# Crurotarsal

> Mediated Wiki article. Canonical URL: https://mediated.wiki/source/Crurotarsal
> Markdown URL: https://mediated.wiki/source/Crurotarsal.md
> Source: https://en.wikipedia.org/wiki/Crurotarsal
> Source revision: 1220514446
> License: Creative Commons Attribution-ShareAlike 4.0 International (https://creativecommons.org/licenses/by-sa/4.0/)

Therian form of crurotarsal ankle. Adapted with permission from [Palaeos](https://web.archive.org/web/20100911212001/http://www.palaeos.com/Vertebrates/Units/430Mammalia/430.500.html#Theria)

A **crurotarsal** joint is one that's situated between the bones of *[crus](/source/Crus_(lower_leg))*, i.e. shin ([tibia](/source/Tibia) and [fibula](/source/Fibula)) and the proximal [tarsal](/source/Tarsus_(skeleton)) bones, i.e. [astragalus](/source/Talus_bone) and [calcaneum](/source/Calcaneus).[1][2]

The ankle joint of [therian](/source/Theria) [mammals](/source/Mammal) ([marsupials](/source/Marsupials) and [placentals](/source/Placentals)) is a crurotarsal joint, with the main joint of ankle bending between the tibia and the astragalus; the calcaneum has no contact with the tibia but forms a heel to which muscles can attach.

A group of [archosauriform](/source/Archosauriformes) [diapsids](/source/Diapsid), [Crurotarsi](/source/Crurotarsi) (including living [crocodilians](/source/Crocodilia) and their extinct relatives) is named after specialized crurotarsal joint in the skeletons of the members of this group, located between their fibula and calcaneum, with a hemicylindrical [condyle](/source/Condyle_(anatomy)) on the calcaneum articulating against fibula.[3] This joint is present in the skeletons of [suchians](/source/Suchia) (including crocodilians) and [phytosaurs](/source/Phytosaur), and was cited as one of the characters supporting uniting these two groups in a clade to the exclusion of [avemetatarsalian](/source/Avemetatarsalia) [archosaurs](/source/Archosaur) ([birds](/source/Bird) and their extinct relatives). However, according to a study published in 2011, suchians are more closely related to Avemetatarsalia than to phytosaurs; there is, however, not enough information to find out whether the aforementioned crurotarsal joint evolved independently in suchians and in phytosaurs, or whether it was already present in the skeleton of their most recent common ancestor (and secondarily lost in avemetatarsalians).[4]

 Crocodilian form of crurotarsal ankle. Adapted with permission from [Palaeos](https://web.archive.org/web/20050405074210/http://palaeos.com/Vertebrates/Units/270Archosauromorpha/270.500.html)

The ankle joint of [pseudosuchians](/source/Pseudosuchia) (including crocodilians) and phytosaurs, passing between the astragalus and calcaneum, is also called crurotarsal joint in the literature.[5][6] In the skeletons of the phytosaurs and most of the pseudosuchians this joint bends around a peg on the astragalus which fits into a socket in the calcaneum (the “crocodile normal” tarsus); only in the skeletons of the [ornithosuchid](/source/Ornithosuchidae) pseudosuchians a peg on the calcaneum fits into a socket in the astragalus (the “crocodile reversed” tarsus).[3] Strictly speaking this ankle is not a crurotarsal joint in the previously discussed sense, as it's situated between the two proximal tarsal bones. However, while calcaneum is not fixed to the fibula, the astragalus is fixed to the tibia by a [suture](/source/Suture_(anatomical)) and thus in practice it functions as an extension of the *crus*.[7]

## References

1. **[^](#cite_ref-1)** Sidney Frederic Harmer & Arthur Everett Shipley (eds.) (1902). *The Cambridge Natural History, vol. X: Mammalia*. The Macmillan Company. p. 43. {{[cite book](https://en.wikipedia.org/wiki/Template:Cite_book)}}: |author= has generic name ([help](https://en.wikipedia.org/wiki/Help:CS1_errors#generic_name))

1. **[^](#cite_ref-2)** Sidney Hugh Reynolds (1913). *The Vertebrate Skeleton (Cambridge Zoological Series)* (Second ed.). Cambridge University Press. p. 344.

1. ^ [***a***](#cite_ref-Sereno_3-0) [***b***](#cite_ref-Sereno_3-1) Sereno, Paul (1991). "Basal archosaurs: phylogenetic relationships and functional implications". *Journal of Vertebrate Paleontology*. **11** (Suppl. 4): 1–51. [doi](/source/Doi_(identifier)):[10.1080/02724634.1991.10011426](https://doi.org/10.1080%2F02724634.1991.10011426).

1. **[^](#cite_ref-NSJ11_4-0)** Nesbitt, S.J. (2011). ["The early evolution of archosaurs: relationships and the origin of major clades"](https://zenodo.org/record/5405335). *Bulletin of the American Museum of Natural History*. **352**: 1–292. [doi](/source/Doi_(identifier)):[10.1206/352.1](https://doi.org/10.1206%2F352.1). [S2CID](/source/S2CID_(identifier)) [83493714](https://api.semanticscholar.org/CorpusID:83493714).

1. **[^](#cite_ref-5)** Dyke G.J. & Kaiser G.W. (eds.), *Living Dinosaurs: The Evolutionary History of Modern Birds*, John Wiley & Sons, London, 2011, p. 406.

1. **[^](#cite_ref-6)** ["Glossary: crurotarsal - Palaeos"](http://www.palaeos.com/vertebrates/glossary/glossaryCr.html#Crurotarsal%20joint).

1. **[^](#cite_ref-7)** Guillaume Lecointre & Hervé Le Guyader (2006). [*The Tree of Life: A Phylogenetic Classification*](https://archive.org/details/treeoflifephylog0000leco/page/386). [Harvard University Press](/source/Harvard_University_Press). p. [386](https://archive.org/details/treeoflifephylog0000leco/page/386). [ISBN](/source/ISBN_(identifier)) [978-0-674-02183-9](https://en.wikipedia.org/wiki/Special:BookSources/978-0-674-02183-9).

---
Adapted from the Wikipedia article [Crurotarsal](https://en.wikipedia.org/wiki/Crurotarsal) by Wikipedia contributors ([contributor history](https://en.wikipedia.org/wiki/Crurotarsal?action=history)). Available under [Creative Commons Attribution-ShareAlike 4.0 International](https://creativecommons.org/licenses/by-sa/4.0/). Changes may have been made.