# Errantia

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Subclass of annelid worms

Errantia Temporal range: Cambrian Stage 4–Recent PreꞒ Ꞓ O S D C P T J K Pg N [1] Alitta virens Scientific classification Kingdom: Animalia Phylum: Annelida Clade: Pleistoannelida Subclass: Errantia Audouin & H Milne Edwards 1832 Orders[2][3] Aciculata Eunicida Phyllodocida Protodriliformia Polygordiidae Protodrilida

**Errantia** is a diverse group of [marine](/source/Marine_life) [polychaete](/source/Polychaete) worms in the phylum [Annelida](/source/Annelida). Traditionally a [subclass](/source/Subclass_(biology)) of the [paraphyletic](/source/Paraphyletic) class [Polychaeta](/source/Polychaeta),[4] it is currently regarded as a [monophyletic](/source/Monophyletic) group within the larger [Pleistoannelida](/source/Pleistoannelida), composed of Errantia and [Sedentaria](/source/Sedentaria).[2] These worms are found worldwide in marine environments and brackish water.

## Phylogeny

The [phylogeny](/source/Phylogeny) of [polychaetes](/source/Polychaete) is slowly being resolved. Errantia and [Sedentaria](/source/Sedentaria) are the two biggest [clades](/source/Clade) of polychaetes, and together they compose clade [Pleistoannelida](/source/Pleistoannelida).[5] Two groups are nested within Errantia: [Aciculata](/source/Aciculata) ([Eunicida](/source/Eunicida) + [Phyllodocida](/source/Phyllodocida)) and [Protodriliformia](/source/Protodriliformia) ([small](/source/Progenesis) meiofaunal worms such as the [Protodrilida](/source/Protodrilida)).[6][3][2]

Historically, the order [Amphinomida](/source/Amphinomida) was part of this subclass. However, [phylogenetic analyses](/source/Phylogenetic_analyses) place [Amphinomida](/source/Amphinomida) inside a basal clade with [Sipunculida](/source/Sipunculida) and *[Lobatocerebrum](https://en.wikipedia.org/w/index.php?title=Lobatocerebrum&action=edit&redlink=1)*, and this clade is the [sister group](/source/Sister_group) to [Pleistoannelida](/source/Pleistoannelida).[3]

Amphinomida Lobatocerebrum Sipunculida Pleistoannelida Errantia Aciculata Eunicida Phyllodocida Protodriliformia Protodrilida Polygordiidae Sedentaria

Some [taxa](/source/Taxon), such as [Spintheridae](/source/Spintheridae) and [Myzostomida](/source/Myzostomida), are still difficult to place due to their long branching, but they likely belong to either Errantia or [Sedentaria](/source/Sedentaria).[3]

## Classification

### Historical

Errantia is, along with [Sedentaria](/source/Sedentaria), one of the two old orders of the [paraphyletic](/source/Paraphyletic) class "[Polychaeta](/source/Polychaeta)". In 1977 the zoologist Kristian Fauchald split Errantia into three orders: [Phyllodocida](/source/Phyllodocida), [Amphinomida](/source/Amphinomida) and [Eunicida](/source/Eunicida), giving way to this classification.[7]

- Order [Amphinomida](/source/Amphinomida) – 2 families ([Amphinomidae](/source/Amphinomidae), [Euphrosinidae](/source/Euphrosinidae))

- Order [Eunicida](/source/Eunicida) - Superfamily Eunicea – 7 families ([Onuphidae](/source/Onuphidae), [Eunicidae](/source/Eunicidae), [Lumbrineridae](/source/Lumbrineridae), [Iphitimidae](https://en.wikipedia.org/w/index.php?title=Iphitimidae&action=edit&redlink=1), [Arabellidae](https://en.wikipedia.org/w/index.php?title=Arabellidae&action=edit&redlink=1), [Lysaretidae](https://en.wikipedia.org/w/index.php?title=Lysaretidae&action=edit&redlink=1), [Dorvilleidae](/source/Dorvilleidae)) - Superfamily incertae sedis – 2 families ([Histriobdellidae](/source/Histriobdellidae), [Ichthytomidae](https://en.wikipedia.org/w/index.php?title=Ichthytomidae&action=edit&redlink=1))

- Order [Phyllodocida](/source/Phyllodocida) - Suborder [Phyllodociformia](/source/Phyllodociformia) – 4 families ([Phyllodocidae](/source/Phyllodocidae), [Alciopidae](/source/Alciopidae), [Lopadorrhynchidae](/source/Lopadorrhynchidae), [Pontodoridae](/source/Pontodoridae)) - Suborder [Aphroditiformia](https://en.wikipedia.org/w/index.php?title=Aphroditiformia&action=edit&redlink=1) - Superfamily [Aphroditacea](https://en.wikipedia.org/w/index.php?title=Aphroditacea&action=edit&redlink=1) – 6 families ([Aphroditidae](/source/Aphroditidae), [Polynoidae](/source/Polynoidae), [Polyodontidae](/source/Polyodontidae), [Pholoididae](https://en.wikipedia.org/w/index.php?title=Pholoididae&action=edit&redlink=1), [Eulepethidae](/source/Eulepethidae), [Sigalionidae](/source/Sigalionidae)) - Superfamily [Chrysopetalacea](https://en.wikipedia.org/w/index.php?title=Chrysopetalacea&action=edit&redlink=1) – 2 families ([Chrysopetalidae](/source/Chrysopetalidae), [Palmyridae](https://en.wikipedia.org/w/index.php?title=Palmyridae&action=edit&redlink=1)) - Superfamily [Pisionacea](https://en.wikipedia.org/w/index.php?title=Pisionacea&action=edit&redlink=1) – 1 family ([Pisionidae](/source/Pisionidae)) - Suborder [Nereidiformia](https://en.wikipedia.org/w/index.php?title=Nereidiformia&action=edit&redlink=1) – 6 families ([Hesionidae](/source/Hesionidae), [Pilargiidae](https://en.wikipedia.org/w/index.php?title=Pilargiidae&action=edit&redlink=1), [Syllidae](/source/Syllidae), [Calamyzidae](https://en.wikipedia.org/w/index.php?title=Calamyzidae&action=edit&redlink=1), [Nereididae](/source/Nereididae), [Antonbmunidae](https://en.wikipedia.org/w/index.php?title=Antonbmunidae&action=edit&redlink=1)) - Suborder [Glyceriformia](https://en.wikipedia.org/w/index.php?title=Glyceriformia&action=edit&redlink=1) – 3 families ([Glyceridae](/source/Glyceridae), [Goniadidae](/source/Goniadidae), [Lacydoniidae](/source/Lacydoniidae)) - Suborder incertae sedis – 5 families ([Iospilidae](/source/Iospilidae), [Nephtyidae](/source/Nephtyidae), [Sphaerodoridae](/source/Sphaerodoridae), [Tomopteridae](/source/Tomopteridae), [Typhloscolecidae](/source/Typhloscolecidae))

## References

1. **[^](#cite_ref-1)** Yang, Xiaoyu; Aguado, M. Teresa; Helm, Conrad; Zhang, Zhiqian; Bleidorn, Christoph (April 2024). ["New fossil of Gaoloufangchaeta advances the origin of Errantia (Annelida) to the early Cambrian"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11004674). *Royal Society Open Science*. **11** (4). [doi](/source/Doi_(identifier)):[10.1098/rsos.231580](https://doi.org/10.1098%2Frsos.231580). [ISSN](/source/ISSN_(identifier)) [2054-5703](https://search.worldcat.org/issn/2054-5703). [PMC](/source/PMC_(identifier)) [11004674](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11004674). [PMID](/source/PMID_(identifier)) [38601033](https://pubmed.ncbi.nlm.nih.gov/38601033).

1. ^ [***a***](#cite_ref-Handbook_Zoology_Annelida_1_2-0) [***b***](#cite_ref-Handbook_Zoology_Annelida_1_2-1) [***c***](#cite_ref-Handbook_Zoology_Annelida_1_2-2) Struck TH (2019). "Phylogeny". In Purschke G, Böggemann M, Westheide W (eds.). *Handbook of Zoology: Annelida*. Vol. 1: Annelida Basal Groups and Pleistoannelida, Sedentaria I. De Gruyter. pp. 37–68. [doi](/source/Doi_(identifier)):[10.1515/9783110291582-002](https://doi.org/10.1515%2F9783110291582-002). [ISBN](/source/ISBN_(identifier)) [9783110291469](https://en.wikipedia.org/wiki/Special:BookSources/9783110291469).

1. ^ [***a***](#cite_ref-AnnelidCurrent_3-0) [***b***](#cite_ref-AnnelidCurrent_3-1) [***c***](#cite_ref-AnnelidCurrent_3-2) [***d***](#cite_ref-AnnelidCurrent_3-3) Weigert A, Bleidorn C (2016). "Current status of annelid phylogeny". *Org Divers Evol*. **16** (2): 345–362. [doi](/source/Doi_(identifier)):[10.1007/s13127-016-0265-7](https://doi.org/10.1007%2Fs13127-016-0265-7).

1. **[^](#cite_ref-4)** ["Encyclopedia of Life"](https://web.archive.org/web/20101115153944/http://www.eol.org/pages/85). Archived from [the original](http://www.eol.org/pages/85) on 2010-11-15. Retrieved 2011-06-25.

1. **[^](#cite_ref-Two_adaptive_routes_Annelida_5-0)** Struck TH, Golombek A, Weigert A, Franke FA, Westheide W, Purschke G, Bleidorn C, Halanych KM (3 August 2015). ["The evolution of annelids reveals two adaptive routes to the interstitial realm"](https://doi.org/10.1016%2Fj.cub.2015.06.007). *Curr Biol*. **25** (15): 1993–1999. [Bibcode](/source/Bibcode_(identifier)):[2015CBio...25.1993S](https://ui.adsabs.harvard.edu/abs/2015CBio...25.1993S). [doi](/source/Doi_(identifier)):[10.1016/j.cub.2015.06.007](https://doi.org/10.1016%2Fj.cub.2015.06.007). [PMID](/source/PMID_(identifier)) [26212885](https://pubmed.ncbi.nlm.nih.gov/26212885).

1. **[^](#cite_ref-Articulating_Archiannelids_6-0)** Andrade, Sónia C.S.; Novo, Marta; Kawauchi, Gisele Y.; Worsaae, Katrine; Pleijel, Fredrik; Giribet, Gonzalo; Rouse, Greg W. (November 2015). ["Articulating "Archiannelids": Phylogenomics and Annelid Relationships, with Emphasis on Meiofaunal Taxa"](https://doi.org/10.1093%2Fmolbev%2Fmsv157). *Molecular Biology and Evolution*. **32** (11): 2860–2875. [doi](/source/Doi_(identifier)):[10.1093/molbev/msv157](https://doi.org/10.1093%2Fmolbev%2Fmsv157). [PMID](/source/PMID_(identifier)) [26205969](https://pubmed.ncbi.nlm.nih.gov/26205969).

1. **[^](#cite_ref-Fauchald_Polychaetes_7-0)** Fauchald, Kristian (3 February 1977), [*The polychaete worms, definitions and keys to the orders, families and genera*](http://www.vliz.be/imisdocs/publications/123110.pdf) (PDF), Science Series, vol. 28, Natural History Museum of Los Angeles County: Los Angeles, CA (USA), pp. 1–188

Taxon identifiers Errantia Wikidata: Q6759542 Wikispecies: Errantia EoL: 39694098 ITIS: 914194 NCBI: 105390 WoRMS: 754174 Aciculata Wikidata: Q2823177 Wikispecies: Aciculata ADW: Aciculata BOLD: 125392 IRMNG: 12063 ITIS: 555670 NBN: NHMSYS0000065669 Paleobiology Database: 91491 WoRMS: 155090

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Adapted from the Wikipedia article [Errantia](https://en.wikipedia.org/wiki/Errantia) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Errantia?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.