indian oceans, with a revised key to the *Etmopterus lucifer* clade". *Marine Biodiversity*. **51** (3). [doi](/source/Doi_(identifier)):[10.1007/s12526-021-01173-0](https://doi.org/10.1007%2Fs12526-021-01173-0). [S2CID](/source/S2CID_(identifier)) [236343587](https://api.semanticscholar.org/CorpusID:236343587).

1. **[^](#cite_ref-Silva2006_8-0)** Schaaf-Da Silva, J.A.; Ebert, D.A. (2006). "*Etmopterus burgessi* sp. nov., a new species of lanternshark (Squaliformes: Etmopteridae) from Taiwan". *Zootaxa*. **1373**: 53–64. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.1373.1.3](https://doi.org/10.11646%2Fzootaxa.1373.1.3).

1. **[^](#cite_ref-Straube2015_9-0)** Straube, N., Leslie, R.W., Clerkin, P.J., Ebert, D.A., Rochel, E., Corrigan, S., Li, C. & Naylor, G.J.P. (2015): [On the occurrence of the Southern Lanternshark, *Etmopterus granulosus*, off South Africa, with comments on the validity of *E. compagnoi*.](http://www.sciencedirect.com/science/article/pii/S0967064514001210) *Deep Sea Research Part II: Topical Studies in Oceanography, 115: 11-17.*

1. **[^](#cite_ref-Knuckey2011_10-0)** Knuckey, J.D.S., Ebert, D.A. & Burgess, G.H. (2011): [*Etmopterus joungi* n. sp., a new species of lanternshark (Squaliformes: Etmopteridae) from Taiwan.](https://aqua-aquapress.com/etmopterus-joungi-n-sp-a-new-species-of-lanternshark-squaliformes-etmopteridae-from-taiwan-2/) [Archived](https://web.archive.org/web/20201021221828/https://aqua-aquapress.com/etmopterus-joungi-n-sp-a-new-species-of-lanternshark-squaliformes-etmopteridae-from-taiwan-2/) 2020-10-21 at the [Wayback Machine](/source/Wayback_Machine) *aqua, International Journal of Ichthyology, 17 (2): 61-72.*

1. **[^](#cite_ref-Ebert2017_11-0)** Ebert D.A.; Papastamatiou Y.P.; Kajiura S.M.; Wetherbee B.M. (2017). ["*Etmopterus lailae* sp. nov., a new lanternshark (Squaliformes: Etmopteridae) from the Northwestern Hawaiian Islands"](https://web.archive.org/web/20181024103252/http://www.science.fau.edu/sharklab/pdfs/epkw17.pdf) (PDF). *Zootaxa*. **4237** (2): 371–382. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.4237.2.10](https://doi.org/10.11646%2Fzootaxa.4237.2.10). [PMID](/source/PMID_(identifier)) [28264297](https://pubmed.ncbi.nlm.nih.gov/28264297). Archived from [the original](http://www.science.fau.edu/sharklab/pdfs/epkw17.pdf) (PDF) on 2018-10-24. Retrieved 2017-09-26.

1. **[^](#cite_ref-ebertvanhees_12-0)** Ebert, David A.; Van Hees, Kelley E. (2018). "*Etmopterus marshae* sp. nov, a new lanternshark (Squaliformes: Etmopteridae) from the Philippine Islands, with a revised key to the *Etmopterus lucifer* clade". *Zootaxa*. **4508** (2): 197–210. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.4508.2.3](https://doi.org/10.11646%2Fzootaxa.4508.2.3). [PMID](/source/PMID_(identifier)) [30485973](https://pubmed.ncbi.nlm.nih.gov/30485973). [S2CID](/source/S2CID_(identifier)) [54122190](https://api.semanticscholar.org/CorpusID:54122190).

1. **[^](#cite_ref-White2017_13-0)** White W.T.; Ebert D.A.; Mana R.R.; Corrigan S. (2017). ["*Etmopterus samadiae* n. sp., a new lanternshark (Squaliformes: Etmopteridae) from Papua New Guinea"](http://www.mapress.com/j/zt/article/view/zootaxa.4244.3.3). *Zootaxa*. **4244** (3): 339–354. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.4244.3.3](https://doi.org/10.11646%2Fzootaxa.4244.3.3). [PMID](/source/PMID_(identifier)) [28610109](https://pubmed.ncbi.nlm.nih.gov/28610109).

1. **[^](#cite_ref-Ebert2011_14-0)** Ebert D.A.; Compagno L.J.V.; De Vries M.J. (2011). "A New Lanternshark (Squaliformes: Etmopteridae: *Etmopterus*) from Southern Africa". *Copeia*. **2011** (3): 379–384. [doi](/source/Doi_(identifier)):[10.1643/CI-09-183](https://doi.org/10.1643%2FCI-09-183). [S2CID](/source/S2CID_(identifier)) [83658773](https://api.semanticscholar.org/CorpusID:83658773).

1. **[^](#cite_ref-Straube2013_15-0)** Straube N.; White W.T.; Ho H.-C.; Rochel E.; Corrigan S.; Li C.; Naylor G.J.P. (2013). ["A DNA sequence-based identification checklist for Taiwanese chondrichthyans"](http://www.mapress.com/zootaxa/2013/f/zt03752p278.pdf) (PDF). *Zootaxa*. **3752** (1): 256–278. [doi](/source/Doi_(identifier)):[10.11646/zootaxa.3752.1.16](https://doi.org/10.11646%2Fzootaxa.3752.1.16). [PMID](/source/PMID_(identifier)) [25229119](https://pubmed.ncbi.nlm.nih.gov/25229119).

1. **[^](#cite_ref-Straube2011_16-0)** Straube, N.; Duhamel, G.; Gasco, N.; Kriwet, J.; Schliewen, U.K. (2011). "Description of a new deep-sea Lantern Shark *Etmopterus viator* sp. nov. (Squaliformes: Etmopteridae) from the Southern Hemisphere. *In*: Duhamel, G. & Welsford, D. (Eds.), *The Kerguelen Plateau: Marine Ecosystem and Fisheries.*". *Société Française d'Ichtyologie*: 137–150.

1. **[^](#cite_ref-17)** ["List of species/Extinct - complete"](http://shark-references.com/species/listAllFossil/E). *shark-references.com*. Retrieved September 25, 2017.

v t e Extant shark species Kingdom Animalia Phylum Chordata Subphylum Vertebrata Class Chondrichthyes Subclass Elasmobranchii Division Selachii Order Carcharhiniformes (Ground sharks) Hemigaleidae (Weasel sharks) Hemipristis Snaggletooth shark (H. elongata) Chaenogaleus Hooktooth shark (C. macrostoma) Hemigaleus Australian weasel shark (H. australiensis) Sicklefin weasel shark (H. microstoma) Paragaleus Whitetip weasel shark (P. leucolomatus) Atlantic weasel shark (P. pectoralis) Slender weasel shark (P. randalli) Straight-tooth weasel shark (P. tengi) Leptochariidae Leptocharias Barbeled houndshark (L. smithii) Proscylliidae (Finback sharks) Ctenacis Harlequin catshark (C. fehlmanni) Eridacnis Cuban ribbontail catshark (E. barbouri) Pygmy ribbontail catshark (E. radcliffei) African ribbontail catshark (E. sinuans) Proscyllium Graceful catshark (P. habereri) P. venustum Pseudotriakidae Gollum Slender smooth-hound (G. attenuatus) Pseudotriakis False catshark (P. microdon) Sphyrnidae (Hammerhead sharks) Eusphyra Winghead shark (E. blochii) Sphyrna Scalloped bonnethead (S. corona) Whitefin hammerhead (S. couardi) Scalloped hammerhead (S. lewini) Scoophead (S. media) Great hammerhead (S. mokarran) Bonnethead (S. tiburo) Smalleye hammerhead (S. tudes) Smooth hammerhead (S. zygaena) Triakidae (Houndsharks) Furgaleus Whiskery shark (F. macki) Galeorhinus School shark (G. galeus) Gogolia Sailback houndshark (G. filewoodi) Hemitriakis Deepwater sicklefin hound shark (H. abdita) Sicklefin hound shark (H. falcata) Japanese topeshark (H. japanica) Whitefin topeshark (H. leucoperiptera) Ocellate topeshark (Hemitriakis Sp.A) Hypogaleus Blacktip tope (H. hyugaensis) Iago Longnose houndshark (I. garricki) Bigeye houndshark (I. omanensis) Lowfin houndshark (Iago Sp.A) Mustelus (Smooth-hounds) M. albipinnis Gummy shark (M. antarcticus) Starry smooth-hound (M. asterias) Gray smooth-hound (M. californicus) Dusky smooth-hound (M. canis) Sharptooth smooth-hound (M. dorsalis) Striped smooth-hound (M. fasciatus) Spotless smooth-hound (M. griseus) Brown smooth-hound (M. henlei) Smalleye smooth-hound (M. higmani) Spotted estuary smooth-hound (M. lenticulatus) Sicklefin smooth-hound (M. lunulatus) Starspotted smooth-hound (M. manazo) Speckled smooth-hound (M. mento) M. minicanis Arabian smooth-hound (M. mosis) Common smooth-hound (M. mustelus) Narrowfin smooth-hound (M. norrisi) Whitespotted smooth-hound (M. palumbes) Blackspotted smooth-hound (M. punctulatus) M. ravidus Narrownose smooth-hound (M. schmitti) Gulf smoothhound (M. sinusmexicanus) Humpback smooth-hound (M. whitneyi) M. widodoi Scylliogaleus Flapnose houndshark (S. quecketti) Triakis Sharpfin houndshark (T. acutipinna) Spotted houndshark (T. maculata) Sharptooth houndshark (T. megalopterus) Banded houndshark (T. scyllium) Leopard shark (T. semifasciata) Carcharhinidae Large family listed below Scyliorhinidae Large family listed below Family Carcharhinidae (Requiem sharks) Carcharhinus Blacknose shark (C. acronotus) Silvertip shark (C. albimarginatus) Bignose shark (C. altimus) Graceful shark (C. amblyrhynchoides) Grey reef shark (C. amblyrhynchos) Pigeye shark (C. amboinensis) Borneo shark (C. borneensis) Copper shark (C. brachyurus) Spinner shark (C. brevipinna) Nervous shark (C. cautus) Whitecheek shark (C. dussumieri) Silky shark (C. falciformis) Creek whaler (C. fitzroyensis) Galapagos shark (C. galapagenisis) Pondicherry shark (C. hemiodon) Finetooth shark (C. isodon) Smoothtooth blacktip shark (C. leiodon) Bull shark (C. leucas) Blacktip shark (C. limbatus) Oceanic whitetip shark (C. longimanus) Hardnose shark (C. macloti) Blacktip reef shark (C. melanopterus) Dusky shark (C. obscurus) Caribbean reef shark (C. perezii) Sandbar shark (C. plumbeus) Smalltail shark (C. porosus) Blackspot shark (C. sealei) Night shark (C. signatus) Spottail shark (C. sorrah) Australian blacktip shark (C. tilstoni) Galeocerdo Tiger shark (G. cuvier) Glyphis (River sharks) Ganges shark (G. gangeticus) Northern river shark (G. garricki) Speartooth shark (G. glyphis) Irrawaddy river shark (G. siamensis) Borneo river shark (Glyphis sp. B) Isogomphodon Daggernose shark (I. oxyrhynchus) Lamiopsis Broadfin shark (L. temminckii) Loxodon Sliteye shark (L. macrorhinus) Nasolamia Whitenose shark (N. velox) Negaprion Sicklefin lemon shark (N. acutidens) Lemon shark (N. brevirostris) Prionace Blue shark (P. glauca) Rhizoprionodon Milk shark (R. acutus) Brazilian sharpnose shark (R. lalandii) Pacific sharpnose shark (R. longurio) Grey sharpnose shark (R. oligolinx) Caribbean sharpnose shark (R. porosus) Australian sharpnose shark (R. taylori) Atlantic sharpnose shark (R. terraenovae) Scoliodon Spadenose shark (S. laticaudus) Triaenodon Whitetip reef shark (T. obesus) Family Scyliorhinidae (Catsharks) Apristurus Flatnose cat shark (A. acanutus) A. albisoma A. aphyodes Atlantic ghost catshark (A. atlanticus) Brown catshark (A. brunneus) Hoary catshark (A. canutus) Flaccid catshark (A. exsanguis) A. fedorovi Humpback cat shark (A. gibbosus) Longfin catshark (A. herklotsi) Smallbelly catshark (A. indicus) A. internatus Broadnose catshark (A. investigatoris) Japanese catshark (A. japonicus) Longnose catshark (A. kampae) Iceland catshark (A. laurussonii) Longhead catshark (A. longicephalus) Flathead catshark (A. macrorhynchus) Broadmouth cat shark (A. macrostomus) Ghost catshark (A. manis) Black roughscale catshark (A. melanoasper) Smalleye catshark (A. microps) Smalldorsal cat shark (A. micropterygeus) Largenose catshark (A. nasutus) Smallfin catshark (A. parvipinnis) A. pinguis Spatulasnout catshark (A. platyrhynchus) Deepwater catshark (A. profundorum) Broadgill catshark (A. riveri) Saldanha catshark (A. saldanha) Pale catshark (A. sibogae) South China catshark (A. sinensis) Spongehead catshark (A. spongiceps) Panama ghost catshark (A. stenseni) Asymbolus Australian spotted catshark (A. analis) A. funebris Western spotted catshark (A. occiduus) Pale spotted catshark (A. pallidus) A. parvus A. rubiginosus Variegated catshark (A. submaculatus) Gulf catshark (A. vincenti) Atelomycterus A. baliensis Banded sand catshark (A. fasciatus) Australian marbled catshark (A. macleayi) Coral catshark (A. marmoratus) Aulohalaelurus New Caledonia catshark (A. kanakorum) Australian blackspotted catshark (A. labiosus) Cephaloscyllium Whitefin swellshark (C. albipinnum) Circle-blotch pygmy swellshark (C. circulopullum) Cook's swellshark (C. cooki) Reticulated swellshark (C. fasciatum) Formosa swellshark (C. formosanum) Australian reticulate swellshark (C. hicosellum) Draughtsboard shark (C. isabellum) Australian swellshark (C. laticeps) Spotted swellshark (C. maculatum) Leopard-spotted swellshark (C. pardelotum) Painted swellshark (C. pictum) Sarawak pygmy swellshark (C. sarawakensis) Flagtail swellshark (C. signourum) Indian swellshark (C. silasi) Speckled swellshark (C. speccum) Balloon shark (C. sufflans) Blotchy swellshark (C. umbratile) Saddled swellshark (C. variegatum) Swellshark (C. ventriosum) Narrowbar swellshark (C. zebrum) Cephalurus Lollipop catshark (C. cephalus) Figaro Australian sawtail catshark (F. boardmani) Northern sawtail catshark (F. striatus) Galeus Antilles catshark (G. antillensis) Roughtail catshark (G. arae) Atlantic sawtail cat shark (G. atlanticus) Longfin sawtail cat shark (G. cadenati) Gecko catshark (G. eastmani) Slender sawtail catshark (G. gracilis) Longnose sawtail cat shark (G. longirostris) Blackmouth catshark (G. melastomus) Southern sawtail catshark (G. mincaronei) Mouse catshark (G. murinus) Broadfin sawtail catshark (G. nipponensis) Peppered catshark (G. piperatus) African sawtail catshark (G. polli) G. priapus Blacktip sawtail catshark (G. sauteri) Dwarf sawtail catshark (G. schultzi) Springer's sawtail cat shark (G. springeri) Halaelurus Arabian catshark (H. alcockii) Speckled catshark (H. boesemani) Blackspotted catshark (H. buergeri) Dusky catshark (H. canescens) Broadhead cat shark (H. clevai) New Zealand catshark (H. dawsoni) Bristly catshark (H. hispidus) Spotless catshark (H. immaculatus) Lined catshark (H. lineatus) Mud catshark (H. lutarius) Tiger catshark (H. natalensis) Quagga catshark (H. quagga) Haploblepharus Puffadder shyshark (H. edwardsii) Brown shyshark (H. fuscus) Natal shyshark (H. kistnasamyi) Dark shyshark (H. pictus) Holohalaelurus H. favus H. grennian Crying izak (H. melanostigma) African spotted catshark (H. punctatus) Izak catshark (H. regani) Parmaturus White-tip catshark (P. albimarginatus) White-clasper catshark (P. albipenis) Beige catshark (P. bigus) Campeche catshark (P. campechiensis) Velvet catshark (P. lanatus) McMillan's catshark (P. macmillani) Blackgill catshark (P. melanobranchus) Salamander shark (P. pilosus) Filetail catshark (P. xaniurus) Shorttail catshark (Parmaturus sp. A) Pentanchus Onefin catshark (P. profundicolus) Poroderma Pyjama catshark (P. africanum) Leopard catshark (P. pantherinum) Schroederichthys Narrowmouthed catshark (S. bivius) Redspotted catshark (S. chilensis) Narrowtail catshark (S. maculatus) Lizard catshark (S. saurisqualus) Slender catshark (S. tenuis) Scyliorhinus Polkadot catshark (S. besnardi) Boa catshark (S. boa) Small-spotted catshark (S. canicula) Yellowspotted catshark (S. capensis) West African catshark (S. cervigoni) Comoro cat shark (S. comoroensis) Brownspotted catshark (S. garmani) Freckled catshark (S. haeckelii) Whitesaddled catshark (S. hesperius) Blotched catshark (S. meadi) Chain catshark (S. retifer) Nursehound (S. stellaris) Izu cat shark (S. tokubee) Cloudy catshark (S. torazame) Dwarf catshark (S. torrei) Order Echinorhiniformes (Bramble sharks) Echinorhinidae Echinorhinus Bramble shark (E. brucus) Prickly shark (E. cookei) Order Heterodontiformes (Bullhead sharks) Heterodontidae Heterodontus Horn shark (H. francisci) Crested bullhead shark (H. galeatus) Japanese bullhead shark (H. japonicus) Mexican hornshark (H. mexicanus) Oman bullhead shark (H. omanensis) Port Jackson shark (H. portusjacksoni) Galapagos bullhead shark (H. quoyi) Whitespotted bullhead shark (H. ramalheira) Zebra bullhead shark (H. zebra) Order Hexanchiformes Chlamydoselachidae Chlamydoselachus Frilled shark (C. anguineus) Southern African frilled shark (C. africana) Hexanchidae (Cow sharks) Heptranchias Sharpnose sevengill shark (H. perlo) Hexanchus Bluntnose sixgill shark (H. griseus) Bigeyed sixgill shark (H. nakamurai) Notorynchus Broadnose sevengill shark (N. cepedianus) Order Lamniformes (Mackerel sharks) Alopiidae Alopias (Thresher sharks) Pelagic thresher (A. pelagicus) Bigeye thresher (A. superciliosus) Common thresher (A. vulpinus) Alopias sp. (A. sp.) Cetorhinidae Cetorhinus Basking shark (C. maximus) Lamnidae Carcharodon Great white shark (C. carcharias) Isurus Shortfin mako shark (I. oxyrinchus) Longfin mako shark (I. paucus) Lamna Salmon shark (L. ditropis) Porbeagle (L. nasus) Megachasmidae Megachasma Megamouth shark (M. pelagios) Mitsukurinidae Mitsukurina Goblin shark (M. owstoni) Odontaspididae Carcharias Sand tiger shark (C. taurus) Odontaspis Smalltooth sand tiger (O. ferox) Bigeye sand tiger (O. noronhai) Pseudocarchariidae Pseudocarcharias Crocodile shark (P. kamoharai) Order Orectolobiformes (Carpet sharks) Brachaeluridae Brachaelurus Blind shark (B. waddi) Heteroscyllium Bluegrey carpetshark (H. colcloughi) Ginglymostomatidae (Nurse sharks) Ginglymostoma Nurse shark (G. cirratum) Nebrius Tawny nurse shark (N. ferrugineus) Pseudoginglymostoma Short-tail nurse shark (P. brevicaudatum) Hemiscylliidae (Bamboo sharks) Chiloscyllium Arabian carpetshark (C. arabicum) Burmese bamboo shark (C. burmensis) Bluespotted bamboo shark (C. caerulopunctatum) Grey bamboo shark (C. griseum) Hasselt's bamboo shark (C. hasseltii) Slender bamboo shark (C. indicum) Whitespotted bamboo shark (C. plagiosum) Brownbanded bamboo shark (C. punctatum) Hemiscyllium Indonesian speckled carpetshark (H. freycineti) H. galei Papuan epaulette shark (H. hallstromi) H. henryi Epaulette shark (H. ocellatum) Hooded carpetshark (H. strahani) Speckled carpetshark (H. trispeculare) Orectolobidae (Wobbegongs) Eucrossorhinus Tasselled wobbegong (E. dasypogon) Orectolobus Floral banded wobbegong (O. floridus) Banded wobbegong (O. halei) Western wobbegong (O. hutchinsi) Japanese wobbegong (O. japonicus) Spotted wobbegong (O. maculatus) Ornate wobbegong (O. ornatus) Dwarf spotted wobbegong (O. parvimaculatus) Network wobbegong (O. reticulatus) Northern wobbegong (O. wardi) Sutorectus Cobbler wobbegong (S. tentaculatus) Parascylliidae (Collared carpet sharks) Cirrhoscyllium Barbelthroat carpetshark (C. expolitum) Taiwan saddled carpetshark (C. formosanum) Saddle carpetshark (C. japonicum) Parascyllium Collared carpetshark (P. collare) Rusty carpetshark (P. ferrugineum) Ginger carpetshark (P. sparsimaculatum) Necklace carpetshark (P. variolatum) Rhincodontidae Rhincodon Whale shark (R. typus) Stegostomatidae Stegostoma Zebra shark (S. fasciatum) Order Pristiophoriformes (Sawsharks) Pristiophoridae Pliotrema Anna's sixgill sawshark (P. annae) Kaja's sixgill sawshark (P. kajae) Sixgill sawshark (P. warreni) Pristiophorus Longnose sawshark (P. cirratus) Tropical sawshark (P. delicatus) Japanese sawshark (P. japonicus) Philippine sawshark (P. lanae) African dwarf sawshark (P. nancyae) Shortnose sawshark (P. nudipinnis) Eastern Australian sawshark (P. peroniensis) Bahamas sawshark (P. schroederi) Order Squaliformes Centrophoridae (Gulper sharks) Centrophorus Needle dogfish (C. acus) Dwarf gulper shark (C. atromarginatus) Gulper shark (C. granulosus) Dumb gulper shark (C. harrissoni) Blackfin gulper shark (C. isodon) Lowfin gulper shark (C. lusitanicus) Smallfin gulper shark (C. moluccensis) Taiwan gulper shark (C. niaukang) Leafscale gulper shark (C. squamosus) Mosaic gulper shark (C. tessellatus) Little gulper shark (C. uyato) Deania Birdbeak dogfish (D. calcea) Rough longnose dogfish (D. hystricosa) Arrowhead dogfish (D. profundorum) Longsnout dogfish (D. quadrispinosum) Dalatiidae Euprotomicroides Taillight shark (E. zantedeschia) Heteroscymnoides Longnose pygmy shark (H. marleyi) Mollisquama Pocket shark (M. parini) Dalatias Kitefin shark (D. licha) Isistius Cookiecutter shark (I. brasiliensis) South China cookiecutter shark (I. labialis) Largetooth cookiecutter shark (I. plutodus) Euprotomicrus Pygmy shark (E. bispinatus) Squaliolus Smalleye pygmy shark (S. aliae) Spined pygmy shark (S. laticaudus) Etmopteridae Aculeola Hooktooth dogfish (A. nigra) Centroscyllium Highfin dogfish (C. excelsum) Black dogfish (C. fabricii) Granular dogfish (C. granulatum) Bareskin dogfish (C. kamoharai) Combtooth dogfish (C. nigrum) Ornate dogfish (C. ornatum) Whitefin dogfish (C. ritteri) Etmopterus (Lantern sharks) New Zealand lanternshark (E. baxteri) Blurred lanternshark (E. bigelowi) Shorttail lanternshark (E. brachyurus) Lined lanternshark (E. bullisi) E. burgessi Cylindrical lanternshark (E. carteri) Tailspot lanternshark (E. caudistigmus) Combtooth lanternshark (E. decacuspidatus) Pink lanternshark (E. dianthus) E. dislineatus Blackmouth lanternshark (E. evansi) Pygmy lanternshark (E. fusus) Broadbanded lanternshark (E. gracilispinis) Southern lanternshark (E. granulosus) Caribbean lanternshark (E. hillianus) Smalleye lantern shark (E. litvinovi) Blackbelly lanternshark (E. lucifer) Slendertail lanternshark (E. molleri) Dwarf lanternshark (E. perryi) African lanternshark (E. polli) Great lanternshark (E. princeps) False lanternshark (E. pseudosqualiolus) Smooth lanternshark (E. pusillus) Dense-scale lantern shark (E. pycnolepis) West Indian lanternshark (E. robinsi) Fringefin lanternshark (E. schultzi) Thorny lanternshark (E. sentosus) Velvet belly lantern shark (E. spinax) Splendid lanternshark (E. splendidus) Tasmanian lanternshark (E. tasmaniensis) Brown lanternshark (E. unicolor) Hawaiian lanternshark (E. villosus) Green lanternshark (E. virens) Miroscyllium Rasptooth dogfish (M. sheikoi) Trigonognathus Viper dogfish (T. kabeyai) Oxynotidae (Rough sharks) Oxynotus Prickly dogfish (O. bruniensis) Caribbean roughshark (O. caribbaeus) Angular roughshark (O. centrina) Japanese roughshark (O. japonicus) Sailfin roughshark (O. paradoxus) Somniosidae (Sleeper sharks) Centroscymnus Portuguese dogfish (C. coelolepis) Shortnose velvet dogfish (C. cryptacanthus) Roughskin dogfish (C. owstoni) Centroselachus Longnose velvet dogfish (C. crepidater) Proscymnodon Largespine velvet dogfish (P. macracanthus) Plunket shark (P. plunketi) Scymnodalatias Whitetail dogfish (S. albicauda) Azores dogfish (S. garricki) Sparsetooth dogfish (S. oligodon) Sherwood dogfish (S. sherwoodi) Scymnodon Smallmouth velvet dogfish (S. obscurus) Knifetooth dogfish (S. ringens) Somniosus Southern sleeper shark (S. antarcticus) Frog shark (S. longus ) Greenland shark (S. microcephalus) Pacific sleeper shark (S. pacificus) Little sleeper shark (S. rostratus) Zameus Japanese velvet dogfish (Z. ichiharai) Velvet dogfish (Z. squamulosus) Squalidae (Dogfish sharks) Cirrhigaleus Roughskin spurdog (C. asper) Mandarin dogfish (C. barbifer) Squalus (Spurdogs) Spiny dogfish (S. acanthias) Eastern highfin spurdog (S. albifrons) S. acutirostris Western highfin spurdog (S. altipinnis) Longnose spurdog (S. blainville) Fatspine spurdog (S. crassispinus) Cuban dogfish (S. cubensis) Eastern longnose spurdog (S. grahami) Japanese spurdog (S. japonicus) Shortnose spurdog (S. megalops) Blacktailed spurdog (S. melanurus) Shortspine spurdog (S. mitsukurii) Bartail spurdog (S. notocaudatus) Western longnose spurdog (S. nasutus) Cyrano spurdog (S. rancureli) Pacific spiny dogfish (S. suckleyi) Order Squatiniformes (Angelsharks) Squatinidae Squatina Sawback angelshark (S. aculeata) African angelshark (S. africana) Eastern Australian angelshark (Squatina albipunctata) Argentine angelshark (S. argentina) Chilean angelshark (S. armata) Australian angelshark (S. australis) Pacific angelshark (S. californica) Sand devil (S. dumeril) Taiwan angelshark (S. formosa) Angular angelshark (S. guggenheim) S. heteroptera Japanese angelshark (S. japonica) Indonesian angelshark (S. legnota) Cortez angelshark (S. mexicana) Clouded angelshark (S. nebulosa) Smoothback angelshark (S. oculata) S. punctata Western Australian angelshark (Squatina pseudocellata) Angelshark (S. squatina) Ornate angelshark (S. tergocellata) Ocellated angelshark (S. tergocellatoides)

Taxon identifiers Etmopterus Wikidata: Q2322953 Wikispecies: Etmopterus ADW: Etmopterus AFD: Etmopterus BOLD: 2984 CoL: 633H8 EoL: 46560241 GBIF: 2421025 iNaturalist: 88262 IRMNG: 1320715 ITIS: 160654 NBN: NHMSYS0021053513 NCBI: 335011 NZOR: e90d77c1-d990-4ea3-8c81-cae4e8f1d725 Open Tree of Life: 975972 Paleobiology Database: 34546 Plazi: D1086F11-FFDE-FF9C-3D98-FD79FB61FD76 WoRMS: 105775

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Adapted from the Wikipedia article [Etmopterus](https://en.wikipedia.org/wiki/Etmopterus) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Etmopterus?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.