# Devernodes > Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Devernodes > Markdown URL: https://mediated.wiki/source/Devernodes.md > Source: https://en.wikipedia.org/wiki/Devernodes > Source revision: 1337734818 > License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/) ***Devernodes***[1] is a genus of flightless [weevils](/source/Weevil) belonging to the family [Curculionidae](/source/Curculionidae), subfamily [Molytinae](/source/Molytinae) and tribe Lymantini. First described in 2018 by [Vasily Vikotorivich Grebennikov](https://species.wikimedia.org/wiki/Vasily_Viktorovich_Grebennikov), it comprises weevil species found in leaf-litter habitats in [China](/source/China), [Malaysia](/source/Malaysia) and [Vietnam](/source/Vietnam).[1] Initially, the genus was believed to belong to the Asian [Stromboscerini](/source/Stromboscerini) tribe of the [Dryophthorinae](/source/Dryophthorinae) subfamily. Later *Devernodes* was placed in the Lymantini tribe due to its unique head constriction separating the eye-bearing rostrum from its head capsule and complete female hemisternites. These features, particularly the former, are unknown in Stromboscerini but are diagnostic features of the Lymantini tribe found in the Americas. It was later confirmed in 2022 through a DNA and morphological analysis that *Devernodes* belongs to the Lymantini tribe.[2] As such, the *Devernodes* represent the first record of *Lymantini* weevils found outside the [Americas](/source/Americas). ## Discovery *Devernodes* was first discovered in 2009 during fieldwork on the forested slopes of Mount Emei Shan in [Sichuan](/source/Sichuan), [China](/source/China). Additional species were also discovered in [Vietnam](/source/Vietnam) and [Malaysia](/source/Malaysia).[1] ## Taxonomy *Devernodes* is a [genus](/source/Genus) of [weevils](/source/Weevil) - [arthropods](/source/Arthropod) in the [insect](/source/Insect) family [Curculionidae](/source/Curculionidae), subfamily *[Molytinae](/source/Molytinae)*, and tribe *Lymantini*. This genus was established in 2018 by Dr. Vasily V. Grebennikov following the discovery of five new species from [Southeast Asia](/source/Southeast_Asia): *Devernodes* *alkippe, Devernodes* *asteria, Devernodes* *chthonia (the type species), Devernodes* *drimo and Devernodes* *methone* [1]*.* The genus *Devernodes* is notable because it represents the first known occurrence of *Lymantini* weevils outside the [Americas](/source/Americas), expanding the known geographic range of the tribe beyond its previously recognised limits. ## Phylogeny The Lymantini tribe initially included several genera found primarily in the [Americas](/source/Americas). The results of a phylogenetic analysis in 2022[2] confirmed that *Devernodes* are more closely related to the American Lymantini subtribe Lymantina than to any Asian weevil tribe, particularly the [Stromboscerini](/source/Stromboscerini) tribe, to which the first specimens were initially assigned to prior to their identification as a separate genus. The presence of *Devernodes* in Southeast Asia, despite all other *Lymantina* being American, suggests what scientists call amphi-Pacific [disjunct distribution](/source/Disjunct_distribution). This term describes closely related organisms that exist on opposite sides of the Pacific Ocean despite the vast geographical distance between them. This puzzle was resolved through a biogeographic hypothesis: The ancestors of *Devernodes* likely dispersed overland across Arctic land bridges (which are now submerged) during warmer periods of the [Cenozoic Era](/source/Cenozoic) (66 million years ago until today), specifically during the [Eocene](/source/Eocene). Thus Lymantini populations were established in Asia before climatic cooling divided the continents and fragmented the populations.[2] One of which eventually evolved into the current genus *Devernodes*. ## Species The genus *Devernodes* comprises five species:[1] - *Devernodes alkipipe* Grebennikov, 2018 (first recorded on [Mount Emei](/source/Mount_Emei) in [Sichuan](/source/Sichuan), [China](/source/China)) - *Devernodes asteria* Grebennikov, 2018 (first recorded in the [Tam Dao](/source/Tam_%C4%90%E1%BA%A3o_National_Park) Mountains of northern [Vietnam](/source/Vietnam)) - *Devernodes* *chthonia* Grebennikov, 2018 (first recorded in the [Tam Dao](/source/Tam_%C4%90%E1%BA%A3o_National_Park) Mountains of northern [Vietnam](/source/Vietnam)) - *Devernodes drimo* Grebennikov, 2018 (first recorded in [Pasoh Forest Reserve](/source/Pasoh_Forest_Reserve) in [Malaysia](/source/Malaysia)) - *Devernodes methone* Grebennikov, 2018 (first recorded at [Tanah Rata](/source/Tanah_Rata), [Malaysia](/source/Malaysia)) ## References 1. Grebennikov, Vasily V. (2018-10-16). ["Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia"](https://www.mapress.com/zt/article/view/zootaxa.4500.3.4). *Zootaxa*. **4500** (3): 363–380. [doi:10.11646/zootaxa.4500.3.4](https://doi.org/10.11646/zootaxa.4500.3.4). [ISSN 1175-5334](https://www.worldcat.org/issn/1175-5334) 1. Grebennikov, Vasily V. & Anderson, Robert S. (2022-12-31). ["Phylogeny, diversity and biogeography of flightless amphi-Pacific lymantine weevils (Coleoptera: Curculionidae: Molytinae)"](https://www.aemnp.eu/acta-entomologica/62-2/1928/phylogeny-diversity-and-biogeography-of-flightless-amphi-pacific-lymantine-weevils-coleoptera-curculionidae-molytinae.html). *Acta Entomologica Musei Nationalis Pragae*. [doi:10.37520/aemnp.2022.023](https://doi.org/10.37520/aemnp.2022.023). [ISSN 1804-6487](https://www.worldcat.org/issn/1804-6487) --- Adapted from the Wikipedia article [Devernodes](https://en.wikipedia.org/wiki/Devernodes) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Devernodes?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.