# Cybaeidae

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Family of spiders

Cybaeidae Temporal range: Palaeogene–present PreꞒ Ꞓ O S D C P T J K Pg N Cybaeus charlesi Dirksia cinctipes in Canada Scientific classification Kingdom: Animalia Phylum: Arthropoda Subphylum: Chelicerata Class: Arachnida Order: Araneae Infraorder: Araneomorphae Family: Cybaeidae Banks, 1892 Diversity 24 genera, 303 species

**Cybaeidae** is a [family](/source/Family_(taxonomy)) of [spiders](/source/Spider) first described by [Nathan Banks](/source/Nathan_Banks) in 1892.[1] There are small to large sized [entelegyne](/source/Entelegynae) spiders, which are [ecribellate](/source/Ecribellate).[2] The [diving bell spider](/source/Diving_bell_spider) or water spider *Argyroneta aquatica* was previously included in this family, but is now in the family [Dictynidae](/source/Dictynidae).[3][4] The distribution of this species resembles closely a typical [Holarctic](/source/Holarctic_realm) biogeography, with all of their species being found in the northern hemisphere. Where they have a tendency to live beneath rocks or woody debris in shaded and cool forest habitats, but they can also be found in caves, ant nests, and moss on tree trunks.[2] They are morphologically defined as spiders which have three claws, a single row of tarsal [trichobothria](/source/Trichobothria), and posterior lateral [spinnerets](/source/Spinneret) which lack a [cribellum](/source/Cribellum).[2]

Most of the cybaeid [genera](/source/Genus) are very species rich, as they have a large number of very short-range endemic species.[2]

## Genera

As of January 2026[\[update\]](https://en.wikipedia.org/w/index.php?title=Cybaeidae&action=edit), this family includes 24 genera and 303 species:[5]

- *[Allocybaeina](/source/Allocybaeina)* Bennett, 2020 – [Japan](/source/Japan), [United States](/source/United_States)

- *[Blabomma](/source/Blabomma)* [Chamberlin](/source/Ralph_Vary_Chamberlin) & Ivie, 1937 – [Korea](/source/Korea), North America

- *[Calymmaria](/source/Calymmaria)* Chamberlin & Ivie, 1937 – North America

- *[Cryphoeca](/source/Cryphoeca)* [Thorell](/source/Tamerlan_Thorell), 1870 – Japan, [Turkey](/source/Turkey), Europe, North America

- *[Cryphoecina](/source/Cryphoecina)* Deltshev, 1997 – [Montenegro](/source/Montenegro)

- *[Cybaeina](/source/Cybaeina)* Chamberlin & Ivie, 1932 – Korea, North America

- *[Cybaeota](/source/Cybaeota)* Chamberlin & Ivie, 1933 – [Russia](/source/Russia), North America

- *[Cybaeozyga](/source/Cybaeozyga)* Chamberlin & Ivie, 1937 – United States

- *[Cybaeus](/source/Cybaeus)* [L. Koch](/source/Ludwig_Carl_Christian_Koch), 1868 – Asia, Europe, North America, [Peru](/source/Peru)

- *[Dirksia](/source/Dirksia)* Chamberlin & Ivie, 1942 – [France](/source/France), North America

- *[Ethobuella](/source/Ethobuella)* Chamberlin & Ivie, 1937 – North America

- *[Guicybaeus](/source/Guicybaeus)* Wang, Chen, Yang & Zhang, 2023 – [China](/source/China)

- *[Mastigusa](/source/Mastigusa)* Menge, 1854 – [Algeria](/source/Algeria), [Georgia](/source/Georgia_(country)), [Iran](/source/Iran), Turkey, Europe, Britain

- *[Neocryphoeca](/source/Neocryphoeca)* Roth, 1970 – United States

- *[Neocybaeina](/source/Neocybaeina)* Bennett, 2023 – United States

- *[Pseudocybaeota](/source/Pseudocybaeota)* Bennett, 2022 – United States

- *[Rothaeina](/source/Rothaeina)* Bennett, 2023 – United States

- *[Sincybaeus](/source/Sincybaeus)* Wang & Zhang, 2022 – China, Japan

- *[Siskiyu](/source/Siskiyu)* [Hedin](https://species.wikimedia.org/wiki/Marshal_Hedin), Ramírez & Monjaraz-Ruedas, 2025 – United States

- *[Symposia](/source/Symposia)* [Simon](/source/Eug%C3%A8ne_Simon), 1898 – [Colombia](/source/Colombia), [Venezuela](/source/Venezuela)

- *[Tuberta](/source/Tuberta)* Simon, 1884 – [Azerbaijan](/source/Azerbaijan), Turkey, Europe, Britain

- *[Vagellia](/source/Vagellia)* Simon, 1899 – [Indonesia](/source/Indonesia)

- *[Willisus](/source/Willisus)* Roth, 1981 – United States

- *[Yorima](/source/Yorima)* Chamberlin & Ivie, 1942 – [Cuba](/source/Cuba), United States

A fossil genus is also placed in this family:[6]

- †*[Vectaraneus](https://en.wikipedia.org/w/index.php?title=Vectaraneus&action=edit&redlink=1)* Selden, 2003 (Isle of Wight)

## References

1. **[^](#cite_ref-banks1892_1-0)** Banks, N. (1892). ["A classification of North American spiders"](https://zenodo.org/record/2460991). *The Canadian Entomologist*. **24** (4): 88–97. [doi](/source/Doi_(identifier)):[10.4039/Ent2488-4](https://doi.org/10.4039%2FEnt2488-4).

1. ^ [***a***](#cite_ref-hedin2025_2-0) [***b***](#cite_ref-hedin2025_2-1) [***c***](#cite_ref-hedin2025_2-2) [***d***](#cite_ref-hedin2025_2-3) Hedin, Marshal; Ramírez, Martín J.; Monjaraz-Ruedas, Rodrigo (2025-02-06). ["Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province"](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11826229). *ZooKeys* (1226): 47–75. [Bibcode](/source/Bibcode_(identifier)):[2025ZooK.1226...47H](https://ui.adsabs.harvard.edu/abs/2025ZooK.1226...47H). [doi](/source/Doi_(identifier)):[10.3897/zookeys.1226.140204](https://doi.org/10.3897%2Fzookeys.1226.140204). [ISSN](/source/ISSN_(identifier)) [1313-2970](https://search.worldcat.org/issn/1313-2970). [PMC](/source/PMC_(identifier)) [11826229](https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11826229). [PMID](/source/PMID_(identifier)) [39959859](https://pubmed.ncbi.nlm.nih.gov/39959859).

1. **[^](#cite_ref-WSC_s8306_3-0)** ["Taxon details *Argyroneta aquatica* (Clerck, 1757)"](https://www.wsc.nmbe.ch/species/8306), *World Spider Catalog*, Natural History Museum Bern, retrieved 2020-12-21

1. **[^](#cite_ref-wheeler2016_4-0)** Wheeler, Ward C.; Coddington, Jonathan A.; Crowley, Louise M.; Dimitrov, Dimitar; Goloboff, Pablo A.; Griswold, Charles E.; Hormiga, Gustavo; Prendini, Lorenzo; Ramírez, Martín J.; Sierwald, Petra; Almeida-Silva, Lina; Alvarez-Padilla, Fernando; Arnedo, Miquel A.; Benavides Silva, Ligia R.; Benjamin, Suresh P.; Bond, Jason E.; Grismado, Cristian J.; Hasan, Emile; Hedin, Marshal; Izquierdo, Matías A.; Labarque, Facundo M.; Ledford, Joel; Lopardo, Lara; Maddison, Wayne P.; Miller, Jeremy A.; Piacentini, Luis N.; Platnick, Norman I.; Polotow, Daniele; Silva-Dávila, Diana; Scharff, Nikolaj; Szűts, Tamás; Ubick, Darrell; Vink, Cor J.; Wood, Hannah M. & Zhang, Junxia (2016). ["The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling"](https://doi.org/10.1111%2Fcla.12182). *Cladistics*. **33** (6): 574–616. [doi](/source/Doi_(identifier)):[10.1111/cla.12182](https://doi.org/10.1111%2Fcla.12182). [PMID](/source/PMID_(identifier)) [34724759](https://pubmed.ncbi.nlm.nih.gov/34724759). [S2CID](/source/S2CID_(identifier)) [35535038](https://api.semanticscholar.org/CorpusID:35535038).

1. **[^](#cite_ref-wsc_5-0)** ["Family Cybaeidae Banks, 1892"](https://wsc.nmbe.ch/lsid/urn:lsid:nmbe.ch:spiderfam:000022). World Spider Catalog. [doi](/source/Doi_(identifier)):[10.24436/2](https://doi.org/10.24436%2F2). Retrieved 2026-01-22.

1. **[^](#cite_ref-selden2001_6-0)** Selden, Paul A. (July 2001). ["Eocene Spiders from the Isle of Wight With Preserved Respiratory Structures"](https://kuscholarworks.ku.edu/bitstream/handle/1808/8359/Selden_2001_Eocene%20spiders%20Isle%20of%20Wight.pdf?sequence=1) (PDF). *Palaeontology*. **44** (4): 695–729. [Bibcode](/source/Bibcode_(identifier)):[2001Palgy..44..695S](https://ui.adsabs.harvard.edu/abs/2001Palgy..44..695S). [doi](/source/Doi_(identifier)):[10.1111/1475-4983.00199](https://doi.org/10.1111%2F1475-4983.00199).

## External links

- [Citizen science observations for Cybaeidae](https://www.inaturalist.org/taxa/52950) at [iNaturalist](/source/INaturalist)

v t e Extant Araneae families Kingdom: Animalia Phylum: Arthropoda Subphylum: Chelicerata Class: Arachnida Suborder Mesothelae Liphistiidae (segmented spiders) Liphistius Heptathelinae Suborder Opisthothelae Mygalomorphae Actinopodidae (mouse spiders and relatives) Anamidae Antrodiaetidae (folding trapdoor spiders) Atracidae (Australian funnel-web spiders) Atypidae (atypical tarantulas or purseweb spiders) Barychelidae (brushed trapdoor spiders) Bemmeridae Ctenizidae (cork-lid trapdoor spiders) Cyrtaucheniidae (wafer trapdoor spiders) Dipluridae (funnel-web tarantulas) Entypesidae Euagridae Euctenizidae Halonoproctidae Hexathelidae (funnel-webs or venomous funnel-web tarantulas) Hexurellidae Idiopidae Ischnothelidae Macrothelidae Mecicobothriidae (dwarf tarantulas) Megahexuridae Microhexuridae Microstigmatidae Migidae (tree trapdoor spiders) Nemesiidae (funnel-web tarantulas) Paratropididae (bald-legged spiders) Porrhothelidae Pycnothelidae Rhytidicolidae Stasimopidae Theraphosidae (true tarantulas) Araneomorphae Non-entelegynes Archaeidae (pelican spiders) Archoleptonetidae Austrochilidae Caponiidae Diguetidae (coneweb spiders) Drymusidae (false violin spiders) Dysderidae (woodlouse hunters) Filistatidae (crevice weaver spiders) Gradungulidae (large-clawed spiders) Huttoniidae Hypochilidae (lampshade spiders) Leptonetidae Mecysmaucheniidae Ochyroceratidae (midget ground weavers) Oonopidae (goblin spiders) Orsolobidae Pacullidae Palpimanidae (palp-footed spiders) Periegopidae Pholcidae (cellar spiders) Plectreuridae Psilodercidae Scytodidae (spitting spiders) Segestriidae (tube-dwelling spiders) Sicariidae (violin spiders, assassin spiders) Stenochilidae Telemidae (long-legged cave spiders) Tetrablemmidae (armored spiders) Trogloraptoridae (Trogloraptor marchingtoni) Entelegynae Agelenidae (araneomorph funnel weavers) Amaurobiidae (tangled nest spiders) Anapidae Ancylometidae Anyphaenidae (anyphaenid sac spiders) Araneidae (orb-weaver spiders) Arkyidae Cheiracanthiidae Cicurinidae Cithaeronidae Clubionidae (sac spiders) Corinnidae (dark sac spiders) Ctenidae (wandering spiders or tropical wolf spiders) Cyatholipidae Cybaeidae Cycloctenidae Deinopidae (net-casting spiders) Desidae (intertidal spiders) Dictynidae Dolomedidae Eresidae (velvet spiders) Fonteferreidae Gallieniellidae Gnaphosidae (flat-bellied ground spiders) Hahniidae (dwarf sheet spiders) Hersiliidae (tree trunk spiders) Homalonychidae Lamponidae (white-tailed spiders) Linyphiidae (sheet weavers or money spiders) Liocranidae (liocranid sac spiders) Lycosidae (wolf spiders) Macrobunidae Malkaridae (shield spiders) Megadictynidae Mimetidae (pirate spiders) Miturgidae Myrmecicultoridae Mysmenidae (spurred orb-weavers) Nesticidae (cave cobweb spiders) Nicodamidae (red and black spiders) Oecobiidae (disc web spiders) Oxyopidae (lynx spiders) Penestomidae Philodromidae (running crab spiders) Phrurolithidae Physoglenidae Phyxelididae Pimoidae Pisauridae (nursery web spiders) (including Halidae) Prodidomidae Psechridae Salticidae (jumping spiders) Selenopidae Senoculidae (bark hunters) Sparassidae (huntsman spiders) Stiphidiidae Symphytognathidae (dwarf orb-weavers) Synaphridae Synotaxidae Tetragnathidae (long jawed orb-weavers) Theridiidae (tangle-web spiders) Theridiosomatidae (ray spiders) Thomisidae (crab spiders) Titanoecidae Toxopidae Trachelidae Trachycosmidae Trechaleidae Trochanteriidae Udubidae Uloboridae (cribellate orb weavers) Viridasiidae Xenoctenidae Zodariidae (ant spiders) Zoropsidae Spider taxonomy Bold are families with more than 1000 species

Taxon identifiers Cybaeidae Wikidata: Q10028 Wikispecies: Cybaeidae ADW: Cybaeidae BioLib: 545 BOLD: 82354 BugGuide: 160687 CoL: 8TS EoL: 8806 Fauna Europaea: 10673 Fauna Europaea (new): e4a55f25-a15a-40c5-82e9-509b953257d4 GBIF: 3404 iNaturalist: 52950 IRMNG: 107721 ITIS: 871521 NBN: NBNSYS0100002701 NCBI: 134569 Open Tree of Life: 264296 Paleobiology Database: 57509 Plazi: 42C7A80B-2431-61A6-84B4-DC809B60D8F4 WSC: urn:lsid:nmbe.ch:spiderfam:0038

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