# Dothideomycetidae > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Dothideomycetidae > Markdown URL: https://mediated.wiki/source/Dothideomycetidae.md > Source: https://en.wikipedia.org/wiki/Dothideomycetidae > Source revision: 1289996274 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) **Dothideomycetidae** is a fungal subclass in the class [Dothideomycetes](/source/Dothideomycetes) The cavities of the [sexual structures](/source/Ascomata) do not have vertical cells ([paraphyses](/source/Paraphyses), [pseudoparaphyses](/source/Pseudoparaphyses) or paraphysoids[1]) growing between the sac-like cells bearing the sexual spores ([asci](/source/Ascus)). ## Description Members of the Dothideomycetidae form small to medium fruiting bodies ([ascomata](/source/Ascomata)) that develop either within the [host](/source/Host_(biology)) tissue or [substrate](/source/Substrate_(biology)) or, in some species, burst through to become visible at the surface; a minority are superficially seated from the outset. Each ascoma may comprise a single cavity (locule) or several interconnected chambers where the spore-bearing sacs ([asci](/source/Ascus)) are housed. A minute pore (the [ostiole](/source/Ostiole)) usually connects the cavity to the outside air, and its short lining filaments () can sometimes be seen under the microscope. The surrounding wall is often built of densely packed, brick-like fungal cells (), but the inner space lacks the long sterile threads () found in many other ascomycetes.[2] The asci themselves display a wide range of outlines—from nearly spherical through egg-shaped and club-shaped to elongated cylinders. Inside, the [spores](/source/Spore) vary from colourless to dark brown and may be single-celled, divided by one or several cross walls ([septa](/source/Septum)), or partitioned in both directions to give a (brick-work) appearance. In addition to [sexual reproduction](/source/Sexual_reproduction), Dothideomycetidae exhibit [asexual](/source/Asexual_reproduction) stages: they may produce [conidia](/source/Conidia) inside flask-like chambers embedded in tissue (coelomycetous [anamorphs](/source/Teleomorph,_anamorph_and_holomorph)) or openly on thread-like [hyphae](/source/Hypha) ([hyphomycetous](/source/Hyphomycetes) anamorphs).[2] ## Classification According to *2024 Outline of Fungi*, the Dothideomycetidae consists of the following orders:[3] - [Arthrocatenales](/source/Arthrocatenales) [4] - [Aureoconidiellales](/source/Aureoconidiellales) [5] - [Capnodiales](/source/Capnodiales) - [Cladosporiales](/source/Cladosporiales) [5] - [Comminutisporales](/source/Comminutisporales) [5] - [Dothideales](/source/Dothideales) - [Mycosphaerellales](/source/Mycosphaerellales) - [Myriangiales](/source/Myriangiales) - [Neophaeothecales](/source/Neophaeothecales) [5] - [Phaeothecales](/source/Phaeothecales) [5] - [Racodiales](/source/Racodiales) [5] ## Genera *incertae cedis* - *[Aenigmatomyces](/source/Aenigmatomyces)* [6] ## References 1. Eriksson OE (1981). "The families of bitunicate ascomycetes". *Opera Botanica*. **60**: 1–220. 1. Adl, Sina M.; Simpson, Alastair G. B.; Lane, Christopher E.; Lukeš, Julius; Bass, David; Bowser, Samuel S.; Brown, Matthew W.; Burki, Fabien; Dunthorn, Micah; Hampl, Vladimir; Heiss, Aaron; Hoppenrath, Mona; Lara, Enrique; le Gall, Line; Lynn, Denis H.; McManus, Hilary; Mitchell, Edward A.D.; Mozley‐Stanridge, Sharon E.; Parfrey, Laura W.; Pawlowski, Jan; Rueckert, Sonja; Shadwick, Laura; Schoch, Conrad L.; Smirnov, Alexey; Spiegel, Frederick W. (2012). ["The revised classification of eukaryotes"](https://onlinelibrary.wiley.com/doi/pdfdirect/10.1111/j.1550-7408.2012.00644.x). *Journal of Eukaryotic Microbiology*. **59** (5): 429–514. [doi:10.1111/j.1550-7408.2012.00644.x](https://doi.org/10.1111/j.1550-7408.2012.00644.x). [PMC 3483872](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3483872). [PMID 23020233](https://pubmed.ncbi.nlm.nih.gov/23020233). Retrieved 2025-05-08. 1. Hyde, K.D.; Noorabadi, M.T.; Thiyagaraja, V.; He, M.Q.; Johnston, P.R.; Wijesinghe, S.N. et al. (2024). ["The 2024 Outline of Fungi and fungus-like taxa"](https://www.researchgate.net/publication/385939154). *Mycosphere*. **15** (1): 5146–6239 [5189]. [doi:10.5943/mycosphere/15/1/25](https://doi.org/10.5943/mycosphere/15/1/25). [hdl:1854/LU-8660838](https://hdl.handle.net/1854/LU-8660838) 1. Piątek, Marcin; Stryjak-Bogacka, Monika; Czachura, Paweł (2024). "Arthrocatenales, a new order of extremophilic fungi in the Dothideomycetes". *MycoKeys*. **108**: 47–74. [doi:10.3897/mycokeys.108.128033](https://doi.org/10.3897/mycokeys.108.128033). [PMC 11362667](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11362667). [PMID 39220356](https://pubmed.ncbi.nlm.nih.gov/39220356) 1. Abdollahzadeh, J.; Groenewald, J.Z.; Coetzee, M.P.A.; Wingfield, M.J.; Crous, P.W. (2020). "Evolution of lifestyles in Capnodiales". *Studies in Mycology*. **95**: 381–414. [doi:10.1016/j.simyco.2020.02.004](https://doi.org/10.1016/j.simyco.2020.02.004). [PMC 7426231](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7426231). [PMID 32855743](https://pubmed.ncbi.nlm.nih.gov/32855743) 1. Ruiz, Rafael F. Castaneda & Kendrick, Bryce (1993). "*Aenigmatomyces*, an enigmatic new genus of fungi from Algonquin Park, Canada". *Mycologia*. **85** (6): 1023–1027. [doi:10.2307/3760686](https://doi.org/10.2307/3760686). [JSTOR 3760686](https://www.jstor.org/stable/3760686) --- Adapted from the Wikipedia article [Dothideomycetidae](https://en.wikipedia.org/wiki/Dothideomycetidae) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Dothideomycetidae?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.